Novitattes PUBLISHED by the AMERICAN MUSEUM of NATURAL HISTORY CENTRAL PARK WEST at 79TH STREET NEW YORK, N.Y

AMERICAN MUSEUM Novitattes PUBLISHED BY THE AMERICAN MUSEUM OF NATURAL HISTORY CENTRAL PARK WEST AT 79TH STREET NEW YORK, N.Y. 10024 U.S.A. NUMBER 2633 OCTOBER 4, 1977

NORMAN I. PLATNICK AND MOHAMMAD U. SHADAB On Amazonian Cryptocellus (Arachnida, Ricinulei) ",-. I.1.-I ., 1,-I I. .-,....AII.I..,,j-..; ,,-....I,m, --..,- ;'- ..,. ":.I. :.-. .1 ..1 ..;..r.II...,,.,:,. ....I..IIo.`,.III:],..iII.-..,-.... I-1.,..,,,.. .:.. .-, .I-I-..I,-...I,,,- I..I..,....--,."II, .Ii..1..i. 1.,. ., --i,X% 1...i1..Ia..j. "I..;.:.i. ...II,ii1.II;.II..I...... I.....,II.,.,,.II9I1.:%I,I.II,...I.;-.1.".I""",.:f:..,.I;-:..,: .,--I.I.:,.I.,..I. .:';-.;."-.. -.iI-. ..II.,..,:I..II.1.I.I..I. ,.. :.,... :.'..I,.,..I ,.I.-..I.II ,..,-...I.I.I:,,1.-'...I111.IIIIII. 11.I-..-I'.....I-II.-.II-1-..I--...rI.-I..II.I.II.-".i...II.,iI.I,.-..-.,1.. .; I;.I-I II...-.I..;I-i,...I,iI-.-,11I.I,:II.IIII;..I. Ii.,,.,,..1II-I.1. .1.,:1..rI,.:.I,2.II. ..I,...III.I ..'--,....I:,II.Iz..,I,. ..,,.I;.:,,,,,.i.iII.I%,,...,iII..II...I.I1., ,...I .III:....I.II.II..I.I-....II., ,IIiI.I. .I.i.. .I.I.11.I. ..--I.II .i; ,....1:.,.II.II.I ..I.-.1. ,III.-%I.I-.I".IIII. .III.-I.-.I.Ii....II...,-.I I..,I.:I-,I I.:I.II ..I.,....,III,.-I,,., e.,.I.I,.;.III-II I.:I.r'IIIiII;1..I..I.I-I,.II....II.I.I-....I.IIIII.I.-,-.rIII..,.:. .III.I.-i.:.-.,mI-.::.II.1,..,.III. .I..1..q.I.6,:11III-..., .,,I. .141,.I.:..II I..1.-,,I..I..II.,-I-..,.,.,-.II..;,.,..-I:I.,,.I.I I..I.I-.I.I.I.I::,,171.II.-.I:-1i'- ,...."....-I .II. Ii ,II..I.,.I:4I..,I1.,I..11II-.I...I....11I.I.-...I...... :..",j.:, .c.lk.-I..I1-1:-II:II,-.:,. -.....I.. ,.I. .,. .: q..- I.IIIII..... IIIII.I.t,,,II.1...:.,II. ..-I... t..,. .I-t.IkII..II.-I.-I.-1.,%,,-.I.I:m.'.IIII...I,II..,I..-.II". ...II .I,:,.,.,.",I..I...I,.I..-I...... 1...I. .I....I-I.I...II",I.I,".,,. .,.II-.II%II.III I.II,.::r,..II.IIi:.:,.II-.-I.I.I...1I.IL,.I-. I..II..I...I.. ,.-... :., -.II: ",,.I...I.I.II..Ili ,....II.II IIII,..1....,,,I.,I,I.1.I..1 ..,.II ...... I-.-.I-i,.II-IIIII".I. .I,I.:..i....I.1I..,.1.-..I'L .,.I,-,.1I:.,I::I-..,".I.II..,I.II.-.,..-I... I.hI.....%II.I..-, ..I,-I.,,I1. ". ,..I, ...;I ,III..-II. liI..11 ..-IL.,,",...... z.. I.1. I;....,-IA:,.,..,,;.,.I.I.IIiI...I.I. ...,..-,.;II.II.vI,..II..I..,I-p,.- I..4.,,..I.....Z,..II.I,...I.-.-.I...,--;.I-,.,,,'I..I-1411 .,.-I.I.11..IIL.I-I....IIII...-:..I.."..,II..-I.-I....-., ,I.I.,t,Ii..IIII.,.,...I.. 11II.I.I.,.III.III... ..I..I ..,.,lI..,,.,141,..I,,I.,.,.,I..I-.., 1..I.IIII .;, III..,,...,...,I,.I.._..,Ii I....II:..II ;.I-", I.....1.I IIIII..I ....I.. ..I..II,-..., .;.I..I.III... I.j.. IIx- i.I-.,.I-I..,IIIw.i....,..:III,....,Ir:1 --.I, II.1.III .I...I- .... 11,.,.:I I:,.I..I.1,I..I.II. ...IIII%.I..III .I.. .I.. .,.I. .L.IIr,..IIII.I.,..I."..el-::, .-.I.%....II.I I.IaI..I.I...I.".I..I.II.:.I1.,,,..,IIil%..,1..-.IIII.-.-II.I.. .fI ..II.tIIIII..,..I..I:...1.,I.7..-.,....I.I... -.I.IIIIq.,I...4iII.1.:17.-;...,I,.....II,II.IL..III.II..; -4111"-1II. ..II.I--.-I.II..-,I.III1,..1..,I..'....'.:., ,- '.11:.L.0I.,..,.,...:....I,.II..I. IIII-I.,,. ,;%..,i--.-..".I. III..r.I..,I.III.I.,III.II. tIj..I...-,...,., ;.....I,-..I.II,.I., ,;pII.I.I,I,. ..1.II. i,.. .-, I.....,..I..,,)..";I .II.I.-,III..4.I.Iv,-II..III;, -,:,:,,.:.I..,IIIzI.-III.,"I,...-1,,I I...-.;.,I.I.I...I.,.,II!.I-I,,,,,-I..,..II.III..:m-..I.,I...;:I.,..,I,I,,IIcIII. i-.....III-1,II".".,..I.,".I.I....I-.II-I.IIIII.;..,,..Ii ,.II.....1.;,.,I.,I-..11.r..-..1.,.III II-.II1.I,,..-,.;-I.II11.-.Ii ,I IIIII.I.I..,II.,.i;I"i-,.-.,-I...'..II---. -.I..11,.%I I.I;....,,.- .I,,,-.I.II- II..I.II.II I..I... II.I...,`...,, .;.I,:;.:. ...,,. ,.-I,.--I- ,.I.'I,..-,..I.:,,-. ...,.:.;II..-I- II.I.I;. I.I.:-..II... .-I..;-...,,.;.I,e.I.,.I, II I..II.., .i,-. ,I.,II-.III.I -,.,, ..II- II-..I .IiIII.7,.'..I....4.,II,....1.....I11.,II.II..I....-.I.. I., ,I.. .,.I,I...... ,, ..I...e..-IIII.,. I.....I..-..rI.....,.-.I....I.. "I...:.;I,.I.-I.II ;.... .,..I....I1...L. I9I..III.- .II.-,II..-i:I.. ,...",II....: IIIIII.., ..II..,...I.,.I..III.1.-Ic.I..;,-.-....:II,....I.I.I.III;; Ir .. -,I. ,!1,I,.,..I.1,.,,,:,, ...,.I-,, .III .r-.IIII.II....,:.I.I.:".-.;I..1.I,.,III:..It.II,...I,..;m,:,I-I..I...I ..,I..1II11,..,%.I-.,.iIi.I.I,. .I-... :I..I,I.-:..- .II I.I..- I:- qI-...., ,%I; 1.I..1. -..I.':I...,,--...,. ,.. I... -IIII..,,j"-..,.I..rII.IIIII.-..,,-.I"II,,II.I...I..-.,I.II"III,I....I....I...III..r,.;Ii.. I...I.,-,-I,.II.I...I..,-.1%II.III... .III....". ,.II I-,.,.I..I:;.I,.,I.I.I.." ;..,,. %".I....II.I .II.I I.qIII.-..,.,...1 I.... -..I,.,. .I-. "I I,...,.,I I.:IIII I,IF.....I.I.II,";I .II,..II.I.I.I.:I,II.II.....I. I...:.- ,III.:II.I.III ,III., .1..I.I.III-.I1. II I.tI,.,..,.I,.::..".,-.....I,.-I..II,I ..1.-II..I..'.I..':IIiI,.,.."., ..1. .;:..,II ....JI.I.I:rx,'.,I....1. -I...1II-:.;,I.,-I,..;. I.II.Ii.;LI....;....., .!,..'.''.,.. ,I.I,:.. ..:,11,.`..,Z,,II.':.:;I.-I...IIII".,: ,III I.IIII,...... :II,, ,.- III.IiI. ,.I ...:. ,....,.,.-.I.I, i- :.,,III.,Iv--..II III.IIIIII.I.III.,..ll.I;..;,.II.;...,.... -.. -"III 11. I...I-..;..I..I-III._-, ,-..-;-..I::,..:,.., .IL' ....IIj...,.II.:..,.i.;.-I-.rIII... ..,,.IIII.I-. ...I....Ii,.Ii,-,.....II..j. i."III.II.. :II.1, ..;..I1 -I., ;-,II.III .I. .I,-1.Iq ..wI.. II, .III .;.I: wI.4-I.-. ,I III..;I.11 1..I..I.,.II-, I.I.,III. .-III,.. ;...,. --II1... .. ,:,..II.. .I-IIII....1.iII-..I.I1,I, .I.I., ...r-I.,II..,- .-..:-;..I.,t,.,.;,.I..,...,t.1.I,II...".,I-. I.I.I.I.,,-::.;l.IIIII.II...II,,,.I.I,I,..,-iII.1..,,I...:.II.III....II.w,,I.,III..,I,III.II..!..,..,.II...I.. ,oiI..1.. IIIII.IZ:: .,. I.:,1.-I..I I.I-.I.I."I.., .,i.IZ.,.. 11..1..-..I I.,.II ...". ,;IIII..-I"I i.II."I..I-..,I. III..iI ...-I. ,I.III-..lr;--II.I..I...I,-Ii,.1.II:..j. ..I11...III II..,,....., I. ,I..7on-.,:..,I 11II.I.w.j.II':,.-:.,.-IIj,II...,.'.I...I..:I., .-..III.I, .II..,I:".I.I.F.,.,.-.:,-.-.7..I I,..I.-I,I-."-II.,I.I.I IIII.II.,..:-I--I.I.",.. IIII.I.,I.I..... -."-::IIII,I.;..-:...II..II-.II.,- .,.,.I .-I11I.I.I...I.,i11...... ,,I..II"-:IIII.:-.1.,'.,-,. :.I".I.r...: .. I.II...-,III.I-I-J...1,III,4..-...I).I II.. I. ..,I-",.t.II-:.I.,..I..! .,....,., .I-..-.II. .,. ..II.I. II. III. III.vII1,,.. ,I ..,..I-7..II.I-.I. ,I. III..IIII.I .t.I,I-. ;.-I.I.I.I,. .I., .. ,.I: ;zI., ,i.-.I IIII:.,ImI.11I,-11,.,I-...-I.,..II.I.;I-II..II.i ,.-,-....;: .I.I-.,II...... -.IC..o.,:; .1IL- ,m, ..I.. ,I,-.I. .,I. .IIIII11Io..I,.I...;,-I,41.1,::"..:.I, .II.- .I.:...III.I .. ... I....I.-I-.I.-III.I...... 'I;I.:II. II... II1, 3... .I.I. ,,I!..k.':.,.I .I,.,...I.1 I.. --I;,..I.-,...I ,.I. ..,II- .. ...I.-,I ..k..I.-..-,-,.1.,..7..,I..II .; IIL-I..,,,I.,.I.II..,..I.--I..,:.,,II.. I. I.I..III..-. I.;:.III .,..I..q...... I...I.-,II.. .11.II..II-...III.-.lII-:": ..I..I.III.. I".-.I :II.IIII..,1I.....I ,II.-II .I.II..,..-III,...II:Iio-.,,,,...I., .1 .I.,.,I..:,.I..... I...I.I.:I.iI.II...I.- i: I, II-I....-..4.;I.I..,;I..II1,..II.-.,,..I -.I.,II1...I.'.,.....I..I ..I,I-... -I.,.,I4.,,: IIi,i..I I.III.I.III I.1..:.III.I...-:.IIII1.II.. ,....I1..II-.II.-;II.. II..I.IIIIIII II. "`.IIIII?I...I. .I;...... I.. ,;,..-I;,.I.I.IIIII,-,:,--1.m;. .:.I.I.II.I-,,...-I....,...1.I....,..I.II,.IIc.,.II.,:,4-..I.I.IIII-,,,II.I.II...III..,-. .I..1r'.III..c...i.....LI...e.I.I...,. .,...-1....I.I .;, ..I.I. -I,I,mI.,I-..I,.I-..I.....-.I.I.-I..I I.;..II.i-1.,..IL....I::,I, ,III-...:.... 11:,.7 ,I.,Ii.I.-..II.1117,..III .IIi...II.1.1.II,.I. :.-I:.,.....I,.,IzI.,....I .I,.1I.IIiI.I.,,.I.,.., ,I.11II. IL.....;1- .1,I I..I..II.:2II...... , ..,.',,,,.;. ":Z, .1I..I I.IIII...I...,..II.I.;..iI.I..I....II.,;. .I..,.i.I-.II.. .: ,..,.I..I- ..III. ..III,.. .:II,,.:..IIII."....-I ..I-I.II.-...I..:. I..,..,"-,. ,.III.I IIII...... ,.:.I .:.I"II ...II-...1. .-I,..."...- '.I.I..... II..,I..I-.,IIII.I.".I-.I.:I-,IiIjI.I11...I- .-.-,I.-.-.I,.,I.l .1,II. ,,, -::.-. :,.. ;-,,:. ...I.... .III.,.....II ,. 1',.,.:rIIII.....-'4, :.,I:...11...II. .I.I.i ,..III-: .II...I,I.II, .I-.I,III.,-I I.I.I..I.II...I.II-I.,Ii,.I...... I. j.:,IIII:...-..I.I ...... :.I., I-.I...I I..k:Z,,I;,II1 I,.-II ..7 ..,II ..:1:iIII. .II. -,..,:. .:., ,,I." .III..q--I.II.-I.I-., ,,....-o I,--.2.. II....%I...I11,.I,.....I.'..I,,-,,I..I,. nI.III-L1.I.:..-41,..,;...I. I"Iil:,.I.,7I-IIIL,.I .III.,: I.II.I...I.-I-,."1II:...1..,1. ,!.,,w .I.II,-,-.I..,:,...1 ..II:..I",.II.. .I...:I.-....,,.,.1..I.Iv. .. .:.:.,I,I. .1.7...,;I ..I...--..;,..,..-,..,.:I....i.....II.:'.III.,,,,...:",;.",,:.I,.'. ....,.I-;.....I..c.I.I..I ...,,,,II,-,--...... ,. ..] L" .,t ,.,I., ....I...I.-:., ....4...-I.II ,,.IIIzI..,,,....-.II..III...I.II.,,:.I-:..-III.,..III...-I,.,I,-.....,.,,.,...... -III.....I.. .I..,"..II...... ,;;.,.,I'..I.. -m1. ...IIII-.I...I..-III.II;..., .,.-..I.i.I ..,..I,.I,I.1.III..I.i...Ij.I,1.I:..I,I....I.,....., -. IIIIt,.:,II..I.-..,:1:;,,;I.I.....,.;.71,III..,..I.,.,..I.I.-,..I,!,I,"...".I..,...,...,.I.-,....II ....I:.I...... j.rI-..I.I,..;-.,;; )..-II.i, .,II ,..I- .IIII...I.I.I-.I.I,.,....I..I..IIIZ,i.,...?..1I:I.,.1411..I.-.% ..II..,,.,:..II--.,I.:..I.II. ..I7.41-.:... I.....,, ..-7r 1,II.I.,-I.:.... I..I11I..II..I.,-I,.r....II...,;.r..%w.T..,:.I-..II.-..-. .. ,..I.., .1.-..., .II -.II.I.II.I.II-II...I.,.rI ,,, i.I.:j..,.",II.I-11II;.,.:,I,....II..I11II-I1I.'.. -IIIIII ,I.I I,.i,IiI..i,.I-I':II4I,II-I.i...... I..1...I.I.., I.I.II ....I.I..I.,.. .,II,',i.I,I..I..--.,;I,,,.--:,.L..I,..'.I....I-,i..I- .I...I.,,-,,-qII-1.,,- :- ,IIII-I....II.. .11I.I.11.I...-..II...I.I-...II:,..:-..Ii.III.,,.1.I-..!I,.. .: .II..-ii, .II11".11..II. .--I ..I..II,I ..I,I1. .. -..I.Ii-II.',.II...IIIIII.....;I..,.....,,;.-.....I..,.,,.-:.:I: .I"..J..IIIIIII II.,.I-II II.II ....,-I..,.,LII,I-.II.,.:.:,..".I....II-_;,I,III.....-..I.p.I:I,II:. 1..I I.I..I,.i,.I.III`.;.I%I.,1, .II .I.,-:.II ...II.: 4.. ,!,., .4 II.III.IIII,,..III..,,.-i.IeIIiiiIiI.I..,.II...i.I,...I1,.I..I;III..I...,...-i-;.I.::.I II..11....1.II.IIII.I.I...,-IIo.'.I.11,.I...14,I-I-.I.II..;" .I,.I.rI.- ,;,II.i-'.... ,. ,,.II..I.,II. ..,,I.%.II. I.I.I;I"; I:..z.;....I-iI.II-...',.;I: ..II,.I.I..,..-I...,.IIII..,IIL'I,....II.;I.I--I .-.I. .I1.i"-..III.I....:.I11III.I.II.,II;11-I;-.IIII.I.,I..III..I I;.. .III...I,".F.,.I.I,..I-..I....1.; .,.., -I.i IIk:II.II.,,.--..,I..II.'. -1..... ,I.II. -I..I:II.-.m....;.:-.I.I:;.;t.II...I,.I-.,III.r.",.I.,I,II-1 .I.:I....IIII,.,. .I.;--....II .I.,....:,II:...;,,.-II., ,-;,II.-. ...I...I..1 .II ---..1...,iI....,..1....I.I,.- .I.,-I..II-I.IIII.:r1. III I.I.III....I,; ,11.., :.;,II....';-,-..,,Iri.11I..,,-I,:...;.., ;.,I. iII-..:;II.II.I I.I"I...;1';I,iI.II.,2..m,... " .,III,-",..!.,2;...1.....;IIiIII-1,:.".I..., ",..I.: .- I..'i. .-I;t.l.q:.,,"I.I...... IIIIII; .I..I.II I.I..II, ..II.II. I...... I...",...III.I1,..:It.,I..L.I.,.. r:.II IIII:-,I.I ,. ...,,. .,,II. ..I ..,i..-..I.I ..I..I.,I.1:III-II.1III.; IIIII.I,.--IIII..1,..-..;.I..,mIII,.II.I.1.,-II..-.I...I-,-,11....; .-I,,I. LI :..4,I. -.:I.III:,.I; .... ,I,I -.I...I.I .;I.;.-,,II.i.,"-I. I.II..I--,11.Iz.I-,.I,.I..'". .`,. .I.I III.IIIIIIII.j.-,11.:,.,I.L,IjIe..,.,:...,.III,.,,..,...I.,. ,,.-.I-.oj-...II..I.I..;.I .....,..,..L'..III.I..., ..I.I.,.IIII..;,IIIi.I,..II .II;II...-1-7-I-;.-.,!....,. ,IIII.II...,-I-,I ..I,...IIII;...o.1,III.,o.-.:IIIiq:,.I.,,.I..:.,II,..-....,."".II I..IIIIIi..I.I;I:.I1.L,I,,I..1,I...-.III.y II.I--II.. -I..III. .II;...i11..,.:..I:..r.I..5...II.I....-.;...!..I,,1.II.:1I-I1, -I..:;,;III',.I. -I....- I:.k,.II..1,,...1.I-...I..III;!.,I;..I,.",.,I.-,I.I.,..-I....vI,.,I.I.i1..I.....".;. ..,.,:.. III.;., I...II .,,,..I.II.iII.I.:.III....I.-II.,I....., .14.".-II,I .I.I.I-I,-.:..11 ,,I.I.,,; -1.,II.II1..II.,.dI,1II.,I,.,....-...I..I I. .:,I,.III.-1i:-., ...,I..,...I:li..II12.I...... ,.I, .,:-I .",I1..t.-I"I...:I.,.11IvI.II..II .III :1;II;;I.IIII,.,..II!I,.-II..:..I..II-..,II.II.I,.%.. -.. ,..I...... IIII .17-I"..i.III I.I.:,,I-,.I.I.,,I.II.-II..Ie.-I.I'. !. .II... I.,1. .. .1..,!,,. i.iI.III .. I..I .II...;. I.,I. .I,.4,I.IIIII..!..,III.I..,;,I.,I,-..1.,.-I-.....III;,II,I-.. 71. .,. ..It,I-II.IiIII.. I-I.,I-1.III.,.I.,III,:, ..,-.mI..IIII..II. II.I...,.:..I.I.-.,.I.,II.-,.I"..I,II.71..,I..,-I....I,.!,.1.,..II-.-7,:-. ,I.II....1, "I:.I II., II.r .I- I...I .I.,.I ,;,i..I,-.II.,I'.I..I,-I,I-I.III..I.. II.I.II.I.:..L.III.II..I.... .I.:-II.I,iI,.I11y.I.,....i. .I:.,4- ...It,..IIII ,II,. I,.::.II;I.I.I.I1..,.;..,...,1.-.,.I-4I.r11IIIII,.I... ..I -I'....,.I-II11.r,.I.-...-.III-..,.,,.I. .11 ..I.I:,III-7 ;IIII. ..:,,1.I...I.I.I..1,I.,.I.II.II.III,.,.I 1..,... III-.IIr.)- I-"-,I....-I...-;.Ic;II..z,.III.III.I-IIII.II:..III.,..II,,".I17111. IiII..I;,I.II...,I:I.....:II 1,I..II..- .II.I IIr..:... ,-.I.II,.-....,..I. :II.I-..I...I,I.,--.I.Im,,.I.,.".14. .II.;I.II.,:.II.I..I,,I .. ..I..,. ,..,.I.., ..q..,.I.,I..III.., ,1. -.z-.I,I.II..I,..I.,I. -I ,I..: ,.I...I1,7.,,,;,.,,,.,...I. ..,;,:.I. ..i.`,:-1IIII.:II.,....,.,m14---,.:..I...,.II1;I. ..I.IIi:..I.IIIIIIIIII..II-I. ..III,.:.. I..-..--..II...I.1.IIII.II.I.I ..'iIIII .I.-..II.,III.I.I.---..II.II.I.-I"I,,:..II."-I.-. .11...-- :...,.. ..I, ,,Id...... -II.I..,-I.I.,II-.,II.III.1.III,.. .II11-.,.IIII.I,,)II ImIIII..I.III.,..I,I.:.",.,,I.I.-I.IIIIIIIII..I.! 11IjI III .II...-':.I.. .II,,,I...II1,III..I.....II" -II-II..'..-I....I,, 1,, .I I.I.1,I.-.. .I'...Jl',II.1.I-..,.,.,w ,..,.,,I.;II,..;I. II..II.-...-..:.11 e,., ..,... .I.I .I..,;. ILIIIII,I,-..,..IIIII,.II.I.-.C,I.-. ,:..I'..",,.-:..,.;..t... .-,.11,,I. .I-I",.... .,.: -1 I. ,1.,. ,-..IIII,IL. .IIe.II.I.I-,I.,I"I. b.. ;7-...... I .-i,..,..I-.,..,..,....I.I...1-...I,I,.....I.;....,.,I..:,I.:,,..11.,,-,,...... 1."II. .,..I.,., ..". ,...;C.I...I--I;..,11..I,.,..I.II-;I.,.,.:.IIII.I.. I:.. .., ...I., .7, I,III. II j.,..I, .i..1.z.IIC.-::,.II ;I.I,. ..II I:.I...e.I.,Iw..I..I-.,L.IILII -. ,I:.I,I.,,I.I; IIII I.....'....I..,,II.:-II.I:7.-.. -:I.I.1.I..-I. ... ,-I....III ...... I ,...I,,. .. .II,,I...I...II ."..,.....I.II.-:..-.I,I...I;. I..-i-;,II..-..-IIIII.I.,IF.I', ., :.II1.,I11 II..,.I....I .;.I,;:1...:-. 1,z. ,,I.i-.1:.I.. .I.II.il...-1.-.-.III'.:,I.1.I...,.....,...... III..i...,,I.II,-,.-.".I ... .,...I- i.I, :III -.II..I,...:..II. .III ;..."I,.II.:;.. ..Ii.I ..I.,.II:..,...II,.I,.141-,,"..,,,11-...r.:. ,,,,., ..I..II..F ,- -.I. :...,I.I...4..i.,I ..;.':.. .II,I ...II.I..Ir:" II-I.,.-...,-j.1111,. II.,I,.;..I,:..I .I.I-I"-.I%-...I"I.,.,.I.p....I..k. I;I... AMERICAN MUSEUM Norntates PUBLISHED BY THE AMERICAN MUSEUM OF NATURAL HISTORY CENTRAL PARK WEST AT 79TH STREET, NEW YORK, N.Y. 10024 Number 2633, pp. 1-17, figs. 1-58 October 4, 1977

On Amazonian Cryptocellus (Arachnida, Ricinulei)

NORMAN I. PLATNICK1 AND MOHAMMAD U. SHADAB2

ABSTRACT A key is provided to the 10 known South provided. Three new species are described: C. American ricinuleid species. The synonymy of whitticki from Guyana, C. becki from Brazil, and Cryptocellus simonis Hansen and Sorensen with C. peckorum from Colombia. In a supplement, C. foedus Westwood is disclaimed. These two the recently proposed genus Heteroricinoides species and C. pseudocellatus Roewer are rede- Dumitrescu and Juvara-Bals is newly synony- scribed; illustrations of the spermathecae of C. mized with Cryptocellus. albosquamatus Cooke and C. lampeli Cooke are INTRODUCTION This paper, the second in a series on the order, established a second species, Cryptocellus arachnid order Ricinulei, reviews the known simonis, for a single male collected in Para, Bra- South American members of the group. Because zil. Since that time six additional South Ameri- of their extreme rarity in collections, previous can species have been described, although one of authors who have reported on South American these (Cryptocellus manni Ewing) has subse- Ricinulei have seldom been able to examine more quently been placed in synonymy (Platnick and than one or two species at a time, with the result Shadab, 1976). Beck and Schubart (1968) ob- that some specimens representing undescribed tained pairs of specimens from both Para and species have been misidentified and that some Amazonas, Brazil, and observed one of these named species have been erroneously synonym- pairs in copula; without consulting the original ized. We have made a concerted effort to exam- material of C. foedus and C. simonis (or the male ine all previously reported South American speci- from Guyana attributed to C. simonis by Whit- mens as well as newly available material; the few tick, 1938), they identified the females from localities from which specimens are known are both Brazilian areas as the former species, the shown in figure 58. males from both areas as the latter species, and Westwood (1874) described the first known synonymized the two. Brignoli (1974), without American ricinuleid, Cryptocellus foedus, on the having seen any specimens, objected strongly to basis of a single female taken by Bates in the this synonymy on the grounds that the behav- Amazonian region of Brazil, and Hansen and ioral observations support only a hypothesis of Sorensen (1904), in their monograph on the conspecificity in one of the populations and that

'Associate Curator, Department of Entomology, the American Museum of Natural History. 2Scientific Assistant, Department of Entomology, the American Museum of Natural History.

Copyright © The American Museum of Natural History 1977 ISSN 0003-0082 / Price $1.45 2 AMERICAN MUSEUM NOVITATES NO. 2633 the resulting distribution of the species is unten- lampeli reveals many of the same modifications able in view of the extremely restricted distribu- found by Pittard and Mitchell (1972) in C. tions presently known for all other members of pelaezi: a large treelike seta and numerous bent the genus. Although these arguments are not con- setae (fig. 5) and a peg seta and shallow oval pits vincing in themselves, direct observation of the bearing peripheral setae (fig. 6). The entire body relevant material indicates that Brignoli is cor- of C albosquamatus is coated with large boat- rect. The male from Guyana noted by Whittick is shaped setae (figs. 7, 8) that seem structurally very different from the Brazilian males, the Para identical with the navicular setae described by and Amazonas specimens are distinguishable in Legg (1976, pl. 1) in African Ricinoides. both sexes, and females from both populations Any serious attempt to investigate the phy- are separable from the holotype of C. foedus logeny of the Ricinulei must await a review of (from an indeterminate Amazonian locality); all the numerous Central American and Mexican these forms are redescribed below. forms, but some preliminary observations can be Restudy of the available South American made. There is, of course, no doubt as to the material has also allowed us to investigate the monophyly of at least the Recent Ricinulei; the value of the female spermathecae, first noted by cucullus, the abdominal coupling mechanism, Brignoli (1974), as a taxonomic character. Differ- and the male copulatory apparatus are all unique ences in these structures can be detected in all to the group, and Hansen and Sorensen (1904) eight species of which females are known; in noted numerous other autapomorphic characters some cases, such as Cryptocellus albosquamatus of the order. The division of the group into two Cooke (fig. 52), they are very distinctive; in genera, Ricinoides (in tropical West Africa) and other cases, such as the complex including C. Cryptocellus (in tropical America) has not been foedus, C. simonis, and C. becki, the differences investigated or seriously questioned since Hansen are slight and somatic morphology provides more and Sorensen presented a list of about a dozen readily usable characters. Illustrations of the distinguishing characters of the two. Those au- spermathecae of all species other than the two thors knew only two very closely related and rela- Colombian ones already treated by Platnick and tively atypical species of Cryptocellus, so it is not Shadab (1976) are provided below. In addition, surprising that some of their suggested characters the one remaining species without a detailed (such as the relative widths of coxae II and III modern description, Cryptocellus pseudocellatus and the shape of the movable finger of the chelic- Roewer, is redescribed below. erae) are no longer valid, particularly for the Enough material of two species, Cryptocellus troglobitic species. Of the basic structural (rather lampeli Cooke and C. albosquamatus, has been than proportional) characters, most (an anterior available to allow some observation by scanning tooth on the chelicerae, a separate dorsal groove electron microscopy. One frequently cited char- for the metatarsal process on the male metatarsus acter, the presence or absence of a notch in the III, a lamina cyathiformis that adducts com- posterior dorsal margin of the basal pygidial seg- pletely against the metatarsal process and en- ment, must be used carefully as some species closes the male tarsal process, and a wide lateral have only slight notches (compare figs. 14, 20, flange on the body of the tarsal process) are re- and 42); only when the notch is complete is the stricted to Ricinoides; Cryptocellus is generally character conspicuous (fig. 1). At least three recognized by the absence of these characters. types of tubercles can be found on C. lampeli; The question thus arises as to whether there are the carapace bears flat tubercles like those found any characters that can be considered synapo- in the cuticular pits of Cryptocellus magnus morphic for Cryptocellus. Of course the genera Ewing (compare fig. 3 with Platnick and Shadab, are geographically disjunct and it is therefore 1976, fig. 5), the palpal trochanter bears low convenient to think of them as sister groups, but conical tubercles (fig. 2) like those found in the South America-Africa vicariance event need Cryptocellus pelaezi Coronado (see Pittard and not be correlated with the earliest dichotomy in Mitchell, 1972, fig. 18), and the first metatarsus the history of the Recent Ricinulei. If no derived bears tall conical tubercles (fig. 4). Examination characters shared uniquely by all the Crypto- of the distal tarsomere of legs I and II of C cellus species can be found, it is possible that the 1977 PLATNICK AND SHADAB: CR YPTOCELL US 3 group is paraphyletic and that some of its mem- the female spermathecae are short and wide (fig. bers are more closely related to Ricinoides than 12); relationships within this group will probably to each other. The navicular setae of C. albo- remain obscure until the male of C. foedus and squamatus, for example, might be regarded as the female of C. whitticki are discovered. The evidence of such a (more recent) transatlantic re- three remaining species (C. peckorum, C. lampeli, lationship. There is at least one character, how- and C. albosquamatus) do not appear to be ever, that might be considered synapomorphic closely related to each other; the first of these for Cryptocellus: the presence of distinct invagi- may be the plesiomorphic sister taxon of the nations of the dorsodistal margins of the final foedus group, judging by its slightly expanded tarsomeres of legs III and IV. Similar invagina- metatarsus III (fig. 54). tions are found in some other arachnids (such as As in our previous collaborations, the senior oncopodid opilionids; Suzuki, 1976, fig. 2), how- author has been responsible for the taxonomic ever, so serious consideration of this question re- analysis and the preparation of the text and the quires an analysis of the arachnid orders to deter- junior author for the preparation of the illustra- mine what the closest relatives of the Ricinulei tions; the scanning electron micrographs were ob- may be; Kraus (1976) has commented aptly on tained with the assistance of Mr. Robert J. Koest- the difficulty of such an analysis. It is worth ler. We are grateful to the following curators and noting that if the presence of a pedicel is re- institutions for supplying material: Drs. R. E. garded as a synapomorphic character, as sug- Crabill, Jr., National Museum of Natural History, gested by Kraus, the Ricinulei will have to be Smithsonian Institution; H. Dybas, Field Mu- included in the group thus formed; the fact that seum of Natural History (FMNH); M. Grasshoff, the pedicel is normally externally invisible be- Natur-Museum Senckenberg (NMS); M. Hubert, cause of the abdominal coupling mechanism does Museum National d'Histoire Naturelle (MNHN); not justify consideration of the order as non- H. W. Levi, Museum of Comparative Zoology, pedicelate. Harvard University (MCZ); E. Taylor and M. W. Some preliminary hypotheses on the inter- R. de V. Graham, Hope Department of Ento- relationships of the South American Ricinulei mology, Oxford University (HDO); and F. can be offered. The "pseudocelli" described by Wanless, British Museum (Natural History) Roewer (1952) in the Peruvian C. pseudocellatus (BMNH). are not the two translucent areas on the margins All measurements cited below are in milli- of the carapace between legs I and II, but eight meters. (out of many) deep pits of the same type found in the Colombian C. magnus (Platnick and KEY TO SOUTH AMERICAN SPECIES OF Shadab, 1976, figs. 1, 2, 4, and 5). The presence CR YPTOCELL US of these pits is probably synapomorphic and the two species are probably sister taxa; the unusual 1. Body and legs covered with large navicular setae (figs. 7, 8); spermathecae triangular straight and massive accessory piece of the male (fig. 52); Guyana. . albosquamatus Cooke tarsal process in C. magnus and the Colombian Body and legs without navicular setae; Cryptocellus glenoides Cooke and Shadab may spermathecae not triangular-...... 2 indicate that the latter species is the plesio- 2. Carapace with deep pits containing tubercles morphic sister group of C. magnus plus C. (Platnick and Shadab, 1976, figs. 1, 2, 4, pseudocellatus. If this is the case, the male of C. 5) ...... 3 pseudocellatus, when discovered, should also Carapace without deep pits ...... 4 prove to have a straight and massive accessory 3. Accessory piece of male tarsal process trifid Four other C. (Platnick and Shadab, 1976, fig. 13); piece. species (C. foedus, simonis, dorsal margin of spermathecae smooth C. becki, and C. whitticki) form a monophyletic (Platnick and Shadab, 1976, fig. 14); group with the following unique characters: the Magdalena, Colombia.... magnus Ewing male cucuilus has a strong depression below its Males unknown; dorsal margin of sper- dorsal margin (fig. 22), the male metatarsus III is mathecae invaginated (fig. 5 1); Caja- greatly expanded (fig. 24), the male trochanters marca, Peru . . . . pseudocellatus Roewer on legs III and IV bear apophyses (fig. 23), and 4. Males ....5...... 5 4 AMERICAN MUSEUM NOVITATES NO. 2633

FIGS. 1-4. Scanning electron micrographs, Cryptocellus lampeli, female. 1. Basal segment of pygidium, showing notch, dorsal view, 500x. 2. Tubercle from palpal trochanter, ventral view, 5000X. 3. Tubercles from carapace, dorsal view, 2000X. 4. Tubercle from dorsal ridge of metatarsus I, 2000X.

Females ...... 10 tubercles (fig. 25); Para, Brazil ......

5. Femur I with proximal and distal prolatero- ...... simonis Hansen and Sorensen ventral knobs (Hansen and Sorensen, Cucullus and median plate of tergite 11 with 1904, fig. 3g); trochanters III and IV with few tubercles (figs. 43, 44); femur III ele- apophyses (as in fig. 23); cucullus with a vated dorsally, with many tubercles (fig. proximal depression (as in fig. 22) . . . .6 47); Amazonas, Brazil ...... Femur I without knobs; trochanters III and ...... becki, new species IV without apophyses; cucullus without 8. Accessory piece of male tarsal process thick proximal depression ...... 8 and straight (Cooke and Shadab, 1973, 6. Apophyses on trochanters III and IV round- fig. 37); Valle, Colombia . . ed (fig. 3 1); Guyana...... glenoides Cooke and Shadab ...... whitticki, new species Accessory piece of male tarsal process thin

Apophyses on trochanters III and IV point- and curved (as in fig. 55)...... 9 ed (figs. 23, 45); Brazil ...... 7 9. Carapace almost devoid of tubercles; Guyana 7. Cucullus and median plate of tergite 11 with ...l...... lampeli Cooke many tubercles (figs. 21, 22); femur III Carapace almost completely covered with not elevated dorsally, with few prolateral tubercles; Amazonas, Colombia ...... 1977 PLATNICK AND SHADAB: CR YPTOCELL US 5

FIGS. 5-8. Scanning electron micrographs. 5, 6. Cryptocellus lampeli, female. 5. Tarsus I, dorsal view, lOOOx. 6. Tarsus II, dorsal view, IOQOX. 7, 8. C. albosquamatus, nymph. 7. Femur I, showing navicular setae, 250X. 8. Single navicular seta, 2500X.

...... peckorum, new species 17); femur I not expanded dorsally (fig. 10. Spermathecae longer than wide (Platnick 19); pygidium unnotched (fig. 20); Paria, and Shadab, 1976, fig. 17); Valle, Brazil . . .. simonis Hansen and Sorensen Colombia. . glenoides Cooke and Shadab Carapace and median plate of tergite 11 with Spermathecae wider than long (as in fig. few scattered tubercles (figs. 37, 39); 53) ...... 11 femur I expanded dorsally (fig. 41); 11. Spermathecae wider laterally than medially pygidium notched (fig. 42); Amazonas,

(fig. 53); Guyana ...... lampeli Cooke Brazil.. becki, new species Spermathecae wider medially than laterally (figs. 12, 18, 40) .12 12. Carapace and median plate of tergite 11 al- Cryptocellus foedus Westwood most completely covered with tubercles Figures 9-14 (figs. 9, 1 1) ...... foedus Westwood Carapace and median plate of tergite 11 with Cryptocellus foedus Westwood, 1874, p. 201, pl. scattered tubercles only ...... 13 37, figs. 5, 5a-5c (female holotype from 13. Carapace and median plate of tergite 1 1 with "Amazonia," somewhere between Belem in numerous scattered tubercles (figs. 15, Para and Sao Paulo de Olivenca in Amazonas, 6 AMERICAN MUSEUM NOVITATES NO. 2633

Brazil, in HDO, examined). Hansen and Soren- I II III IV Palp sen, 1904, 155, pl. 9, figs. 2a-2e. p. Coxa 0.82 1.04 0.94 0.86 0.40 Trochanter I 0.58 0.65 0.47 0.61 0.50 Diagnosis. Males of C. foedus are unknown; Trochanter II 0.54 0.65 0.39 females may be distinguished from those of C. Femur 1.12 1.62 1.09 1.22 0.86 simonis and C. becki by the much larger number Patella 0.73 0.94 0.81 0.77 of tubercles on the carapace, cucullus, and medi- Tibia 0.97 1.37 0.88 0.94 1.40 an plate of tergite 11 (figs. 9-1 1), the ventrally Metatarsus 1.06 1.53 0.95 0.96 but not dorsally expanded femur I (fig. 13), and Tarsus 0.49 1.51 0.83 0.86 0.14 the shallowly notched pygidium (fig. 14). Total 5.78 8.66 6.51 6.87 3.69 Female. Total length 4.64. Carapace 1.85 long, 1.83 wide between legs II and III, where Second legs not greatly widened; femur I about widest, dark red with distinctly darkened bor- twice, femur II about three times as long as wide. ders, with translucent pale yellow areas near mar- Tarsal claws stout, evenly curved. Spermathecae gins between legs I and II; surface, except for wide, short (fig. 12). margins, rather evenly and completely covered Male. Unknown. with tubercles (fig. 9). Cucullus 0.90 long, 1.13 Material Examined. Only the holotype. wide, dark red, with long setae and numerous Distribution. The type specimen was collected scattered tubercles most densely packed below by H. W. Bates in the Amazon basin. A label with dorsal margin; lateral lobes slightly protuberant the specimen gives the date 1861, which was thus (fig. 10). Left chelicera: movable finger laterally reported by Hansen and Sorensen (1904) as the thin, flattened posteriorly, slightly widened date of collection. Edward Clodd's memoir of transversely, armed with seven teeth increasing in Bates (in Bates, 1892) indicates that Bates re- length distally; fixed finger armed with four turned to England by at least 1860, having de- teeth of which most distal is much the largest. parted from Brazil in June 1859. We have been Sternal region with coxae I not meeting trito- unable to find any reference to the specimen in sternum; coxae II meeting along their posterior Bates's narrative of his expedition, so it would four-fifths, their suture line only slightly longer appear that the type locality of this species can than that of coxae III; coxae IV almost meeting only be said to be somewhere along the Amazon, distally. Abdomen 2.99 long, 2.63 wide near with limits set only by the extent of Bates's front of tergite 12, where widest, coloration as in itinerary, cited above. carapace except for brown articular membranes, with numerous tubercles evenly coating all tergal Cryptocellus simonis Hansen and Sorensen plates (fig. 11) and U-shaped area covering all of Figures 15-28 sternite 13 and lateral areas of sternite 12; me- Cryptocellus simonis Hansen and Sorensen, dian plates of tergites 11-13 and corresponding 1904, p. 156, pl. 9, figs. 3a-3i (male holotype sternites with paired lateral depressions, depres- from Le Para [=Belem], Para, Brazil, in sions of sternite 11 triangular, filled with tuber- MNHN, examined). cles; median plates of all tergites wider than long. Cryptocellus foedus (misidentification): Beck Pygidium with shallow notch in posterior dorsal and Schubart, 1968, p. 75 (in part; specimens margin of basal segment (fig. 14). Pedipalp coxae from Para only). light red, distal segments orange; coxae and Diagnosis. Females of C. simonis may be dis- trochanters with scattered ventral and dorsal tu- tinguished from those of C. foedus and C. becki bercles, femora with few proximal tubercles. Leg by the moderate number of tubercles on the formula 2431. Legs dark red, lighter distally, carapace, cucullus, and median plate of tergite 11 coated with short setae; coxae with few small (figs. 15-17), the unexpanded femur I (fig. 19), tubercles at margins; femur I expanded ventrally and the unnotched pygidium (fig. 20). Males of but not dorsally (fig. 13); distal leg segments C. simonis may be distinguished from those of C. with relatively few scattered tubercles. becki by the larger number of tubercles on the 1977 PLATNICK AND SHADAB: CR YPTOCELL US 7 cucullus and median plate of tergite 11 (figs. 21, their suture line about one and one-third times as 22), the smaller number of prolateral tubercles long as that of coxae III; coxae IV not meeting. on femur III (fig. 25), and the lack of a distinct Abdomen 3.02 long, 2.54 wide near middle of widening below the tip of the tarsal process (fig. tergite 12, where widest; coloration, setation, 26), and from those of C. whitticki by the much and ornamentation as in carapace except for longer apophyses on trochanters III and IV brownish orange articular membranes, with nu- (fig. 23). merous tubercles evenly distributed except on Female. Total length 4.57. Carapace 1.80 bare patch anteromedially on median plate of long, 1.67 wide at front of leg III, where widest, tergite 11 (fig. 17); median plates of tergites dark red laterally and posteriorly, lighter medi- 11-13 with paired lateral glabrous depressions, ally and anteriorly, with very small translucent corresponding sternites with similar tuberculate yellow areas near margins between legs I and II; depressions, with numerous tubercles on dark U- surface, except for midline, with scattered large shaped band covering all of sternite 13 and lat- tubercles (fig. 15) and short recumbent white eral areas of sternites 11 and 12, surrounding setae. Cucullus 0.83 long, 0.97 wide, dark red lighter smooth median areas of those sternites; proximally, lighter distally, evenly coated with median plates of all tergites wider than long. large tubercles and white setae becoming longer Pygidium with narrowed dorsal margin of basal distally; lateral lobes only slightly protuberant segment but without distinct notch (fig. 20). (fig. 16). Left chelicera: movable finger laterally Pedipalps orange; coxae and trochanters with thin, flattened posteriorly, slightly widened ventral tubercles, femora with proximal dorsal transversely, armed with seven teeth of which and ventral tubercles. Leg formula 2431. Legs most proximal is smallest, others subequal in dark red proximally, lighter distally, coated with length; fixed finger armed with four teeth of fine white setae; femur I not expanded dorsally which most distal is much the largest. Sternal or ventrally (fig. 19); segments with relatively region with coxae I not meeting tritosternum; few scattered tubercles except on dorsal ridges coxae II meeting along their posterior two-thirds, and sides of metatarsi, where numerous.

FIGS. 9-14. Cryptocellus foedus, female. 9. Carapace, dorsal view. 10. Cucullus, anterior view. 11. Median plate of tergite 1 1, dorsal view. 12. Spermathecae, posterior view. 13. Femur I, prolateral view. 14. Pygidium, dorsal view. 8 AMERICAN MUSEUM NOVITATES NO. 2633

17 X. ; Jngt... _..~~~~~~~~~~~~~~~~~~~~~~~~~~~~~~~~~~~,)S~~~~~~~~~~~i.oa..g

FIGS. 15-20. Cryptocellus simonis, female. 15. Carapace, dorsal view. 16. Cucullus, anterior view. 17. Median plate of tergite 11, dorsal view. 18. Spermathecae, posterior view. 19. Femur I, prolateral view. 20. Pygidium, dorsal view.

I II III IV Palp able finger laterally thick, flattened posteriorly, Coxa 0.72 1.10 0.88 0.81 0.43 greatly widened transversely, armed with seven Trochanter I 0.47 0.61 0.50 0.63 0.50 teeth of which most proximal is smallest, others Trochanter II - 0.47 0.57 0.36 becoming longer distally, last four grouped in Femur 1.10 1.73 1.14 1.19 0.86 two pairs; fixed finger armed with four teeth of Patella 0.70 0.83 0.65 0.78 - which most distal is much the largest. Sternal Tibia 0.90 1.32 0.90 0.94 1.22 region with coxae I not meeting tritosternum; Metatarsus 0.94 1.58 0.83 0.99 - coxae II meeting along their posterior eight- Tarsus 0.47 1.30 0.79 0.90 0.14 ninths, their suture line almost twice as long as Total 5.30 8.47 6.16 6.81 3.51 that of coxae III; coxae IV meeting. Abdomen 3.35 long, 2.38 wide near middle of tergite 12, Second legs no)t noticeably widened; femur I where widest, coloration lighter than carapace, three times, fern nur II five times as long as wide. all tergites with numerous tubercles most densely Tarsal claws thiin, evenly curved. Spermathecae packed on median plates and on tergite 11 (fig. wide, short (fig. 18). 21); setation sparse; articular membranes beige; Male. Total Ilength 5.08. Carapace 2.05 long, median plates of tergites 11-13 with paired tuber- 1.69 wide at baick of leg II, where widest, dark culate lateral depressions, corresponding sternites red anteriorly a]nd laterally, lighter medially and with similar depressions, with numerous tuber- posteriorly, wit]h very small translucent yellow cles on U-shaped band covering all of sternite 13 areas at marginss between legs I and II; surface, and posterior one-third of sternite 12 and sides except for anterior end, almost completely of sternites 11 and 12; additional tubercles along covered with nuimerous tubercles, with fine white anterior and posterior margins of sternite 11; me- setae clustered toward margins. Cucullus 0.76 dian plates of all tergites wider than long. Pygidi- long, 1.19 wide, dark red proximally, grading to um with narrowed dorsal margin of basal seg- yellow distally; distal half with scattered tuber- ment but without distinct notch. Pedipalps cles; surface wiith long fine setae; lateral lobes orange; coxae, trochanters, and ventroproximal strongly protubierant, distal margin invaginated edge of femora with scattered tubercles. Leg between lobes; Iproximal margin curved, followed formula 2341. Legs light red proximally, lighter by deep depression (fig. 22). Left chelicera: mov- distally, sparsely coated with fine white setae; 1977 PLATNICK AND SHADAB: CR YPTOCELL US 9 segments with rel atively few scattered tubercles trochanter of leg III with prolateroventral spur, except ventrally cin metatarsus I and prolaterally first trochanter of leg IV with retrolateroventral on femur III (fig. 25), where numerous. spur (fig. 23). Second legs not noticeably widened; femur I two and one-half, femur II four I II III IV Palp times as long as wide. Tarsal claws thin, evenly Coxa 01.58 1.24 1.01 0.78 0.42 curved. Copulatory apparatus as in figures 24, Trochanter I 01.83 0.79 0.47 0.71 0.47 26-28. Trochanter II - - 0.73 0.61 0.40 Material Examined. The holotype male (de- Femur .24 1.74 1.55 1.58 0.89 scribed above) and a male and female from Patella 0).83 0.97 0.79 0.83 - Utinga, near Belem, Para, Brazil (L. Beck, 1966, Tibia .01 1.48 1.01 1.06 1.30 NMS). Metatarsus 1 .30 1.84 1.19 1.01 - Variation. The male from Utinga has slightly Tarsus 0).47 1.62 1.37 1.19 0.16 fewer tubercles on the carapace and abdomen Total 6.26 9.68 8.12 7.77 3.64 and slightly more tubercles on the prolateral surface of tibia III. Legg with niimern ii sexiil mAdificationv femiir Distribution. Known only from Para, Brazil. I with ventral spur at proximal end and two con- nected spurs at distal end on prolateral side of Cryptocellus whitticki, new species venter; tibia and metatarsus I each with prolat- Figures 29-36 eroventral rounded knob; femur II with slight Cryptocellus simonis (misidentification): Whit- prolateroventral knob at proximal end; second tick, 1938, p. 479.

ii_1 2

FIGS. 2 1-28. Cryptocellus simonis, male. 21. Median plate of tergite 11, dorsal view. 22. Cucullus, anterior view. 23. Trochanters III and IV, ventral view. 24. Leg III, anterior view. 25. Femur III, prolateral view. 26. Tarsal process, anterior view. 27. Tarsal process, posterior view. 28. Leg III, posterior view. lo AMERICAN MUSEUM NOVITATES NO. 2633

4 4

-~~~~ ra

FIGS. 29-36. Cryptocellus whitticki, male. 29. Median plate of tergite 11, dorsal view. 30. Cucullus, anterior view. 31. Trochanters III and IV, ventral view. 32. Leg III, anterior view. 33. Femur III, prolateral view. 34. Tarsal process, anterior view. 35. Tarsal process, posterior view. 36. Leg III, posterior view. Type. Male holotype from New River district, distal half with few scattered tubercles, margins elevation 750 feet, Rupununi, Guyana (January- with long white setae; lateral lobes protuberant, March 1938; C. A. Hudson), deposited in BMNH. distal margin slightly invaginated between lobes; Etymology. The specific name is a patronym proximal margin followed by deep depression in honor of R. J. Whittick, who first reported on (fig. 30). Left chelicera: movable finger laterally the holotype. thick, widened transversely, armed with 10 teeth Diagnosis. Females of C. whitticki are un- of which two most proximal are reduced and two known; males may be distinguished from those most distal are largest; fixed finger armed with of C. simonis and C. becki by the shorter and four teeth of which most distal is much the long- more rounded apophyses on trochanters III and est and two most distal are bifid. Sternal region IV (fig. 31), the smaller number of tubercles on with coxae I not meeting tritosternum; coxae II the cucullus and median plate of tergite 11 (figs. meeting along their entire length, their suture 29, 30), and the absence of prolateral tubercles line only slightly longer than that of coxae III; on femur III (fig. 33). coxae IV meeting anteriorly. Abdomen 3.49 Female. Unknown. long, 2.45 wide near middle of tergite 12, where Male. Total length 5.40. Carapace 2.19 long, widest; coloration lighter than carapace, tergites 1.91 wide at middle of leg III, where widest, dark with few tubercles (fig. 29); scattered long thin red around margins, lighter medially, with small white setae; articular membranes light yellow; translucent pale areas at margins between legs I median plates of tergites 11-13 with paired lat- and II; surface almost devoid of tubercles, with eral depressions and greatly elevated median short white setae. Cucullus 0.76 long, 1.19 wide, areas, corresponding sternites with similar depres- dark red proximally, grading to yellow distally; sions, with only a few scattered tubercles poste- 1977 PLATNICK AND SHADAB: CR YPTOCELL US I11 riorly; median plates of all tergites wider than Material Examined. Only the holotype; the long. Pygidium with narrowed dorsal margin of BMNH houses a tritonymph male taken by the basal segment but without distinct notch. Pedi- same collector in Guyana (no specific locality) in palp coxae light red, other segments orange, June 1938, that may well belong to this species. without tubercles; coxae with pair of short thick Distribution. Known only from Guyana. white setae along inner margin. Leg formula 2341. Legs light red proximally, lighter distally, Cryptocellus becki, new species sparsely coated with long white setae; distal seg- Figures 37-50 ments with few scattered tubercles except under metatarsus I, where numerous. Cryptocellus foedus (misidentification): Beck and Schubart, 1968, p. 75 (in part; specimens I II III IV Palp from Amazonas only). Coxa 0.83 1.19 1.06 0.86 0.36 Trochanter I 0.76 0.83 0.74 0.67 0.50 Types. Male holotype and female paratype Trochanter II 0.72 0.61 0.47 from Reserva Ducke, near Manaus, Amazonas, Femur 1.44 2.02 1.62 1.69 0.86 Brazil (1966; L. Beck), deposited in NMS. Patella 0.94 1.12 0.86 0.86 Etymology. The specific name is a patronym Tibia 0.97 1.58 0.94 1.10 1.30 in honor of the collector of the type specimens. Metatarsus 1.18 1.69 1.19 1.12 Diagnosis. Females of C. becki may be dis- Tarsus 0.47 1.63 1.33 1.08 0.18 tinguished from those of C. foedus and C. Total 6.59 10.06 8.46 6.87 3.67 simonis by the small number of tubercles on the carapace, cucullus, and median plate of tergite 11 Legs with sexual modifications as in C. simonis (figs. 37-39), the dorsally and ventrally expanded except femur II prolateroventral knob absent and femur I (fig. 41), and the distinctly notched apophyses of trochanters shorter, rounded (fig. pygidium (fig. 42). Males of C. becki may be 33). Second legs not noticeably widened; femur I distinguished from those of C. simonis by the three times, femur II four times as long as wide. smaller number of tubercles on the cucullus and Tarsal claws thin, evenly curved. Copulatory ap- median plate of tergite 11 (figs. 43, 44), the paratus as in figures 32, 34-36. larger number of prolateral tubercles on femur

642

FIGS. 37-42. Cryptocellus becki, female. 37. Carapace, dorsal view. 38. Cucullus, anterior view. 39. Median plate of tergite 11, dorsal view. 40. Spermathecae, posterior view. 41. Femur I, prolateral view. 42. Pygidium, dorsal view. 12 AMERICAN MUSEUM NOVITATES NO. 2633

f~~44C3b 4

'-4 5

K .- 48

FIGS. 43-50. Cryptocellus becki, male. 43. Median plate of tergite 11, dorsal view. 44. Cucullus, anterior view. 45. Trochanters III and IV, ventral view. 46. Leg III, anterior view. 47. Femur III, prolateral view. 48. Tarsal process, anterior view. 49. Tarsal process, posterior view. 50. Leg III, posterior view.

III (fig. 47), and the presence of a distinct widen- posterior two-thirds, their suture line slightly ing below the tip of the tarsal process (figs. 46, longer than that of coxae III; coxae IV meeting 48), and from those of C. whitticki by the much anteriorly. Abdomen 3.56 long, 2.63 wide be- longer apophyses on trochanters III and IV (fig. hind middle of tergite 12, where widest, colora- 45). tion as in carapace except for beige articular Female. Total length 5.00. Carapace 1.84 membranes, with scattered tubercles sparse ante- long, 1.69 wide between legs II and III, where riorly (flg. 39), becoming quite dense posteriorly widest, dark red along margins, lighter medially, and on U-shaped area covering all of sternite 13 with round white translucent area at margins be- and lateral areas of sternite 12; median plates of hind leg I; surface with few scattered tubercles tergites 11-13 and corresponding sternites with (fig. 37). Cucullus 0.83 long, 1.14 wide, dark red, paired lateral depressions; median plates of all with long setae and few tubercles; lateral lobes tergites wider than long. Pygidium with distinct protuberant (fig. 38). Chelicerae missing but as notch in posterior dorsal margin of basal segment indicated by Beck and Schubart (1968) movable (fig. 42). Palpi missing but indicated by Beck and finger slightly widened transversely, armed with Schubart (1968) as bearing a few tubercles on seven teeth increasing in length distally; fixed fin- trochanters only. Leg formula 2431. Legs dark ger armed with four teeth of which most distal is red proximally, lighter distally, with sparse short much the largest and most proximal is reduced to white setae and relatively few evenly scattered a low lobe. Sternal region with coxae I not meet- tubercles; femur I greatly expanded dorsally and ing tritosternum; coxae II meeting along their ventrally (fig. 41). 1977 PLATNICK AND SHADAB: CR YPTOCELL US 13

I II III IV Palp with short white setae, with few scattered tubercles on anterior more on es- Coxa C)50 0.96 0.83 0.81 0.47 legs, posterior legs, Trochanter I C).46 0.81 0.58 0.73 - pecially on femur III (fig. 47). Trochanter II 0.54 0.61 Femur 1.12 1.82 1.15 1.30 I II III IV Palp Patella 0.61 1.01 0.68 0.73 Coxa 0.70 1.28 1.08 0.81 0.43 Tibia 0.97 1.26 1.02 1.08 Trochanter I 0.86 0.92 0.50 0.65 0.54 Metatarsus 1.03 1.73 0.92 1.01 Trochanter II 0.79 0.86 0.36 Tarsus 0.49 1.58 0.88 0.99 Femur 1.40 2.02 1.62 1.62 0.86 Total 5.18 9.17 6.60 7.26 Patella 0.77 1.04 0.78 0.76 Tibia 0.96 1.80 0.94 0.97 1.40 Second legs not greatly widened; femur I about Metatarsus 1.11 1.80 1.08 1.08 two and one-half, femur II four times as long as Tarsus 0.50 1.72 1.37 1.10 0.14 wide. Tarsal claws thin, evenly curved. Sperma- thecae wide, short (fig. 40). Total 6.30 10.58 8.16 7.85 3.73 Male. Total length 5.08. Carapace 2.05 long, Legs with sexual modifications as in C. simonis 1.87 wide at back of leg II, where widest, dark except femur II prolateroventral knob absent and red around margins, lighter medially, with small apophyses on trochanters longer, more pointed translucent yellow areas at margins between legs (fig. 45). Second legs not noticeably widened; I and II, with scattered tubercles concentrated at femur I three times, femur II five times as long as sides and short white setae. Cucullus 0.79 long, wide. Tarsal claws thin, evenly curved. Copu- 1.15 wide, dark red proximally, grading to latory apparatus as in figures 46, 48-50. orange distally, with scattered tubercles concen- Material Examined. Only the types. trated distally; distal margin with long white Distribution. Known only from Amazonas, setae, slightly invaginated and depressed between Brazil. protuberant lateral lobes; proximal margin followed by deep depression (fig. 44). Left chelic- Cryptocellus peckorum, new species era: movable finger tremendously widened, Figures 54-57 sickle-shaped, armed with eight teeth of which two most proximal are greatly reduced and Type. Male holotype from Berlese sample of others increase in length distally; fixed finger forest litter at Leticia, Amazonas, Colombia armed with five teeth of which most distal is (February 24-28, 1974; S. and J. Peck), depos- much the largest. Sternal region with coxae I not ited in MCZ. meeting tritosternum; coxae II meeting along Etymology. The specific name is a patronym their entire length, their suture line about one- in honor of the collectors of the type. third longer than that of coxae III; coxae IV Diagnosis. Females of C. peckorum are un- meeting anteriorly. Abdomen 3.25 long, 2.29 known; the moderately expanded metatarsus III wide near middle of tergite 12, where widest; (fig. 54) is diagnostic of the males, which may coloration as in carapace except for orange artic- also be distinguished from those of C. simonis, C. ular membranes; tergites with scattered tubercles becki, and C. whitticki by the lack of apophyses (fig. 43) and short white setae; median plates of on trochanters III and IV, from those of C. mag- tergites 11-13 raised medially between paired nus and C. glenoides by the thin, curved acces- lateral depressions, corresponding sternites with sory piece of the male tarsal process (fig. 55), depressions and scattered posterior tubercles; and from those of C. lampeli by the carapace median plates of all tergites wider than long. being almost completely covered with tubercles Pygidium with shallow notch in dorsal margin of (as in fig. 9). basal segment. Pedipalp coxae light red, other Female. Unknown. segments orange; coxae and trochanters with few Male. Total length 3.58. Carapace 1.31 long, scattered tubercles. Leg formula 2341. Legs light 1.44 wide near middle of leg III, where widest, red proximally, lighter distally, sparsely coated dark red, with small white translucent areas at 14 AMERICAN MUSEUM NOVITATES NO. 2633

FIGS. 51-57. 51-53. Spermathecae, posterior views. 51. Cryptocellus pseudocellatus. 52. C. albosquamatus. 53. C. lampeli. 54-57. C. peckorum, male. 54. Leg III, anterior view. 55. Tarsal process, anterior view. 56. Tarsal process, posterior view. 57. Leg III, posterior view. margins between legs I and II; surface almost bercles. Leg formula 2341. Legs dark red proxi- completely covered with tubercles, with few mally, grading to orange distally, with numerous setae. Cucullus 0.56 long, 0.81 wide, light red, scattered tubercles and long white setae. with scattered tubercles near distal margin and short setae; lateral lobes only slightly protuber- I II III IV Palp ant; depression below proximal margin lacking. Coxa 0.47 0.83 0.61 0.54 0.34 Left chelicera: movable finger laterally thick but Trochanter I 0.32 0.54 0.47 0.50 0.36 not transversely widened, armed with seven teeth Trochanter II 0.49 0.43 0.22 increasing in length distally; fixed finger armed Femur 0.79 1.33 0.95 1.08 0.61 with four teeth of which most distal is much the Patella 0.50 0.83 0.59 0.58 largest. Sternal region with coxae I not meeting Tibia 0.73 1.03 0.71 0.74 0.92 tritosternum; coxae II meeting along their poste- Metatarsus 0.79 1.15 0.74 0.72 rior two-thirds, their suture line almost as long as Tarsus 0.29 1.19 0.90 0.63 0.13 that of coxae III; coxae IV meeting anteriorly. Total 3.89 6.90 5.46 5.22 2.58 Abdomen 2.23 long, 1.69 wide near middle of tergite 12, where widest; coloration as in cara- Legs without sexual modifications. Second legs pace except for white articular membranes; ter- noticeably widened; femur I two and one-half, gites almost completely covered with tubercles, femur II three and one-half times as long as wide. with few setae; median plates of tergites 11-13 Tarsal claws thin, evenly curved. Copulatory ap- with lateral depressions and median elevations, paratus as in figures 54-57. corresponding sternites with numerous tubercles Material Examined. One protonymph, one and setae; median plates of all tergites wider than deutonymph, one tritonymph female, and two long. Pygidium with notch in dorsal margin of adult males taken with the holotype (MCZ) and basal segment. Pedipalp coxae light red, other one protonymph taken in a Berlese sample of segments orange; coxae and trochanters with tu- tropical forest litter 18 km. north of the type 1977 PLATNICK AND SHADAB: CR YPTOCELL US 15 locality (same dates, S. Peck; FMNH) probably portion of carapace width posteriorly, rows each belonging to this species. with from one to seven pits; four or five in longi- Distribution. Known only from Amazonas, tudinal row along anterior lateral margins. Cucul- Colombia. lus 0.97 long, 1.47 wide, light red, with long white setae and four transverse rows of about 10 Cryptocellus pseudocellatus Roewer tubercles each distally, without pits, with moder- Figure 51 ately protuberant lateral lobes. Left chelicera: Cryptocellus pseudocellatus Roewer, 1952, p. movable finger laterally thin, slightly widened 39, figs. 4, 4a-4c (female holotype from Santa transversely, armed with seven teeth of which Rosa, Rio Chinchipe, Cajamarca, Peru, in most proximal is greatly enlarged; fixed finger NMS, examined). Beck and Schubart, 1968, p. armed with six teeth of which most proximal is 77, figs. 5, 12, 17, 22. shortest and most distal longest. Sternal region with coxae I meeting tritosternum with point Diagnosis. Males of C. pseudocellatus are un- rather than median edge; coxae II meeting along known; females may be distinguished from those their posterior five-sixths, their suture line more of C. magnus (the only other species with deep than twice as long as that of coxae III; coxae IV cuticular pits on the carapace) by the invaginated meeting anteriorly. Abdomen 3.85 long, 2.95 dorsal margin of the spermathecae (fig. 51). wide near middle of tergite 12, where widest, Female. Total length 5.87. Carapace 1.91 coloration and setation as in carapace except for long, 1.90 wide between legs II and III, where brownish orange articular membranes, with tu- widest, dark red posteriorly, lighter anteriorly, bercles along anterior margins of tergite and with translucent yellow areas at margins between sternite 10 and deep pits arranged as follows: legs I and II; surface with short white setae, tergite 10, median plate none, lateral plates one longer at margins, with tubercles in transverse at middle of inner margin; tergite 11, median row along posterior margin and deep pits ar- plate 10 along anterior margin, 11 along poste- ranged as follows: median longitudinal row of rior margin, median longitudinal row of three, about six commencing at level of translucent lateral plates five along inner margin, two along areas, becoming deeper posteriorly but not reach- posterior margin; tergite 12, median plate eight ing posterior margin; about 10 procurved rows along anterior margin, 10 along posterior margin, arranged concentrically, occupying increasing median longitudinal row of three, lateral plates five along inner margin, two along posterior margin; tergite 13, median plate seven along anterior margin, numerous scattered over posterior half, lateral plate about six along inner margin; median plates of tergites 11-13 with paired lateral depressions, corresponding sternites with similar depressions and lateral longitudinal rows of four, three, and two deep pits, respectively; median plate of all tergites wider than long. Pygidium without notch in posterior dorsal margin of basal segment. Pedipalp very light red; coxae and trochanters with scattered tubercles; coxae each with two thick white setae posteriorly along inner margin; tibia with two rows of ventral tu- FIG. 58. Map of northern South America, bercles at distal end. Leg formula 2431. Leg light showing known localities of Cryptocellus magnus (1), C. glenoides (2), C. pseudocellatus (3), C. red, lightest distally, coated with long white peckorum (4), C. lampeli (5), C. albosquamatus setae; segments with relatively few scattered (6), C. whitticki (7), C. becki (8), and C. simonis tubercles except under metatarsi II, where nu- (9). merous. 16 AMERICAN MUSEUM NOVITATES NO. 2633

I II III IV Palp Those authors quite properly regard genitalic Coxa 0.83 1.11 0.95 0.88 0.40 characters as being of great significance, and Trochanter I 0.49 0.72 0.50 0.50 0.36 justify the establishment of Heteroricinoides Trochanter II 0.59 0.58 0.40 primarily on features of the female spermathe- Femur 1.15 1.98 1.24 1.44 0.99 cae. They note that the spermathecae of H. bor- Patella 0.71 1.00 0.78 0.72 doni differ from those of C. cubanicus and the Tibia 0.86 1.48 0.95 1.04 1.58 three Mexican species illustrated by Brignoli Metatarsus 1.01 1.55 0.99 1.08 (1974) in being globose and undivided rather Tarsus 0.50 1.69 0.94 1.01 0.14 than tubular and bifurcated. As we have shown Total 5.55 9.53 6.94 7.25 3.87 above and elsewhere (Platnick and Shadab, 1976), however, all known South American fe- Second legs not greatly widened; femur I slightly males, including those of the type species of Cryptocellus, resemble H. bordoni in having more than three times, femur II almost four globose and undivided spermathecae. Thus, even times as long as wide. Tarsal claws thin, evenly curved. Spermathecae with invaginated dorsal if one accepts the structure of the spermathecae margin (fig. 51). as a valid generic character, H. bordoni would Male. Unknown. belong to the group including the type species of Material Examined. Only the holotype. Cryptocellus and it is the other group, including Distribution. Known only from Cajamarca, C cubanicus and the Mexican species, that would Peru. require a new generic name. We therefore consider Heteronicinoides to be a synonym of Cryptocellus (NEW SYNONYMY). SUPPLEMENT Cryptocellus bordoni (Dumitrescu and Juvara- After the manuscript of the present paper had Bals) will key out to couplet three in the key been set in type, a paper by Dumitrescu and given above, but may be distinguished from both Juvara-Bals (1977) appeared in which a new C. magnus and C. pseudocellatus by the presence genus and species of ricinuleid (Heteroricinoides of a deep ventral notch in the posterior border of bordoni) are described on the basis of one male the basal pygidial segment and by details of the and three female specimens collected in the genitalia (Dumitrescu and Juvara-Bals, 1977, figs. Cueva de Cerro Verde, Rio Guasare, Maracaibo, ID, 6, and 9). The ventral pygidial notch was Estado Zulia, Venezuela. Those authors present claimed as unique to C. bordoni by its describers; an extremely detailed description, and we agree it is most strongly developed in that species with their conclusions that the Venezuelan speci- but does occur as a shallower invagination in mens represent a new species most closely related other Neotropical species (figs. 14, 20). Other to Cryptocellus magnus Ewing (and C manni characters of C. bordoni cited by Dumitrescu and Ewing, our synonymy of which with C magnus Juvara-Bals are also shared with various species of they were unaware). However, it is our judgment Cryptocellus: the enlarged cylindrical processes that both the lack of sufficient comparative of the male copulatory apparatus with both C. material (their having seen specimens of only one magnus and C glenoides, and the elongated tu- other species of Cryptocellus, C cubanicus bercles on the distal half of the palpal tibia and Dumitrescu and Juvara-Bals, 1973) and the un- the relative lengths of the tarsomeres of leg II critical acceptance of erroneous statements and with C magnus. poor descriptions in the literature (such as The presence of deep cuticular pits in C. Ewing's mistaken indication that C. magnus has bordoni, C. magnus, and C. pseudocellatus indi- no tubercles on the carapace, Roewer's inade- cates that these three species probably form a quate description of the cuticular pits of C monophyletic group, and the presence of ele- pseudocellatus, and Ewing's failure to note the vated tubercles on the distal half of the palpal deutonymph rather than tritonymph tarsomere tibia in C. magnus and C. bordoni indicates that formula of C manni) have led Dumitrescu and they are probably sister species (which is con- Juvara-Bals to an erroneous conclusion in their sistent with their close geographical proximity). erection of a new genus for H. bordoni. Thus, one could consider the latter two species, 1977 PLATNICK AND SHADAB: CR YPTOCELL US 17 all three species, or (if the arguments presented nida-Ricinulei). Premier representant de in the Introduction about the relationships of la fam. Ricinuleidae de Cuba. In C. glenoides are correct) these three species plus Resultats des Expeditions Biospeo- C. glenoides as the only members of a valid logiques Cubana-Roumaines a Cuba. genus. However, such a decision might (and Bucharest, pp. 259-278, figs. 1-9. probably would) result in a paraphyletic group 1977. Position systematique de Heterorici- noides bordoni n.g. n.sp. dans la famille containing the remaining species of Cryptocel- Ricinuleididae (Arachnida). Bol. Soc. lus, since it has not yet been demonstrated that Venezolana Espeleologia, vol. 7, pp. those four species singly or together constitute 147-180, figs. 1-13. the sister group of all other Cryptocellus. Thus, Hansen, H. J., and W. Sorensen until a phylogeny of the species groups of 1904. On two orders of Arachnida. Cam- Cryptocellus can be established on the basis of bridge, 182 pp., 9 pls. shared derived characters, we regard attempts to Kraus, Otto split the genus as premature. It should also be 1976. Zur phylogenetischen Stellung und pointed out that if Dumitrescu and Juvara-Bals's Evolution der Chelicerata. Ent. Ger- manica, vol. 3, pp. 1-12, figs. 1-8. indication that the male copulatory apparatus of Legg, Gerald Ricinoides feae lacks the circular loop of the 1976. The external morphology of a new spe- "filament chitineux" of the male tarsal process cies of ricinuleid (Arachnida) from proves to apply also to the other known species Sierra Leone. Zool. Jour. Linnean Soc., of Ricinoides, the presence of the loop may vol. 59, pp. 1-58, figs. 1-43, pls. 1-6. prove to be (along with the tarsal invaginations Pittard, Kay, and Robert W. Mitchell mentioned in the Introduction) a synapomorphy 1972. Comparative morphology of the life for Cryptocellus. stages of Cryptocellus pelaezi (Arach- nida, Ricinulei). Graduate Studies LITERATURE CITED Texas Tech Univ., no. 1, pp. 1-77, figs. 1-130. Bates, Henry Walter Platnick, Norman I., and Mohammad U. Shadab 1892. The naturalist on the river Amazons, re- 1976. On Colombian Cryptocellus (Arach- print of the unabridged edition with a nida, Ricinulei). Amer. Mus. Novitates, memoir of the author by Edward no. 2605, pp. 1-8, figs. 1-17. Clodd. London, 395 pp. Roewer, Carl F. Beck, Ludwig, and Herbert Schubart 1952. Neotropische Arachnida Arthrogasta, 1968. Revision der Gattung Cryptocellus zumeist aus Peru. Senckenbergiana, vol. Westwood 1874 (Arachnida: Ricinulei). 33, pp. 37-58, figs. 1-26. Senckenbergiana Biol., vol. 49, pp. Suzuki, Seisho 67-78, figs. 1-22. 1976. Report on a collection of opilionids Brignoli, Paolo M. from Pasoh Forest Reserve, West Ma- 1974. On some Ricinulei of Mexico with laysia. Nature and Life in Southeast notes of the morphology of the female Asia, vol. 4, pp. 9-38, figs. 1-13. genital apparatus (Arachnida, Ricinu- Westwood, J. 0. lei). Accad. Naz. Lincei Quaderno, no. 1874. Thesaurus Entomologicus Oxoniensis. 171, pp. 153-174, figs. 1-3. Oxford, 205 pp., 40 pls. Cooke, John A. L., and Mohammad U. Shadab Whittick, R. J. 1973. New and little known ricinuleids of the 1938. On a second specimen of Cryptocellus genus Cryptocellus (Arachnida, Ricinu- simonis Hans. & Sqr. [Arachnida]. lei). Amer. Mus. Novitates, no. 2530, Ann. Mag. Nat. Hist., ser. 11, vol. 2, pp. pp. 1-25, figs. 1-39. 479-480, 1 fig. Dumitrescu, Margareta, and Ilinca Juvara-Bals 1973. Cryptocellus cubanicus n.sp. (Arach-

3 ",--.III....I I..I.I.-.,",-....e- .,. ",-I-I.,,I... ..; ...i, " ...", ..IIII1., ..I .n .II I:'I.,I.r:% "I,II ,,., .It.., ..I.,,4.-... ,..I-,...,...,".I N,..I I-II,,,,11":I,..I .-11I, . .-..-.-,.."..1. -.. I1,-IiI...... -.II.1,'..IIII..II .:... .I..,-..I I-III.iII .,.I.,. :.II .I..IIf ,:..".,.,-I-,..12i-wII. ..I..-.I.III,.-I.I.,...1,IIII-.A .I.I.I,I...I...... -.11Ij..'!,.II.ll-,I....I..I..,.:.II..,..I,..-.II". .-I.IIIII--I.I1.II:,-.III--....-...':I..,II.-,I1 .;-,..I-.I1:II:,I .I...I-.. ...II;.,I. ..III... ,,.."I ..II.I.I .II I;IIIIIiI-I...: ". !,-r.-.;..II.,- .-.I..1 .I-.-... ..II;,., -iI,I2, ....I .-.:.III,,.II-IIIo.III..,-I,...:",.,....I..I. ;-.-,III..- ,-.1. ..LI-I.II.-. I:t...I.I.II-11..II:,II.I.II,!....";.II ,1.I .zII...II.I -.....I.1.]. ..II,, '. 1.,..I.II,. ..: ,-LI.II. TIII.. I;I--1I .II.I.IIII.II.I..I....II.,I1, ,I,,.. II.I..I..I-11,,.. .I-I,..-..I.-III v.,IIII..::,..,:--,I.I... III.I-.I.II.,II...I..o IIII-II.:.II o, II.II"... ..II-. II.I..."I...,II.I, -.IIII.-..II..,- .I..,I...;I..;.I..17.I..I",I.I:,-I.I.,,,. ....-.II.... ;:....1I;I..,zI.,..6I,11q,,I.-I I.III.IIIIII..II. I.-II. :I ,. ..I.II1. .-,., ..1...,;.1I.IIIII..I.I.; .11I.I:--.III I.wII.....-,.;,I...,I-"."'- ,..,... IIII... .. I, ..I.- ..,.I I,.I.I.II-,,,"I,..I..'.,,,-.II-.II..III..1,..A.,II.-Itc:,... .4..I..I. .eI;. ,i.iI,-.;II.:I. .; ,-.,.41..t.,.I:....: ..-I.II.,..I-.",.-.,I. -...,II.,..II..I .11..,.....;I....1,..,;I-I. .- .11:L. ..I.II ..IIII..I,i,.,...... ,-..rl.. ,I.I. ...I.. ;,:...,. .,i"I..I I? 'i .--I..I., ",.. .II.I I.I.-,....IIII..I..II.,.III..1I,.II,,.%I..,III .iI.., I..- :.II -.r, I.-.,I11,-..III---11,..II.. ,1, ,II-,-II.,I.!..I.I.-II. I..II.II..11 1.,I...."I.1, ,,.I,."II,tI1,.,I,....i. ..II-II. III.,.-I. .4, I...... I.I.II.-7 :-,9IIwII,7ll..I..,:II...f,e...... ;":II.- I..I, .. ,;I.I ,:II..4,.. .I.,.I-I 1,.-.III.IIpII...-,.....III.I.,.I:,,..'mi...., t,AI,.,:....I..III,I,I.-,...,,..,.:...-i. ...I...I.i";. 'v".. .II;., elI...,II. .I.. ...III.,....III.I.I.II.II.IIII.. .II.III..,.I.,:I.1.;.I.I:li,I;F 1,-,"I.I .I....,.III'.-. I.,I.I-II.I.I:1...:,,.I..,.:I r...:.I III..I..,I,II..: iIIIIi-.III..---:..I.I.;.II..I I. ...I. I.II .I.;.,I-."-..I,.,7, .,.....;-III-. I...-,:,II...,III.I.I.,.rIII.,II III.I.I.,,.v.-I,I.II..I,,.:I..;:."I..,,14:.- ,,I ..-I;.11....II,II.IIIIII..I.1...l-.Ili ,.I III.,,..,,I.1,I..;,...II-IIII.lI..,;.:1. -,,...,.-1;I,A I...III.I .:.,!,I.I;....II.IIII I.I...III "..",. .III-,..1I.-I..I-I,..;.,`.II..I.II.II." ,.,.,::....I.II-I. I.,IiI.,I.IIII.III..IIII I,I,,...II...."I...I.I I.II.,II I."I., .,IIII,I;::,:. ,I.I.II.II..I II ,-I:,.: .,,.II ,.II,-.I.I.,.II.I I...r....I.,..I..I.....1..,...,-.I,.I,.II....III.., ,...,,.I.,,..Ij.,,.11.-I,--,:,,,.-,,:. II..I.IIIiI'.,,II..,...I.I ., "I.. II%.I..,-4.:I...I .II-..I..,...,..II...-.,,I1..... :..,...I.... II-.. ..-iI-...... II.,I..I,I.I-i ;,I.,I-:. :,.Iq.,. .,I.-".-I.I,.;....k. ,I.I-...IIII,I.....,. ..I.,..4"- ;I:.I II. ..II., -II.."..II,,., ,..:.IiIII"i.:..I:I I".I.II..IItI...... I.I.1-I.:1.I,..I ,,..,,,I,I. --..i. .:-.I. ..I.411 .....i.I.IIII ,;I-.II.II,...,I -.12... ..I..I ...I..,;.'.,,.,,..,. .,-.I...... ,I..IiI:I:I,.-I-, ..III... ..;..II-II..1I.I....zIIII"-.,,,i-...I..I:.:! ..I.II-,I.II.I;...,,.....,I.. .I11.11 :I .,I.,I..".I'..IAIII.I. IIIII I. :I.I;..,.,,. -1III ..., ,1 j.:'.....- .1-.11 c. ... II.....J...I... III I.II.II..I"-,.III,I.I.II'.,, .i;.i ,..II.. .j.,'I .-II-.II.,:...... " -I'.w,.., 11.;II::.....I..I.I-..I1e .I..I.I.I..I;.... 1.I....IIIII.I.:I.-1..I.:-11 ..--:-,.I I.....,.,.: .I.I....I I..I ..I.. ::,- .:.'I,...V-.I...,. I.;.,.., 7,... tIII-.,.I.:II..III..:,.r,I..,..-,-I,.l.I..I..,,IIr-I.]I.jIJ.. 1....-. .i..-.iI.-.I..I.,.I.III0:...... I II,.,. I.III- ,I :I,:,. ,I-,,,.. :.!- :,-..'.,1..I,:I..I. IIItII .II. I.1,,,-...:,..",6.;.,.,,.,... ,.:.,I....I..,,. I.:.,I... 1...,...... I...":.II Im- .. 7.,..: ..I-.:I.. ;II.II ,.....i-IrII-, .I,.:,",II..-1..II.:,,. -I -.II .,I.II.I...I '.II I.II...IciI...... I..IIIII.j...I-II,w.i ...,::,..-,..I...11..y.',7-.I.-I...:r-,,.:.--I.I,,II..I.II.-II i.,.-.II,I-..I- -I-.-.I..I..I.%..,..-.....,. ..1..p.l.rIIII.i.-.-1...I II.I.i,I.II..II.. ,.I11,-III ..I.I; II....,.w, .I11 ..I. i;." :I .Ii.I.I...IliIIII,;.,:.-I.-..;cI, ..I.,...,1..,.I11..,.I,....I.-. .,.I::.,-... ..III.. .II.II.IIII,.. -.I.:;,..jI.II.I..II ,.I.-i-I,..I.;.,..p.I.I...l..Ii-% .I .I...j,.. ,..I.wI11I. .I, .,.;. ..11.IIet, .- .I.- ..I ..I.;I..I..1;I,.`.I., ',.I-I .....I-I.IIII,.II.'; ...IIi.I:.-lll..I..I..,.II.II:.,,.,,..I. .I,., ,.,I.1%.:I.III. .II.II...I-II.I...p.II::.,...: ,I..-.:., .,IIII.:.,-I.) .-,. ;.,... .III,....- .. ..;,v -.; ,.".I.I .I I. IqI..II.,..I..,.,II.;.I.i...;- I-I.II:..I.I.,1.I.....,I,-I. mI.... II,,.II.,I.11.I.I,.-,.,,.,,...IIIII.I,...I.., .....II.,,-:.... III.I-I .I.,T.-,"i-,.-,..I.-I.".I..I I.,II.,.,,..I;II..I-1ro,.w;.,%I.-,;...,.4.I.r.-:.-II..i%,. I.III..LI...q.i.I...II...1..I.,I:..II ..I..1;I..-...III.,..II,.....:..,.-,.::.IjI.I.I II..II-...,j.,.II.,.;.,,",-I.I...I.i,..; "I-;... :.IIII ,IIqII. ...I.,.,-..::,.1. ;.;.II...". r.-?I.I:IT.: .. I..III II.I.I., .I.I-I. ;,.-..II,...... I .I.;IIj....,;..II-.,,.I-11,..I..,..,... ..:.,.:.I.I.,.,,I....;.I..I.II.,. ,.,I, ..III.I. II;..I.I..:,.,,,:.7,II.I-.....II-...1..-"I.,.I...... II ,.,.I ...I..II..i,".IIt.,.--.I.:"-.: ,,-I...I,...I. I..-IIII II..I.....:,Ip..I1.."1I,...;.Iq.,1....-1..",I,.I..i...,.I..:III...,.,-,II....I..I.... :.,. ,:-,`.II.I1I,,IAI,. .I.I,,. I..I..I ...6 ,I,l.....I I1.I.II11 .I: .,",I.I....1. ,-I. I.I.II...I.--.'...... ,,...4..II.1..I.,..:.,...'.-,.I-.-,.,-..:.",.,.,:...III1....I.. III.II-III:..,I..I.I.I. ,.I..II..II.I'., ,. .,.....-,.--.w I1. :.-,:i.I1.- .IIII-III.::.t.I.14111. I.I..I...,.. 11-III.I.i 1.I.-.1.1 i",..,. ".I..- .:,.., ,III,:. ,I..II I.,-.III-I-.. ..II...II :: I.IV..,.I..... II..- ..II%.I...II.I:;i,.,..F,I,I.-I;..II.-I,I,IIII.1II1. -.....-...1.II .II." .11.I.- -III:.,11..III,.,.,I.,,-11;II.I.I...III.,-,.III.-..I. %, .II.II.,I."r.-:.,.III.I.4 I?,IIIII..I..I.I,,.,Ii...I." ...-1.1,I.1.II. -II.II.I..I.IIII,..IIiI.I-.,:..I.-..,-..I;.2,..I.: I.1, .I.,-,,..I...... IIIII.III... ..I..I-...L.II.. ,i.II.I I.-I .;....:,,IIII....,j .II..,..:.I I,I.I.II....;II III..41::.I.II..7.r.".,-II.1.I1,III,.. .'...1..J-1....!% ..--II. I7I,, :.....,I.II--41i,I..1 .I,.II-.I..III..,.,I.I..-I.II..I.I..I.I,....I..:i.,I..I.I.I..I.11 IIIIII.-I.I,III..II.1, I....I.."I ,....-. II.II...III-III.II-III.... ,ii.,...;.7,:1.,.. I41 .,-.;1;..II..I.,.,,I .IIIII".I.:.-,I...-.,...,1... .vwI,,,.I. ,.;. .. .I.II..I.II. .II.II,;I.--".. ."-I .II- ,.,,i..r,Iw.1.,7.III.,I.-II.I..III .I..I.I.II.-.III.IIII...,,:,.--I:;I..1.II.IIIII-,.I,. 7. .:.I.II.. IIII.1.I.I-I-.II.I.%.-.17.I.,.:I ..II;Ill.II I.; I...I.I.II-,.I,.II....1.:..'...... II,.I,.:...."I.-,.I.III,I:III1,I"II-I1,.Iq,,;"I5'.1,1,I. .II.--;..I.I..i.I..I.."II.I.....II.,,,.II.I.... .I.I.. I. .II..:.1...: ,--..-..I II;... .IIII.IIIII .;-IIIII-...I..I...I..IIr,.II.,.II .-..I.,t..I...I.4.IIIII.I..I.-I-I. r.iIIII.....:II ;.I--i.. II.'.... I-I-. iIII I.I-I...... ;II .I..1.:I.I.-I I-..IIIII.I,-..-II..,..I,-,:.III..I.4.1.-. -II... .I.:III: .I.I...I,: ..II..I. II.II.".IIeIII.I-.-I.Ili..',..i..I.Io, ..-..II.I.I...I .1..I.IIII.III.I..;I, ;I7..c,.,:1,, -., :.I .lIII.....I I.II;III.I:...... II..-.I.II- I-II.III.III.I ..I..II... -..,,,III.I..IIo ..;I.-.,..,I..II,... ., .-, -.... II".,.III-.....III-.I-I.--1."w... I.III.'..I.. I,..I.....-..I.iI.....II.I,I..I::; II;IIII....I,.III,..I..;...,...... '..! ...-,I. -.,.I. 11.- ..-- .--,....I .-.II,.,;..".II.:,...... I.II...;-.-I,.,..I.I1. .-I.-,..-. ,,,I."I...I III...-III.I.-,,I""IIIIII.-,I.I.IIIiII..I.,.II).,.III.I.I.II.Iil-I.I..II..,11 ,.... ."I, qIII.I-:I..:II..-.1,.1,I.II.I. --.I-...... :II. .-I- I..I..l.:...... III.,..,1.III .,.'...,I...., II.:,.I..:..IIIIII.I..1..L-;.IIIII Ir..,I-0 -,,III. ...;.,.I-...I,,.,.I. -..II-:, i,:..I.,.....II ....I...;.,..I.;.",I., ,,,III.....,I...... iI.....-I:I-.-.-6.. .-,I. .II...,.,.-....I: .-. :...IIII.., .-.. ,--II.I.. .I..,I..,..:.I'I,...mI.I.....r.I .,,:'....II,.1-.1z,,".II.,..IT..,I., .. I..II I.I...I:.,,...,II..I.;I. I.. ,,.....iI II .,I.... ,., :,,, ,,"IIIII .l "I, .,I,.:.4-1.:,.LI....,I..I.I I.qII.I.iI.%.d.....II..;-...,III11..IIII,.II, ,.,-I.I%.II.IImw.,-,.I..IIII.I:II.I II.I... .I.I.II.,:.-.I...... ,.,"...,.,..-I...,.-...,,..II.,-.-I-.:I..,II.!.ImI,;. ,....' ..I.I.III."I..II. I.II.:I. ,I1,.I..I..,.II-...': ...I",,, .III.:....,.,.,t II.I, ...I,.-,...II.I.., ..I,.;..-....:.,.':II0-.': .,' .....;.II. .,,.,.. .,.- -., -I I.-I. .. ..I.....III".11-I..1.I...II..II. I...114..-,.I.r,.-,,I.I..w..III.III.,a:.I II,. .II.II.I-,I.,...,,I...; I-- ':...., I.,.II..I.I.I. II:I11.,.-.--,,.I1i...,l...... ",,I.-1I....p1171.,....., .,.11-I:.I I.,:II-IIII...,..-.....-.!.,.1,....I".-.,;,..I.,.;..,....II..IIiI..-,I..m:I:,.,_iI.IIIII.I .-IIII.I .IIIII.- .;I'A-,-iI.III.II.II.,..;...I.....i,.:. 1,!.1.I..I-II -II...I.-Ii.I;.-,.I.I.,...,:,..III.I.,.. .7-.. ..I..1.I-..ll..1..,II...I'.II IIIIII..I.I.4:.,.,.,.: .II4III, .I.,.I-I..:IIid.I.. i-.,I,, I..1,I,I-.I.I I.II.,..III.....I.....--II,,,-II.II",,...j,,,.,I II,III.II .,:.,.., .- I.II.III. ..;:.-I-.r.,.,-i.-1.-I.I..,... Ii-., ,.III.I"-.II. .I..,iIII.IIII.7.-:I,:II...I-I.... ,..:",, ...,..II.I III.I-II.-m...,.,,.I,,.I.II-...,":;-:"-...III..II.,.III .u0..,;,..III... II....I. .,II.,,.I.II, II.I..I..I. !.,, ".".,..II..,. .. ,.,-..,..IIII.I. .III.I.-I.-II..i.. .IIIII i-.I..,1..I.,I:I...11..I...,.-..:1:,1--.1,,I.I-..-.,...,I"1.I. I.,1.-.,-: ,..I-i,.:..".I, ...... ,.... I.II.;...II..I.I.-..I,.,I--,-,..-II.I,. ..-. ,.riI...II,I.,.-,..I..;. I,:-1....::7...... ,-,, ,:. ...III.qIII...-II1.!I,.I--.%..I.I.,r..:II.I,I....':IIo.I..I:,;.I.,,.I.:.,.,.II,..I I1.I. ..., ,-"m.".,.,I.,lI,.I,I,.,,.-.;... .:1i 1-.I..II.:..I :IIII..... IqI."I.:-.I.I- I.IIIIIII.I11II.17...... ,,,,,Ii...o-.I.- -.II.II...I,:.IIIII II..I"I.41.I..I..-. ..I.I ,1,IIIjI -.-,. .I.. -III, ..:.III...... ,Iw.I1....I.Iy..,.,,I,".:1;. I.II..-...,. .`II.I,..,.,,r,.1 I.... III.:...- ..II- I--II...I.I-,-,.I..III,.....-1. .-.1..III.I..1.II-1..I.IIiIIII-1I.II,-I...-II ...---,IIII,--I.,. .II.I,I.I.:. .,.. ..II..,,....-.... .-:-.. ,,.,. .i.,..-.,-..I,. :,::11.: :,.....I.I.. ...-;,.,.I:I.I-.,I.III..I;-.,II..:;-p.-I...,...:--;...,..wII.,.III..,,.I. :. ..I....I...I...I..,I.."."I,,,:- %..-.-t.I--IIII III..r.I:.II..II.,I:.1..V..11..I,-..,...... i,...r;.1..11... I.:,.I ..II-,.I,-.I...... I.I.I..,..I..I..II I.I.II-.-I I...,-II..I. .II.,II...II..-...II, .,., .....I ....I..IjII....11.-. -., 4I.I.r....I.....I..iI..I..1..,.-I..I..II .-.I-.I...I. .I... .iII.II. I... ..,-,,,11.I--,11; ...... II.I. .-.I%...7"..I..Ic..I.q..I.:,.ZII.....-`... .,.I .., .-.:,I...... I.-.I-.:..-I.I._.I1,,.:1,..IIIII,.J-,I1.. Z,,:i",I;.."';-II.. .I,.1,III11,.:,,,..I.6..Ij ;.I.1I:..I-..I ..II....:IIII.I..I.iI.II.IIIIII.,.II.-.II..I.II., .....II ..I.II. II.I..,..II11,.w..:...II1."II...... I;,.1...I..l..I..,...1,.I.III ..;.1.,,., j..., I1.., I... ,..jI .-. ,,. .I: ,.I. ,-I.; .I...I I.I:4;....I:II"I".,II.,-.,.. ..-...1...-.,...I.:;.-I.--. ..-,a...Ii...i:. I..1P. .: .;,.. -:-,,.....I.., ,.:-.:'...... ,, ..,. .II..:.:,.II.,.,..,,...J."I-...... :I.,11.IV.I-.., I.I.;.....It....m...... ;,I.I.I-..t....1.II..4.. ...;,I,.-.,...I.:,.I-.I.,",...I;-..I'.I.,I.,w..I.,I...IIi IIIIII.,.II...I:-.,..i.I.11I.IAI:.-.",...... ,I...II-:.,'. :. ,.-,.I-, I,, ,11I.....-. I.I..,, I,%..,-..., ,.IIII i..II.; III.,,,-..I, ,.....,I.III.,,...,...I,I,i,I,.,d...I ,...-,,-... q.. II. -I.1I...,I..,."," ...,.,I.I,.II...---..I.-..-.I II-...I. ...I .- ...I ..I.." I.II--I.,I.. ..II..,,I?.-I.11..I.-..III,...III::1.. 7...III I..-....1, 1,....IL.I....,IIi.. .I.I.I...... I0I.-;.I...17.-II -;Ic...It. I. ,,I ..II....I. ,.:,...I.--.. III.. I,.,...-I.!,:. .I. -,.I, -I:..II.I..Ie .I.. rlI...1411 ... I.II-II,.I...,-II.,,I.I1,.I....I.II.I.;I.....,-:I. I...,. -" .1. ..,.1I.- ,I...... '..;...,III...I,I:.1.IIIII..II. .-...-I I.....-.!,..,. .IWII..,II1;.I ,.,,... ;a;.IIIII...,II.I.,I..4..I-.,....I...I...I-I...I..I.-."..I., ..,II...... I.I.:.,.I-II...o,.i....1;...... : ... I.I.III.IrI...I. II.. .. i,,-, .II..:, -,II.I..,..':I .:I.: ,s"7.. --.:.; I."I ,..I.II.-..I.I.;..1.I.....I...I.;"..:,:I ..;..,..;....II.I ...., .IiI1. ..I.I.I -.. .II.I.I,I.I-I.1.I..II.,,"...... -.II. I..., .I,,. 11.I..,I .I,,.....,..,I.I...I..- .... 11...,.I..-,I...I.. ,:,,. .. .I...... -.III.:-.:,...,.4g.II.11,.I.:.-., ....I,... r.,.III-j.II..-. ..III.,q..II.-....Iv,,,....IIII....II.,.,1...,I -.III,III..I I...,,.,.III..III III...... ,,...,., ..I..II I-;:.I.::,-.-I., ,.. I-.III. ,II. I.III.I....II.I.,..I.I .I... ".-I.I..l.,:.,.II.IIm.,. -I,.I.I -...I..-.. I1. ..I-...... I....-..II.II.:,,I..i...:,.. .;.. .I11.II.e,.IIII. .I.....;:.II.-,:,..,:II1.II... .. -..,,I.II ..I..1 :11, .I..I.., ,.III- .-.-,;I.:..I..%I..,..,zI--I .I,II..II.II-.;I.I.I. .1i..II.I...:. :I ..i..I..II. II.I...II.-.,,....-IIII.,..-..I,I.II.I.i .. ,.II...I.I.I..II .p.II-...11I,,..I... ,. I;.II,II:.r,.... I..1..I.:,. ..-IIII,o-... I.II.II.-...'..I.-.I,,II,IrII..,,,.I.I1. .,.,II" ,II-,.,..',I ..,.;I I..III,-oI.I.I...:.....,.1I-I.,;1.,:..1I,,,'.' :1, I. .jI.III,b.,III: ..I11l..I... ..:., ...... ,. ,. i,.,. ,-.', .,. F.,., I%II.,.,I-I.,.II..III IIII,I1.I.II."..,--..I14::. ,,.. ,.I.I,, .III. ...I.,.i.I.i.,I -I,,...I,"...,:.., ., ,.,II-I..,I.I. Im;..I.,.I;.I..,..I...-...I.I,.I.].I;I...I. II.,...I..I.].2,--II,.I,,,,.r-,,;.:.I.II. ,.- .,. .,... "II11I:.I... I11.,.I..'t,r,.I ..I..I,:..I.:r...;.I..1.L.1 I.IIili -,, ':.I,:..I.l.III,...... II ..I .-..III.11o;'d.IIg,..,;..,.. 4..:..,.I ;I .II..III.7,r-,..I-..II:I...I ..,.-1 ,...,".I,.I-..I.II...... I,I :..I..iI.III I.. ..I.;I..l..,.,.. `.,;..-,.. III.I..II...II..I.I.III....I,,I.I.I..I.. ,,, ...,II,II.....,. L-.I:,.,III-. .I...I..,.,?,.I" .I.-.-..I,.....I...-,. I.I :.,...--.I. .: .,'.zv....,I.--i,....,..I....,..-.... ;I.I.,II11,.:..,:..,.. :i....,....II.t.. ..IIII-I.II..II..."..".li...... II.;I-.:-I..,e"I,....I-I.:.II:.,4 111.. II"I..,,-:.".1-.III1;.,.-.I14.I;.I. .,-.".:..-.II.1. .II II I.II.u-.w.-I':,I..IrIII.,.III..^...VII.I.-I..'III.1,.I,I.....II.,.;.,.,Ii..1,III-III.II II.I....I.,.I..I;-,.I.I.I.I..,II.,l ,..I..I..,..,", .,, ..II.-,,,...:,.z.I.,.....I..;. ,-.., ,, ..I..II.I .I.;II .I.: .'-. .I,.I.'..,.I.41.,I.-. ...11 .., .., -.: -,...,,.II I.I .I:,,..,,..,..II..I I...I I.wI.I,.,II4d.,.mi .:...,,I,,I III.;.I.....,I,;.-I.III..,...,. ,.I I..r..II.,.....I.':..I.IIII..IIII-.I,:..II I. ..:III.I1I.1.III.. II.I.I..,:I"III.II.-,-.I:.:,.m..,I ,..,,,,:.,...-Ir ..I.II.III...I ;,IIIII, ...I..I. IIII.I..Io.,.:.iIII, I.I..I.,I-I-...-,I. ..m.-..r.II.I...,- 4.I.,.:I..;..,,,-.%...I:I;.I.II.11I.I.II. ,I.,!. ,,,,II.I.I.,..,, ...,. .-.. I": ...,I II.....I...... -....;I...iI,...,-.,,-,. ...III.:;r,- .;, .1I.I