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ISSN 0373-580 X Bol. Soc. Argent. Bot. 41 (3-4): 327 - 330

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ISSN 0373-580 X Bol. Soc. Argent. Bot. 41 (3-4): 327 - 330 ISSN 0373-580 X Bol. Soc. Argent. Bot. 41 (3-4): 327 - 330. 2006 Nuevas citas de Pteridofitas para el distrito chaqueño Serrano de Córdoba (Argentina) RITA MORERO1,4, MELISA A. GIORGIS2, ELÍAS R. de la SOTA3 y GLORIA E. BARBOZA2 Summary: New records of ferns from the Chaqueño Serrano District of Córdoba (Argentina). In this contribution, three new taxa for the local fern-flora, are reported for the first time: Microgramma vacciniifolia (Langsd. & Fisch.) Copel. (Polypodiaceae), Phlebodium areolatum (Willd.) J. Sm. (Polypodiaceae) and Thelypteris dentata (Forssk.) E. P. St. John (Thelypteridaceae). Also morphological characters, habitat and geographic distribution are given. Key words: Ferns, Microgramma vacciniifolia, Phlebodium areolatum, Thelypteris dentata, new records, Córdoba. Resumen: En esta contribución se mencionan por primera vez, tres taxones para la flora de helechos de Córdoba, Microgramma vacciniifolia (Langsd. & Fisch.) Copel. (Polypodiaceae), Phlebodium areolatum (Willd.) J. Sm. (Polypodiaceae) y Thelypteris dentata (Forssk.) E. P. St. John (Thelypteridaceae). Además, se comentan sus caracteres morfológicos, hábitat y distribución geográfica. Palabras claves: Helechos, Microgramma vacciniifolia, Phlebodium areolatum, Thelypteris dentata, nuevas citas, Córdoba. Introducción mencionar entre los relevamientos florísticos realizados anteriormente, a Bianco & Cantero (1988), quienes citaron 46 entidades para el sector suroeste En el presente trabajo se dan a conocer tres y a Arana et al. (2004), que ampliaron el registro a un especies de Pteridofitas, no mencionadas total de 83 taxones específicos e infraespecíficos para anteriormente para la provincia de Córdoba. Los la provincia. ejemplares fueron hallados en distintos viajes de Es interesante destacar que los tres taxones fueron colección realizados al norte y oeste de dicha encontrados en el norte del distrito Chaqueño provincia y corresponden a los siguientes taxones: Serrano, perteneciente a la Provincia Biogeográfica Microgramma vacciniifolia (Langsd. & Fisch.) Chaqueña (Cabrera & Willink, 1980). Si bien esta Copel. (Polypodiaceae), Phlebodium areolatum región cuenta con estudios relativos a la composición (Willd.) J. Sm. (Polypodiaceae) y Thelypteris dentata florística de la vegetación (Kurtz, 1904; Sayago, 1969; (Forssk.) E.P. St. John (Thelypteridaceae). Zak & Cabido, 2002), sólo Kurtz (1904) hace mención Estos nuevos registros contribuyen a actualizar de algunas especies de helechos mientras que los la información relacionada con la diversidad de los restantes no abordan la flora pteridológica de la helechos de Córdoba y a documentar la extensión del misma. Dada la topografía y la diversidad bioclimática área de distribución de elementos neotropicales. Cabe de esta región (Sayago 1969), debería explorarse aún 1 Museo Botánico de Córdoba. Facultad de Ciencias Exactas, más, con el fin de obtener un listado actualizado de Físicas y Naturales. UNC. Vélez Sarsfield y Duarte Quirós, las distintas especies de helechos, lo que permitiría 5000 Córdoba. 2 Instituto Multidisciplinario de Biología Vegetal/ Universidad establecer posibles afinidades con las floras Nacional de Córdoba y CONICET. Casilla de Correo 495, pteridológicas de otras regiones biogeográficas de la 5000 Córdoba, Argentina. Argentina. 3 Facultad de Ciencias Naturales y Museo, Paseo del Bosque A continuación se presentan las tres nuevas citas, s/nº 1900 La Plata. CONICET, Avda. Rivadavia 1917, 1033 Buenos Aires. mediante una descripción de sus aspectos 4 e-mail: [email protected] morfológicos, haciendo especial referencia a las 327 Bol. Soc. Argent. Bot. 41 (3-4) 2006 características del indumento, venación y esporas, Mato (Myrcianthes cisplatensis) y epipétrico (fig. 1). que no habían sido consideradas en detalle en las Material examinado. ARGENTINA. Prov. floras existentes. Córdoba. Dpto. Sobremonte, en las proximidades de San Francisco del Chañar, 39º55’S 63º49’W, 573 m.s.m., 1. Microgramma vacciniifolia (Langsd. & Fisch.) 21/II/2006, Giorgis, Cabido, Cingolani & Gurvich 1044 Copel., Gen. fil. 185. 1947. (CORD). Polypodium vacciniifolium Langsd. & Fisch., Pl. Voy. Russes Monde 8, t. 7. 1810. Tipo: Brasil. Isla 2. Phlebodium areolatum (Willd.) J. Sm., J. Bot. Santa Catarina, Langsdorff 6/7 (Holotypus: LE!). (Hooker) 4: 59. 1841. Iconografía.- Sota de la, 1960: 66-67, figs. 8, 9. Polypodium areolatum Humb. & Bonpl. ex Willd., Descripción.- Plantas con rizomas dorsiventrales, Sp. pl. 5: 172. 1810. Tipo: Venezuela. Sucre, Cumaná, largamente rastreros, de 3-4 mm de diám., con escamas Humboldt & Bonpland s.n. (Holotypus B-W 19645- peltadas, adpresas, aovado-deltoides, con base 01 01). ensanchada y ápice filiforme, paucidentadas, 4-6 (10) Iconografía.- Sota de la, 1960: 79, fig. 11 (sub mm de long., discoloras o, concoloras y blanquecinas Phlebodium aureum var. areolatum (Willd.) Baker); en las zonas más jóvenes del rizoma. Frondes Sota de la, 1977: 142, fig. 57 (sub Phlebodium aureum subtrimórficas, con dos tipos de frondes estériles, (L.) J. Sm.) unas suborbiculares y otras oblongas y erectas, Descripción.- Plantas con rizomas robustos, ambas fotosintéticas, de 1-6 cm de long. por 8-18 mm rastreros, cubiertos de escamas castaño-rojizas, de lat. Láminas fértiles lineares, de 3,5-13 cm de long. peltadas, con la base redondeada, discolora y ápice por 4-7 mm de lat., predominando en la epidermis filiforme, márgenes paucidentados. Frondes abaxial escamas foliares semejantes a las del rizoma, distanciadas, de 15-55 (100) cm de largo. Pecíolos aunque más pequeñas, y otras de base reducida y castaño-oscuros, ½ de la longitud total del fronde, con 2-5 prolongaciones filiformes y tortuosas. glabros. Láminas de consistencia cartácea a Venación de las láminas estériles reticulada, subcoriácea, deltoideas, con base truncada, glabras, generalmente las aréolas adyacentes a la costa con verdes o glaucas, profundamente pinnatipartidas, una venilla libre inclusa y hacia el margen aréolas senos redondeados. Segmentos subascendentes, el menores sin venillas internas, raramente venas libres. par basal descendente, ápices agudos u obtusos, Venación de las láminas fértiles representada por márgenes ondulados. Venación reticulada, con aréolas aréolas costales fértiles, continuando hacia el margen marginales y costales estériles y aréolas medias venillas libres. Soros superficiales, circulares, con mayores en tamaño y fértiles. Soros circulares, uno parafisos filamentosos, terminales sobre una venilla por aréola costal, en la unión de 2 a 4 (raramente 1) libre o en la unión de dos venillas conniventes. venas conniventes. Esporas monoletes, amarillentas, Esporas monoletes, elipsoideas en vista polar y con oblongo-elípticas en vista polar, exosporio la superficie verrucosa. tuberculado y perisporio liso con algunas Distribución y hábitat.- Crece como epífito y en perforaciones (Giúdice et al., 2004). ciertas ocasiones como epipétrico. Habita en países Distribución y hábitat. Phlebodium areolatum limítrofes como Bolivia, Paraguay, Uruguay y Brasil. es una especie tropical endémica de América, que En la Argentina, es un componente del Bosque habita desde México hasta Argentina. En nuestro país Chaqueño Oriental en las provincias de Corrientes, crece en las provincias del noroeste (Jujuy, Salta, Chaco, Formosa y Santa Fe; también se encuentra en Catamarca, Tucumán) y en Misiones. Es un elemento las selvas de Misiones, extendiéndose raramente al que crece como epífito y menos frecuentemente como sur hasta el Delta del Paraná y Punta Lara, en la epipétrico (Sota de la, 1977). En Córdoba, se lo ha provincia de Buenos Aires (Sota de la, 1960). hallado adoptando este último hábitat, en Obs.: Hasta el momento ninguna especie del afloramientos rocosos en el margen del río de Los género Microgramma C. Presl ha sido citada en la Tártagos, en la localidad de Cerro Colorado (Dpto. literatura pteridológica para Córdoba. Microgramma Río Seco). Esta cita documentaría el límite más austral vacciniifolia se encuentra en el norte de nuestra de distribución de este taxón en el continente provincia, cerca del límite con Santiago del Estero, en americano (Fig. 1). el Dpto. Sobremonte, donde habita en el Bosque Obs.: En la Argentina habitan dos especies de Serrano maduro, como epífito sobre ejemplares de Phlebodium, P. areolatum y P. aureum, esta última 328 R. Morero et al., Nuevas citas de Pteridofitas para Córdoba. Fig. 1. Ubicación de las nuevas citas de Pteridofitas en la Provincia de Córdoba. corresponde a Thelypteris dentata, para Phlebodium areolatum y para Microgramma vacciniifolia. recientemente citada, en su sentido estricto, por Amer. Fern J. 26 (2): 44. 1936. primera vez para la Argentina (Meza Torres et al., Polypodium dentatum Forssk., Fl. Aegypt.- 2005). Ambas se diferencian por los siguientes Arab.:185. 1775. Tipo: Yemen, Forsskål s.n. (Holotypus caracteres: P. areolatum presenta soros 1-seriados y C, n.v.) escamas rizomáticas más pequeñas de 4-9 mm long., Iconografía.- Abbiatti, 1964: 541, 547. mientras P. aureum, posee láminas con 2-3 hileras de Descripción.- Plantas con rizomas rastreros, 8-15 soros a ambos lados de la costa y escamas rizomáticas mm de diám., con escamas castañas, brillantes, de 11-18 mm de long. pilosas. Frondes fasciculadas, de 0,25-1,20 m de Aparentemente los filopodios de Phlebodium longitud. Pecíolos castaños, 1/3 de la lámina, con aerolatum serían funcionales, ya que esta especie indumento de pelos y algunas escamas en la base. pierde la totalidad de sus frondes durante el período Láminas elípticas,
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