Amietia angolensis RdS 871 KC179832 Gephyromantis sculpturatus 2002 0187 KC179828 Hoplophryne rogersi RdS 949 KC180215 Limnonectes dabanus AMCC 193358 PT495 Ptychadena anchietae RdS 905 KC179935 Strongylopus fuelleborni RdS 994 KC179951 Tomopterna tuberculosa RdS 880 KC179959 Lithobates sp RdS 541 KC179938 77 Polypedates leucomystax K 3031 KC179932 100 Polypedates megacephalus K 3221 KC179933 Arthroleptis xenodactyloides RdS 864 KC179838 97 Arthroleptis poecilonotus FieldNumber MTR18328 PT304 70 97 Arthroleptis tanneri RdS 929 KC180092 Hyperolius guttulatus AMNH Herp 52973 PT252 64 61 Afrixalus paradorsalis AMNH Herp 44550 PT249 98 Afrixalus uluguruensis AMNH Herp 153306 PT248 98 Tachycnemis seychellensis UMMZ 189290 PT408 100 Heterixalus betsileo UMMZ 211032 PT382 64 Heterixalus tricolor UMMZ 237146 PT394 Hemisus marmoratus RdS 916 KC180102 100 Hemisus marmoratus FieldNumber MTR18344 PT298 87 Hemisus sudanensis T471 KC179889 Breviceps mossambicus RdS 903 KC180103 100 98 Breviceps gibbosus AMNH Herp 144951 PT243 84 Breviceps montanus AMNH Herp 144961 PT245 99 Callulina kisiwamsitu RdS 936 KC180213 100 Callulina kreffti FieldNumber RAN31516 PT397 100 Probreviceps macrodactylus RdS 863 KC179934 78 Spelaeophryne methneri FMNH HERP 255879 PT225 100 Spelaeophryne methneri MW 1850 KC179946 Adelastes hylonomos ROM44169 PT458 Hoplophryne rogersi ZMH 14541 PT526 Melanobatrachus indicus IND18 Dyscophus insularis AMNH Herp 173883 PT314 99 Dyscophus antongilii ZSM 1053 2003 KC179872 99 Dyscophus guineti KC180161 Phrynomantis bifasciatus FieldNumber MTR15016 PT287 100 Kaloula pulchra RdS 1200 KC179940 77 Phrynomantis microps MVZ Herp 249480 PT540 Otophryne robusta ROM 39679 KC179927 99 Otophryne robusta ROM42963 PT459 99 Otophryne steyermarki ROM 39677 KC179928 99 Otophryne steyermarki ROM42961 PT461 94 Synapturanus mirandaribeiroi SMNS 12078 KC179953 Synapturanus salseri 55 KC179954 100 85 Synapturanus salseri FieldNumber AF237 PT271 87 Synapturanus sp MW 1004 KC179952 Kalophrynus limbooliati KUHE 53284 PT470 Kalophrynus palmatissimus KUHE 52454 PT473 Kalophrynus pleurostigma ATL010 Kalophrynus stellatus KUHE 35230 PT480 100 Kalophrynus barioensis KUHE 53125 PT467 53 Kalophrynus subterrestris KUHE 53145 PT476 Kalophrynus pleurostigma AK 1097 KC179903 100 Kalophrynus interlineatus CAS HERP 232488 PT168 98 Kalophrynus interlineatus USFS 34285 KC179902 Scaphiophryne brevis AMNH Herp 173891 PT312 100 Scaphiophryne brevis voucher 2002 B2 gi 169665052 gb EU341117 96 Scaphiophryne calcarata ZSM 115 2002 KC179830 84 Scaphiophryne calcarata FieldNumber RAN31520 PT392 99 100 Scaphiophryne calcarata voucher 2002 B24 EU341118 Scaphiophryne gottlebei UMMZ 220724 PT395 Scaphiophryne marmorata voucher 2002 G63 EU341116 90 Scaphiophryne boribory voucher 2002B56 EU341114 62 Scaphiophryne madagascariensis KC179944 69 Scaphiophryne madagascariensis voucher FGMV 2002 2119 EU341115 80 Scaphiophryne sp TNHC 64007 KC822475 100 74 Scaphiophryne madagascariensis FGMV 2002 2120 KC179829 Scaphiophryne madagascariensis PT402 79 Scaphiophryne sp pet trade 2 KC180047 Anodonthyla montana MV 2001 530 KC179835 Anodonthyla boulengerii AMNH Herp 173852 PT313 94 Anodonthyla vallani FieldNumber FGZC2103 PT365 80 Anodonthyla jeanbai FieldNumber FGZC2405 PT363 58 Anodonthyla nigrigularis FieldNumber FGZC155 PT359 77 Anodonthyla theoi FieldNumber ZCMV5463 PT364 Plethodontohyla sp1 ATL026 88 Rhombophryne coronata FieldNumber 2001 A1 PT375 Rhombophryne alluaudi ZSM 3 2002 KC179930 74 Rhombophryne laevipes FieldNumber FGZC1052 PT374 Rhombophryne testudo ZSM 474 2000 KC179943 67 56 Rombophryne minuta AMNH Herp 181903 PT317 Rhombophryne mangabensis FieldNumber ZCMV890 PT377 Stumpffia roseifemoralis AMNH Herp 168381 PT265 97 100 94 Stumpffia hara FieldNumber FGZC1813 PT380 Stumpffia pygmaea FieldNumber Schachtel2001 F21 PT381 95 Stumpffia sp2 AMNH Herp 181904 PT315 92 Stumpffia psologlossa ATL029 100 99 Stumpffia gimmeli FieldNumber Schachtel2001 G6 PT379 99 Stumpffia sp1 AMNH Herp 167386 PT261 Plethodontohyla mihanika AMNH Herp 173856 PT311 57 Plethodontohyla bipunctata AMNH Herp 181895 PT320 69 Plethodontohyla brevipes FieldNumber ZCMV555 PT372 57 Rhombophryne matavy FieldNumber FGZC917 PT376 98 Plethodontohyla guentheri FieldNumber FGZC2814 PT373 99 Plethodontohyla notosticta AMNH Herp 167301 PT262 60 Platypelis grandis ATL028 Platypelis grandis ZSM 652 2003 KC179929 78 Plethodontohyla sp ZSM 649 2003 KC179931 Cophyla berara FieldNumber 2000 E16 PT367 93 Cophyla occultans AMNH Herp 167233 PT240 57 Platypelis barbouri AMNH Herp 181892 PT316 71 100 Platypelis pollicaris AMNH Herp 167235 PT241 Platypelis tetra AMNH Herp 181902 PT318 Platypelis milloti AMNH Herp 167217 PT260 79 99 Cophyla sp1 UMMZ 211167 PT404 63 Platypelis sp1 UMMZ 211271 PT405 Micryletta inornata AK 1090 KC179914 Micryletta inornata CAS HERP 247206 PT177 100 Micryletta inornata FMNH 255121 KC179915 82 Micryletta inornata K 3246 KC179918 Micryletta inornata K 1956 KC179916 85 Micryletta inornata K 3068 KC179917 Kaloula cf taprobanica IND05 Uperodon sp T1 KC179960 84 100 Uperodon systoma IND06 Ramanella obscura MM 5980 KC179936 94 64 Ramanella sp nov IND13 88 Ramanella variegata 0019C KC179937 96 100 Ramanella variegata IND15 Metaphrynella sundana FMNH HERP 231203 PT236 Kaloula pulchra USFS 34083 KC179906 55 99 Kaloula mediolineata KUHE 35178 PT479 Kaloula baleata KUHE 32313 PT468 85 100 Kaloula baleata voucher JAM3232 KC822570 65 Kaloula baleata ROM 32925 KC180032 100 Kaloula baleata ROM 32932 KC179904 Kaloula conjuncta conjuncta voucher ACD769 KC822537 95 Kaloula picta USFS 56931 KC179905 Chaperina fusca ATL009 100 Calluella yunnanensis FMNH 232988 KC179840 99 Chaperina fusca RMB 3031 KC179843 Calluella yunnanensis CAS HERP 242184 PT164 99 Microhyla achatina RMB 2620 KC179909 66 99 Calluella guttulata ATL008 Glyphoglossus molossus CAS HERP 210056 PT167 53 Glyphoglossus molossus USFS 34043 KC179883 100 Glyphoglossus molossus USFS 34044 KC179884 98 Microhyla nanapollexa AMNH Herp 163686 PT484 100 Microhyla petrigena KUHE 53743 PT483 Microhyla butleri CAS HERP 210751 KC179911 81 Microhyla heymonsi CAS HERP 210748 KC179912 89 Microhyla mantheyi KUHE 15726 PT471 Microhyla ornata CAS HERP 230957 KC179913 92 Microhyla rubra CAS HERP 215851 PT176 Microhyla berdmorei CAS HERP 204876 KC179910 58 Chaperina fusca RMB 3053 KC179844 98 Microhyla achatina MZBAmp16402 PT475 Gastrophrynoides immaculatus UKMHC HC 279 PT545 Oreophryne monticola FieldNumber MZB Amp 16265 PT478 Oreophryne brachypus ATL011 78 Oreophryne sp PT509 Oreophryne sp 2289 KC179926 78 Oreophryne anulata H 1366 KC179925 95 Oreophryne sp 9731 KC179924 100 Albericus exclamitans ABTC 98306 PT425 Choerophryne proboscidea ABTC 98918 PT441 100 100 Albericus siegfriedi ABTC 112401 PT453 100 Albericus tuberculus ABTC 98318 PT426 Cophixalus sp CCA 476L KC179864 Cophixalus balbus ABTC 111930 PT448 100 95 99 Cophixalus sp 7741 KC179863 Cophixalus crepitans ABTC 17596 PT523 99 Cophixalus hosmeri ABTC 15335 PT498 Genyophryne thomsoni ABTC 112116 PT452 100 Barygenys flavigularis ABTC 98904 PT513 98 Barygenys nana ABTC 98904PT439 Austrochaperina adelphe ABTC 29463 PT501 Austrochaperina derongo 2293 KC179839 99 Austrochaperina rivularis ABTC 46045 PT510 61 Oxydactyla alpestris ABTC 112410 PT455 Copiula sp 3235 KC179866 58 100 Copiula oxyrhina 2721 KC179865 56 100 Copiula oxyrhina ABTC 98451 PT428 Liophryne rhododactyla ATL013 Liophryne schlaginhaufeni 2380 KC179907 95 100 Liophryne schlaginhaufeni ABTC 43842 PT505 Sphenophryne cornuta ABTC 46144 PT511 100 Sphenophryne sp 2541 KC179947 Callulops personatus ABTC 98910 PT440 100 Callulops robustus 45452 KC179841 51 90 Callulops robustus ABTC 45377 PT506 80 Mantophryne lateralis ABTC 42828 PT502 Hylophorbus infulatus ABTC 98539 PT431 93 55 Hylophorbus rainerguntheri ABTC 98304 PT424 98 Hylophorbus rufescens ABTC 42916 PT503 76 Asterophrys turpicola ABTC 90180 PT521 100 Metamagnusia slater ABTC 45425i PT507 100 Metamagnusia slateri 2099 KC179842 Xenorhina obesa 6139 KC179962 100 100 Xenorhina obesa ABTC 44142 PT529 Xenorhina fuscigula ABTC 112404 PT454 99 Xenorhina mehelyi ABTC 45459 PT508 100 Xenorhina sp 3137 KC179961 Xenorhina sp 3689 KC179963 Ctenophryne aterrima QCAZ 17124 KC179923 100 Ctenophryne aterrima QCAZ17119 PT678 Melanophryne sp T4 KC179908 100 Ctenophryne aequatorialis KU 202919 KC822479 1 64 100 Ctenophryne aequatorialis KU 202911 PT546 87 Ctenophryne aequatorialis QCAZ 31243 KC179922 93 Ctenophryne geayi MPEG 25397 PT019 Ctenophryne geayi voucher AMNH A166444 DQ283383 100 90 Ctenophryne geayi ROM 40138 KC179867 59 Ctenophryne geayi WED 56812 KC179868 Chiasmocleis bassleri MPEG 27764 PT032 100 GQ891752 1 Chiasmocleis bassleri MHNSM 21576 60 Chiasmocleis haddadi voucher MNHN 3733 20110139 KF621170 100 Chiasmocleis hudsoni MPEG 18547 PT053 Chiasmocleis antenori voucher MTR28350 KF621205 64 Chiasmocleis antenori voucher QCAZ 4902 38719KF621148 Chiasmocleis antenori QCAZ 23842 KC179957 100 54 Chiasmocleis magnova KC179856 84 Chiasmocleis carvalhoi MNCN ADN 3713 PT198 Chiasmocleis tridactyla voucher JMP1948 KF621192 100 57 Chiasmocleis avilapiresae MPEG 27768 PT034 100 Chiasmocleis shudikarensis MPEG 28328 PT002 GQ891750 1 Chiasmocleisdevriesi MHNSM 21540 Chiasmocleis papachibe voucher MPEG 3803 30684 KF621184 100 GQ891751 1 Chiasmocleisventrimaculata MHNSM 21539 82 100 81 Chiasmocleis royi voucher KU 3001 215542 KF621186 100 Chiasmocleis ventrimaculata KU 215540 KC179861 Chiasmocleis albopunctata MPEG 29414 PT143 100 Chiasmocleis albopunctata JMP 218 KC179847 100 Chiasmocleis albopunctata C 621 KC179850 67 100 Chiasmocleis albopunctata C 565 KC179848 67 Chiasmocleis albopunctata C 572 KC179849 100 Chiasmocleis leucosticta T5 KC179855 Chiasmocleis schubarti CFBH 9331 KC179858 88 53 Chiasmocleis alagoanus C 2682 KC179845 73 100 Chiasmocleis alagoanus C 2683 KC179846 Chiasmocleis capixaba C 1437 KC179851 100 Chiasmocleis capixaba CFBH 22990 PT329 94 Chiasmocleis lacrimae FieldNumber LSH067 PT286 schirchi CFBH T 71 DQ283095 100 Dasypops schirchi CFBH 22938 PT332 71 Dasypops schirchi T6 KC179869 100 microps CFBH 24140 PT340 99 Myersiella microps M 1131 KC179919 100 Myersiella microps M 2358 KC179920 75 Myersiella microps Zaher KC179921 Stereocyclops histrio FieldNumber FSU 42 PT284 Hyophryne histrio MNRJ 38931 KC179890 Stereocyclops incrassatus A 399 KC180014 96 100 63 Stereocyclops incrassatus FieldNumber MRT6045 PT273 100 Stereocyclops incrassatus CFBH 9334 KC179948 61 86 Stereocyclops incrassatus CFBH 9335 KC179949 Stereocyclops incrassatus MUFAI 2482 KC179950 78 Stereocyclops incrassatus MUFAI 2483 KC180045 Arcovomer passarellii A 466 KC179837 94 100 Arcovomer passarellii A 1452 KC179836 88 Arcovomer passarellii FieldNumber MTR12362 PT281 Altigius alios KU 215544 KC179834 100 Hamptophryne alios FieldNumber APL15915 PT321 JMP 216 KC179886 99 84 Hamptophryne boliviana MPEG 27797 PT043 95 95 Hamptophryne boliviana 44 2003 KC180036 Hamptophryne boliviana WED 57567 KC179887 100 Hamptophryne boliviana WED 57569 KC179888 Dermatonotus muelleri MPEG 29412 PT148 100 Dermatonotus muelleri voucher AMNH A168436 DQ283329 98 Dermatonotus muelleri D 1344 KC179871 77 86 Dermatonotus muelleri JMP KC179985 83 Dermatonotus muelleri T7 KC179870 QCAZ 56485 Incerta sedis Elachistocleis panamensis AJC 988 KC179857 Elachistocleis ovalis NKAG 6488 KC179877 73 Elachistocleis bicolor ZVCB 10599 KC179874 94 Elachistocleis helianneae MPEG 29419 PT059 87 60 91 Elachistocleis sp1 CAS HERP 231748 PT165 78 Elachistocleis ovalis E53 KC179876 59 100 98 Elachistocleis ovalis E54 KC179879 81 Elachistocleis ovalis E71 KC179880 Elachistocleis bicolor E47 2003 KC179875 Elachistocleis piauiensis MPEG 27839 PT155 97 Elachistocleis cesarii CFBH 20007 PT335 96 Elachistocleis cesarii voucher ZUEC DCC3301 JN604511 98 53 Elachistocleis ovalis USFS 8793 KC179878 Elachistocleis ovalis voucher AMNH A141136 DQ283405 7798 Elachistocleis surinamensis E51 KC179873 66 Relictivomer pearsei JDL 26194 KC179942 84 Relictivomer pearsei JDL 26585 KC179941 elegans RdS 726 KC180151 DQ283426 Gastrophryne olivacea RdS 295 KC179881 100 Gastrophryne carolinensis CAS HERP 214349 PT166 94 97 Gastrophryne carolinensis RdS 363 KC180166 Gastrophryne mazatlanensis ATL003 Gastrophryne olivacea voucher ATH 476 KC822473 100 96 Gastrophryne usta UTA A 61716 JAC 24021 KC179882 100 UTA A 61705 PT486 Gastrophryne pictiventris 8850 KC179885 78 KU 291248 KC179892 Hypopachus barberi ENS 8580A KC179893 97 58 Hypopachus barberi UTA A 50963 ENS 8049 KC179897 74 Hypopachus barberi UTA A 55222 JAC 19613 KC179894 99 Hypopachus variolosus JHM 666 KC179900 95 Hypopachus variolosus UTA A 53757 MSM722 KC179901 Hypopachus variolosus UTA A 51790 ENS9818 KC179898 96 Hypopachus variolosus UTA A 60454 JAC 24359 KC179899 75 Hypopachus variolosus UTA A 50968 MSM 93 KC179895 62 Hypopachus variolosus FieldNumber JAC27995 PT485 96 Hypopachus variolosus UTA A 57702 JAC 23325 KC179896

0.5 0.4 0.3 0.2 0.1 0.0

Edited, based on Default Bush+