AP Biology The Origin and Early History of Life

1 Origin of Life Hypotheses

• Special creation

– Supernatural or divine origin

• Extraterrestrial origin

– Panspermia: Life from elsewhere in universe infected Earth - Testable

• Spontaneous origin

- Life originated from inanimate materials

- Testable

2 Reducing

• Life most likely emerged under high- temperature conditions.

– Early atmosphere is often referred to as a reducing atmosphere.

ƒ Exact conditions unknown

™ Ample availability of atoms (gain electrons = reduction)

™ Wouldn’t take as much energy to form C-rich molecules

™ Very little

3 Origin of Life - Location

• Ocean’s edge

– Bubble hypotheses

• Under frozen seas

– Problematic due to necessary conditions

– It was warm

• Deep in Earth’s crust

– Byproduct of volcanic activity

• Within clay

– Positively-charged clay

• Deep-sea vents

– Conditions suitable for Archaea

4 Miller-Urey Experiment

• Attempted to reproduce early reducing atmosphere and produce organic compounds from inorganic materials

– Hydrogen-rich

– Electrical discharge

ƒ Succeeded in producing amino acids

5 Miller-Urey Experiment

• 15% of C in CH4 converted into simple C compounds • C compounds combined to form formic acid, urea & amino acids (glycine, alanine)

6 Chemical Evolution

• Ongoing debate concerning actual path

– RNA World - Molecules could not have consistently formed without a mechanism of heredity.

– Protein World - Replication would be impossible without enzymes.

– Peptide-Nucleic Acid World - RNA is too unstable, thus a precursor must have existed.

7 Cell Origin Theories

• Bubble theories

– Certain molecules spontaneously form bubbles.

ƒ would serve to shield hydrophobic regions from contact with water

™ supports ocean’s edge scenario

8 Cell Origin Theories

• Oparin’s Bubble Hypothesis

– Primary - protobionts allowed chemical complexity to develop

ƒ Many different bubble scenarios have been proposed.

™ no agreement about composition or how the process occurred

9 Bubble Hypothesis

10 Earliest Cells

• Microfossils have been found in rocks as old as 2.5 billion years old.

– resemble prokaryotes

ƒ lack nucleus of more complex eukaryotes

11 Archaebacteria

• extreme-condition prokaryotes

– lack peptidoglycan in cell walls

ƒ methanogens

ƒ extreme halophiles

ƒ extreme thermophiles

– thought to have split from Bacteria 2 bya.

12 Bacteria

• second major group of prokaryotes

– strong cell walls

– simpler gene structure

– contains most modern prokaryotes

ƒ includes photosynthetic bacteria

™

13 First Eukaryotic Cells

• Eukaryotes probably arose about 1.5 bya.

– Internal membrane-bound structures such as mitochondria and chloroplasts are thought to have evolved via endosymbiosis.

ƒ Energy-producing bacteria were engulfed by larger bacteria.

™ beneficial symbiotic relationship

14 Endosymbiosis

15 First Eukaryotic Cells

• Sexual reproduction

– Eukaryotic cells can reproduce sexually, thus allowing for genetic recombination.

ƒ Genetic variation is the raw material necessary for evolution.

• Multicellularity

– arisen many times among eukaryotes

ƒ fosters specialization

16 Extraterrestrial Life

20 • Universe has 10 stars similar to our sun.

– Conditions may be such that life has evolved on other worlds in addition to our own.

ƒ ancient bacteria on .

™ largest moon of Jupiter, Europa, covered with ice.

¾ liquid water may be underneath

17 18