What a Data Paper Includes? We Considered Both the Topography of the Island and the Habitats of Mollusks During Our Field Investigation

DATA PAPERS AS INCENTIVES FOR OPENING BIODIVERSITY DATA: ONE YEAR OF EXPERIENCE AND PERSPECTIVES FOR THE FUTURE

A peer-reviewed open-access journal ZooKeys 261: 1–13 (2013) Molluscan fauna of Gueishan Island, Taiwan 1 doi: 10.3897/zookeys.261.4197 DATA PAPer www.zookeys.org Launched to accelerate biodiversity research

Molluscan fauna of Gueishan Island, Taiwan

1,2 2 1 2 Lyubomir Penev Vishwas Chavan Teodor Georgiev Pavel Stoev Chih-Wei Huang , Ta-Wei Hsiung , Si-Min Lin , Wen-Lung Wu

1 Department of Life Science, National Taiwan Normal University, No. 88, Sec. 4, Tingzhou Rd., Wenshan Dist., 11677, Taipei, TAIWAN, R.O.C. 2 Biodiversity Research Center, Academia Sinica, No. 128 Academia Road Sec. 2, Nankang Dist., 11529, Taipei, TAIWAN, R.O.C.

Corresponding author: Wen-Lung Wu ([email protected])

Academic editor: V. Chavan | Received 26 October 2012 | Accepted 14 January 2013 | Published 24 January 2013

Citation: Huang C-W, Hsiung T-W, Lin S-M, Wu W-L (2013) Molluscan fauna of Gueishan Island, Taiwan. ZooKeys 261: 1–13. doi: 10.3897/zookeys.261.4197

Abstract This dataset records the occurrence and inventory of molluscan fauna on Gueishan Island, the only active volcanic island in Taiwan, based on the literature survey and field investigation conducted between 2011 and 2012. The literature review involved seven studies published from 1934 to 2003, which collectively reported 112 species from 61 genera and 37 families of Mollusca on Gueishan Island. Through our field investigation, we identified 34 species from 28 genera and 23 families. Fourteen of these species were new records on Gueishan Island: Liolophura japonica, Lottia luchuana, Nerita costata, Nerita rumphii, Diplom- matina suganikeiensis, Littoraria undulata, Solenomphala taiwanensis, Assiminea sp., Siphonaria laciniosa, Laevapex nipponica, Carychium hachijoensis, Succinea erythrophana, Zaptyx crassilamellata, and Allopeas Institute for Global Biodiversity Pensoft Publishers, National Museum pyrgula. In Total, there are 126 species from 71 genera and 45 families of Mollusca on Gueishan Island. These data have been published through GBIF [http://taibif.org.tw/ipt/resource.do?r=gueishan_island] and integrated into the Taiwan Malacofauna Database (http://shell.sinica.edu.tw/).

Keywords Mollusca, Gastropoda, Bivalvia, Cephalopoda, Polyplacophora, Taiwan, Gueishan Island Project details Biodiversity and Information Facility, So�ia, Bulgaria of Natural History Project title: Investigation of molluscan fauna of Gueishan Island, Taiwan. Personnel: Chih-Wei Huang (collection identifier, data collector, data manager, data publisher), Ta-Wei Hsiung (collection identifier, data collector, data manager), Yen-Chen Lee (collection identifier), Si-Min Lin (Project Director), Wen-Lung Wu (Project Director, data manager).

Copyright Chih-Wei Huang et al. This is an open access article distributed under the terms of the Creative Commons Attribution License 3.0 (CC-BY), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited. Ecosystem Research Copenhagen, and Pensoft

2 Chih-Wei Huang et al. / ZooKeys 261: 1–13 (2013)

Funding: Academia Sinica; National Science Council, Executive Yuan, and Pensoft Publishers, Denmark Publishers, So�ia, R.O.C.(Taiwan); Forest Bureau, Council of Agriculture, Executive Yuan, R.O.C.(Taiwan). Study area descriptions/descriptor: Gueishan Island is located about 10 km from Taiwan. The island was formed via volcanic activity about 1.65 Ma ago and experienced multiple volcanic eruption events until 20 ka ago (Juang et al. 2011). It is considered the only active volcanic island near Taiwan. The land area of the island is about 2.85 km2, and the highest peak of the island is 398 meters above sea level. There are two lakes on the island, one of which consist of brackish water (Head Lake) and the other of freshwater (Tail Lake) (Figure 1). Humans colonized Gueishan Island in mid-19th century, by forming a So�ia, Bulgaria Bulgaria small village. Later in 1977, all residents were moved back to Taiwan due to military re- quirements for the island. The fauna of this island were not investigated systematically until 2000, when the island came under the management of the Northeast and Yilan Coast National Scenic Area Administration, Tourism Bureau, MOTC and was open to tourists. Design description: Island species are vulnerable to extinction due to their relatively small population size and limited access to resources. The number of species on an island represents a dynamic equilibrium between immigration and extinction. Volcanic islands provide particularly interesting cases of island biogeography, in that their biota is erased by volcanic activity and recolonized from neighboring regions. Species on Gueishan Is- land may have under gone several cycles of extinction after volcanic eruption, followed by recolonization from Taiwan when the sea-level dropped during glacial periods. Hu-

Figure 1. Location of Gueishan Island and field sampling sites of this study.

Molluscan fauna of Gueishan Island, Taiwan 3 man activity may also have provided opportunities for colonization of mollusks, either intentionally or accidentally. Investigations of molluscan fauna have been previously conducted on Gueishan Island, but these did not involve a comprehensive examination of land snails. We performed a literature survey using diverse databases, in order to col- lect previously identified reports on molluscan fauna of Gueishan Island. In addition, we performed field sampling of mollusks in marine, freshwater and terrestrial environments during 2011 and 2012 to establish the inventory of molluscan fauna of Gueishan Island. What is a Data Paper? What a Data Paper includes? We considered both the topography of the island and the habitats of mollusks during our field investigation. We focused on the terrestrial environment, as the majority of the earlier investigations examined non-terrestrial habitats. In total, our literature survey and field investigation identified 126 species from 71 genera and 45 families of Mollusca on Gueishan Island. This dataset provides basic information on the island’s biodiversity.

Taxonomic coverage

General taxonomic coverage description: The coverage of this dataset includes 126 species from 71 genera and 45 families of Mollusks of marine, freshwater and terrestrial environments on Gueishan Island (Table 1). It includes Class Gastropoda (88.10%), Class Bivalvia (8.73%), Class Cephalopoda (1.59%), and Class Polyplacophora (1.59%). The top five representative families are Cypraeidae (20 species, 15.87%), Trochidae (13 species, 10.32%), Muricidae (11 species, 8.73%), Neritidae (8 species, 6.35%), and Littorinidae (5 species, 3.97%) (Figure 2).

Table 1. Species inventory of mollusks of Gueishan Island, Taiwan.

Taxa References CLASS POLYPLACOPHORA ORDER NEOLORICATA FAMILY CHITONIDAE †Liolophura japonica (Lischke, 1873) § Liolophura sp. (Hwang and Lee 2003) CLASS CEPHALOPODA ORDER OCTOPODA FAMILY ARGONAUTIDAE Argonauta hians (Lightfoot, 1786) (Wu 2002) Octopus sp. (Hwang and Lee 2003) CLASS BIVALVIA ORDER VENEROIDA FAMILY CORBICULIDAE § (National Museum of Marine Biology and Aquarium Corbicula fluminea (Müller, 1774) 2003) FAMILY CARDIIDAE Tridacna crocea Lamarck, 1819 (Hwang and Lee 2003) Tridacna gigas (Linnaeus, 1758) (Hwang and Lee 2003)

4 Chih-Wei Huang et al. / ZooKeys 261: 1–13 (2013) • A Data Paper is a scholarly journal publication whose The Data Paper should include several important elements Tridacna maxima (Roeding, 1798) (Jung and Lai 1999) ORDER UNIONOIDA FAMILY UNIONIDAE Cristaria discoidea (Lea, 1834) (Hayasaka and Tan 1934) ORDER ARCOIDA FAMILY ARCIDAE Barbatia foliate (Forskal, 1775) (Hwang and Lee 2003) ORDER OSTREOIDA FAMILY OSTREIDAE Crassostrea gigas (Thunberg, 1793) (National Museum of Marine Biology and Aquarium 2003) (National Museum of Marine Biology and Aquarium Saccostrea mordax (Gould, 1850) 2003, Hwang and Lee 2003) FAMILY PECTINIDAE primary purpose is to describe a dataset or a group of (usually called metadata, or “description of data”) that Chlamys irregularis (Sowerby, 1842) (Jung and Lai 1999) ORDER PTERIOIDA FAMILY PTERIIDAE Pinctada margaritifera (Linnaeus, 1758) (Jung and Lai 1999), Hwang and Lee 2003 Pteria penguin (Roeding, 1798) Hwang and Lee 2003 CLASS GASTROPODA ORDER PATELLOGASTROPODA FAMILY PATELLIDAE Cellana grata (Gould, 1859) § (Jung and Lai 1999) Cellana toreuma toreuma (Reeve, 1854) § (Jung and Lai 1999, Hwang and Lee 2003) FAMILY LOTTIIDAE Collisella heroldi heroldi (Dunker, 1861) (Hwang and Lee 2003) †Lottia luchuana (Pilsbry, 1901) § Notoacmea schrenckii schrenckii (Jung and Lai 1999, Wu 2002, Hwang and Lee 2003) datasets, rather than to report a research investigation. document, for example: (Lischke, 1868) ORDER VETIGASTROPODA FAMILY HALIOTIDAE Haliotis diversicolor (Reeve, 1846) (Jung and Lai 1999) FAMILY TROCHIDAE Calliostoma unicum (Dunker, 1860) (Jung and Lai 1999) Chlorostoma turbinatum (Jung and Lai 1999) A.Adams, 1853 Chlorostoma argyrostoma argyrostoma (Jung and Lai 1999, Wu 2002, Hwang and Lee 2003) (Gmelin, 1791) Monodonta labio (Linnaeus, 1758) (Jung and Lai 1999, Hwang and Lee 2003) Monodonta perplexa Pilsbry, 1889 § (Jung and Lai 1999, Hwang and Lee 2003) Stomatella planulata (Lamarck, 1816) (Jung and Lai 1999) Tectus conus (Gmelin, 1791) (Jung and Lai 1999) As such, it contains facts about data, not hypotheses and • methods of data collection; Tectus pyramis (Born, 1778) (Jung and Lai 1999) Trochus chloromphalus A. Adams, 1853 (Jung and Lai 1999) Trochus hanleyanus Reeve, 1842 (Jung and Lai 1999, Wu 2002, Hwang and Lee 2003) Trochus maculates Linnaeus, 1758 (Jung and Lai 1999, Hwang and Lee 2003) Trochus sacellum Philippi, 1854 (Jung and Lai 1999, Hwang and Lee 2003) Trochus stellatus Gmelin, 1790 (Jung and Lai 1999, Hwang and Lee 2003) FAMILY TURBINIDAE arguments in support of the data, as found in a conventional • taxa covered;

Molluscan fauna of Gueishan Island, Taiwan 5

Astralium haematragum (Menke, 1829) (Hwang and Lee 2003) Lunella coronate (Gmelin, 1818) (Hwang and Lee 2003) ORDER NERITIMORPHA FAMILY NERITIDAE Nerita albicilla Linnaeus, 1758 (Jung and Lai 1999, Hwang and Lee 2003) †Nerita costata Gmelin, 1791 § Nerita plicata Linnaeus, 1758 § (Jung and Lai 1999) †Nerita rumphii Recluz,1841 § research article. Its purposes are three-fold: • spatial and temporal ranges and geographical coverage; Nerita chamaeleon Linnaeus, 1758 (Hwang and Lee 2003) Nerita ocellata Leguillou, 1841 (Jung and Lai 1999) Nerita polita Linnaeus, 1758 (Jung and Lai 1999) Nerita undata Linnaeus, 1758 (Jung and Lai 1999) ORDER CAENOGASTROPODA FAMILY DIPLOMMATINIDAE †Diplommatina suganikeiensis § (Pilsbry & Hirase, 1905) FAMILY PLANAXIDAE Planaxis sulcatus (Born, 1778) (Hwang and Lee 2003) FAMILY POTAMIDIDAE (National Museum of Marine Biology and Aquarium Batillaria zonalis (Bruguiere, 1792) 2003) FAMILY THIARIDAE • collectors and owners of the data; Tarebia granifera (Lamarck, 1822) § (National Museum of Marine Biology and Aquarium 2003) Thiara scabra (Muller, 1774) § (National Museum of Marine Biology and Aquarium 2003) Thiara tuberculata (Muller 1774) § (National Museum of Marine Biology and Aquarium 2003) FAMILY CYPRAEIDAE Cypraea annulus Linnaeus, 1758 (Jung and Lai 1999, Hwang and Lee 2003) Cypraea arabica Linnaeus, 1758 (Jung and Lai 1999) Cypraea asellus Linnaeus, 1758 (Jung and Lai 1999) Cypraea caputserpentis Linnaeus, 1758 (Jung and Lai 1999, Hwang and Lee 2003) Cypraea caurica Linnaeus, 1758 (Jung and Lai 1999) Cypraea chinensis Gmelin, 1791 (Jung and Lai 1999) Cypraea clandestine Linnaeus, 1758 (Jung and Lai 1999) Cypraea eglantine Duclos, 1833 (Hwang and Lee 2003) Cypraea erosa Linnaeus, 1758 (Jung and Lai 1999) • to provide a citable journal publication that brings • software used to create or view the data. Cypraea gracilis Gaskoin, 1849 (Jung and Lai 1999) Cypraea helvola Linnaeus, 1758 (Jung and Lai 1999) Cypraea labrolineata Gaskoin, 1849 (Jung and Lai 1999) Cypraea lynx Linnaeus, 1758 (Hwang and Lee 2003) Cypraea moneta Linnaeus, 1758 § (Jung and Lai 1999, Hwang and Lee 2003) Cypraea onyx Linnaeus, 1758 (Jung and Lai 1999) Cypraea poraria Linnaeus, 1758 (Jung and Lai 1999) Cypraea testudinaria Linnaeus, 1758 (Jung and Lai 1999) Cypraea tigris Linnaeus, 1758 (Jung and Lai 1999) Cypraea ziczac Linnaeus, 1758 (Jung and Lai 1999) Cypraea errones Linnaeus, 1758 (Wu 2002) FAMILY OVULIDAE scholarly credit to data publishers;

6 Chih-Wei Huang et al. / ZooKeys 261: 1–13 (2013)

Calpurnus verrucosus (Linnaeus, 1758) (Hwang and Lee 2003) Ovula ovum Linnaeus, 1758 (Hwang and Lee 2003) FAMILY LITTORINIDAE • to describe the data in a structured human-readable form; The metadata can be used to form a Data Paper manuscript. Littoraria pintado (Wood, 1828) (Jung and Lai 1999) †Littoraria undulate (Gray, 1839) § Littoraria scabra scabra (Linnaeus, (Hwang and Lee 2003) 1758) Nodilittorina pyramidalis (Quay & § (Jung and Lai 1999, Hwang and Lee 2003) Gaimard, 1833) Nodilittorina vidua (Gould, 1859) § (Jung and Lai 1999, Hwang and Lee 2003) FAMILY ASSIMINEIDAE †Solenomphala taiwanensis § (Habe, 1942) †Assiminea sp. § FAMILY BURSIDAE Bursa granularis (Roeding, 1798) (Jung and Lai 1999) FAMILY RANELLIDAE • to bring the existence of the data to the attention of the Cymatium aquatile (Reeve, 1844) § (Jung and Lai 1999) Cymatium mundum (Gould, 1849) (Jung and Lai 1999) Cymatium pileare (Linnaeus, 1758) (Jung and Lai 1999) Cymatium lotorium (Linnaeus, 1758) (Hwang and Lee 2003) FAMILY COLUMBELLIDAE Pyrene punctata (Bruguiere, 1789) (Jung and Lai 1999) Pyrene testudinaria testudinaria (Hwang and Lee 2003) (Link, 1806) FAMILY FASCIOLARIIDAE Peristernia nassatula (Lamarck, 1822) (Wu 2002) FAMILY NASSARIIDAE Nassarius glans (Linnaeus, 1758) (Jung and Lai 1999) Nassarius papillosus (Linnaeus, 1758) (Jung and Lai 1999) scholarly community. An important feature of Data Papers is that they should Telasco velatus (Gould, 1850) (Jung and Lai 1999) FAMILY MURICIDAE Chicoreus torrefactus Sowerby, 1841 (Wu 2002) Chicoreus brunneus (Link, 1807) (Hwang and Lee 2003) Drupa morum Roeding, 1798 (Jung and Lai 1999) Drupa ricina ricina (Linnaeus, 1758) (Jung and Lai 1999, Wu 2002, Hwang and Lee 2003) Drupa rubusidaea Roeding, 1798 (Jung and Lai 1999, Hwang and Lee 2003) Ergalatax contractus (Reeve, 1846) (Jung and Lai 1999, Hwang and Lee 2003) Mancinella mancinella § (Jung and Lai 1999) (Linnaeus, 1758) Morula uva (Roeding, 1798) (Wu 2002) Purpura panama (Roeding, 1798) (Jung and Lai 1999, Wu 2002) Tenguella granulate (Duclos, 1924) §, (Jung and Lai 1999) Thais clavigera (Kuster, 1860) (Wu 2002, Hwang and Lee 2003) always be linked to the published datasets they describe, FAMILY TURBINELLIDAE Vasum ceramicum (Linnaeus, 1758) (Hwang and Lee 2003) and that link (a URL, ideally resolving a DOI) should be Molluscan fauna of Gueishan Island, Taiwan 7

FAMILY CONIDAE Conus flavidus Lamarck, 1810 (Wu 2002, Hwang and Lee 2003) Conus lividus Hwass, 1792 (Hwang and Lee 2003) Conus textile Linnaeus, 1758 (Jung and Lai 1999, Wu 2002) Conus striatus Linnaeus, 1758 (Wu 2002) ORDER HETEROBRANCHIA FAMILY APLYSIIDAE Aplysia juliana Quoy & (Hwang and Lee 2003) Gaimard, 1832 Aplysia oculifera Adams & (Hwang and Lee 2003) Reeve, 1850 Dolabrifera dolabrifera (Rang, 1928) (Hwang and Lee 2003) published within the paper itself. Conversely, the metadata FAMILY PHYLLIDIIDAE Phyllidia pustulosa Cuvier, 1804 (Hwang and Lee 2003) Phyllidia varicose Lamarck, 1801 (Hwang and Lee 2003) FAMILY SIPHONARIIDAE †Siphonaria laciniosa § (Linnaeus, 1758) FAMILY PLANORBIDAE †Laevapex nipponica (Kuroda, 1947) § FAMILY ELLOBIIDAE †Carychium hachijoensis § Pilsbry, 1902 The future FAMILY VERONICELLIDAE Vaginulus alte (Ferussac, 1821) § (Wu 2002) describing the dataset held within data archives should FAMILY SUCCINEIDAE †Succinea erythrophana Ancey, 1883 § FAMILY CLAUSILIIDAE †Zaptyx crassilamellata Kuroda, 1941 § FAMILY ACHATINIDAE Achatina fulica Bowdich, 1822 § (Wu 2002) FAMILY SUBULINIDAE †Allopeas pyrgula (Schmacker & § Boettger, 1891) FAMILY PHILOMYCIDAE Meghimatium bilineatum (Wu 2002) (Benson, 1842) FAMILY CAMAENIDAE Coniglobus melleus (Pfeiffer, 1865) (Kuroda 1938, Kuroda 1941) include the bibliographic details of the Data Paper once that FAMILY BRADYBAENIDAE Acusta despecta (Sowerby, 1839) (Kuroda 1938, Kuroda 1941) Aegista mackensii (Adams & § (Jung and Lai 1999, Wu 2002) Reeve, 1850) Aegista osbeckii (Philippi, 1847) § (Kuroda 1938, Kuroda 1941, Wu 2002) Bradybaena similaris (Ferussac, 1822) § (Kuroda 1941) †New records on Gueishan Island § Collected by our field sampling is published, including a resolvable DOI.

8 Chih-Wei Huang et al. / ZooKeys 261: 1–13 (2013) Elaborating formats for descriptions of various kinds of data At the time of submission of the Data Paper manuscript, the (ecological, morphological, genomic, environmental, etc.). data described should be freely available online in a public

Figure 2. Taxonomic coverage. A Class B The top five representative families. Numbers in pie charts represent number of species. Taxonomic ranks Link to various platforms (GBIF, Scratchpads, Dryad, EDIT, Phylum: Mollusca Class: Bivalvia, Cephalopoda, Gastropoda, Polyplacophora Order: Arcoida, Caenogastropoda, Heterobranchia, Neoloricata, Neritimorpha, Octopoda, Ostreoida, Patellogastropoda, Pterioida, Unionoida, Veneroida, Vetigastropoda repository under a suitable data license, so that they can CBOL, etc.) to provide automated data paper submission.

Molluscan fauna of Gueishan Island, Taiwan 9

Family: Achatinidae, Aplysiidae, Arcidae, Argonautidae, Assimineidae, Bradybae- be retrieved for reuse, resampling and redistribution by nidae, Bursidae, Camaenidae, Cardiidae, Chitonidae, Clausiliidae, Columbelli- dae, Conidae, Corbiculidae, Cypraeidae, Diplommatinidae, Ellobiidae, Fascio- lariidae, Haliotidae, Littorinidae, Lottiidae, Muricidae, Nassariidae, Neritidae, Octopodidae, Ostreidae, Ovulidae, Patellidae, Pectinidae, Philomycidae, Phyl- lidiidae, Planaxidae, Planorbidae, Potamididae, Pteriidae, Ranellidae, Siphonari- idae, Subulinidae, Succineidae, Thiaridae, Trochidae, Turbinellidae, Turbinidae, Unionidae, Veronicellidae Genus: Achatina, Acusta, Aegista, Allopeas, Aplysia, Argonauta, Assiminea, Astralium, Barbatia, Batillaria, Bradybaena, Bursa, Calliostoma, Calpurnus, Carychium, Cellana, Chicoreus, Chlamys, Chlorostoma, Collisella, Coniglobus, Conus, Cor- bicula, Crassostrea, Cristaria, Cymatium, Cypraea, Diplommatina, Dolabrifera, Drupa, Ergalatax, Haliotis, Laevapex, Liolophura, Littoraria, Lottia, Lunella, anyone for any purpose, subject to one condition at most - Mancinella, Meghimatium, Monodonta, Morula, Nassarius, Nerita, Nodilitto- rina, Notoacmea, Octopus, Ovula, Patella, Peristernia, Phyllidia, Pinctada, Pla- naxis, Pteria, Purpura, Pyrene, Saccostrea, Siphonaria, Solenomphala, Stomatella, Succinea, Tarebia, Tectus, Telasco, Tenguella, Thais, Thiara, Tridacna, Trochus, Vaginulus, Vasum, Zaptyx.

Spatial coverage

General spatial coverage: The spatial coverage of the literature and our field investigation ranged from a latitude of 24°49'48"N to 24°51'0"N and a longitude of 121°55'48"E to 121°57'36"E. It includes the marine, intertidal, freshwater and terrestrial environment of Gueishan Island, Taiwan (Figure 1) that of proper attribution using scholarly norms. Coordinates: 24°49'48"N and 24°51'0"N Latitude; 121°55'48"E and 121°57'36"E Longitude

Temporal coverage:

1934–2012.

Methods Sampling description:

Literature survey: We searched for publications (including journals, project reports, theses and books) associated with the molluscan fauna of Gueishan Island from the following databases: (1) the National Digital Library of Theses and Dissertations in Taiwan (http://ndltd.ncl.edu.tw) (this contains details of theses and dissertations pub- Examples of data papers

10 Chih-Wei Huang et al. / ZooKeys 261: 1–13 (2013) lished since 1956, but did not contain publications relevant to this study); (2) the National Bibliographic Information Network (http://nbinet3.ncl.edu.tw) (this catalog integrates information from National Central Library and 74 other libraries containing all publications with a Taiwan ISBN and selected government project reports; three publications (Wu 2002, National Museum of Marine Biology and Aquarium 2003, Hwang and Lee 2003) from this database met our requirement); (3) the Government Research Bulletin (http://grbsearch.stpi.narl.org.tw/GRB/) (this contains government project reports made since 1993, but did not contain reports relevant to this study); (4) Google Scholar (http://scholar.google.com.tw/) (this contains a wide range of resources, from journals and books to webpages, and it provided two relevant journal articles (Chen and Fu 2007, Lee and Chen 2010)); (5) The Taiwan Malacofauna Da- tabase (http://shell.sinica.edu.tw/) (this database contains taxonomy, distribution and references of all mollusks occurred in Taiwan, and provided six relevant publications (Lee and Wu 1998, Jung and Lai 1999, Wu 2002, National Museum of Marine Biol- ogy and Aquarium 2003, Hwang and Lee 2003, Chen and Fu 2007)). In addition, three relevant publications (Hayasaka and Tan 1934, Kuroda 1938, Kuroda 1941) were identified from citations in Wu (2002). In total, we identified ten relevant publications. Three of these publications (Lee and Wu 1998, Chen and Fu 2007, Lee and Chen 2010) were excluded because they described specimens acquired from fishing ports that had been captured by shrimp fishing or bottom trawling boats near Gueis- han Island, without information of the precise sampling location. The seven remaining publications were used to establish the occurrence and inventory data. Sampling sites, names of collectors and the scientific name of each species were recorded using Mi- crosoft EXCEL 2010. All of the publications mentioned above can be accessed in the National Central Library and the National Taiwan Library. Field Sampling: The topology of Gueishan Island and the types of mollusk habitat were considered for field investigation. Visual search was conducted for mollusks in intertidal, freshwater and terrestrial environments (Figure 1). The surface of rocks on the coastline and man-made concrete structures in port were searched for marine mollusks during low tide. Leaf litter and rocks under or near water around Tail Lake (the only freshwater habitat on island) were inspected for freshwater mollusks. We inspected from leaves, trunks, leaves litter, rocks and rotten woods for land snails along three trails: one trail around Tail Lake, another leads to the highest peak (401 Highland) on the island, and the other leads to the northern part of the island. We surveyed for land snails during their active periods: during and after rainfall, early morning, and night. GBIF and Pensoft pioneered a work�low between the GBIF’s Integrated Publishing Toolkit (IPT) and Pensoft’s journals ZooKeys, At least one living individual or dead shells of each species was collected as voucher specimens. Living organisms were brought back to laboratory, fixed via freezing in a -80°C freezer, and subsequently transferred to 95% ethanol for long term preservation. Quality control description: Latitude, longitude and altitude of sampling sites were recorded using Garmin GPSmap 60CSx with uncertainty of less than 10 meters. Sampling sites were georeferenced (WGS84). All the specimens collected during the field investigation were identified independently by Huang and Hsiung. Seven earlier studies described the mollucan fauna of Gueishan Island, but these publications lack PhytoKeys, MycoKeys, Nature Conservation, etc. to automatically export metadata into the form of a data paper manuscript,

Molluscan fauna of Gueishan Island, Taiwan 11 clear photos or other information for identifying specimens. Species identification was performed using the following guide books and publications about Taiwan malacofauna: Pace (1973), Lai (1990, 1998), Lee and Chen (2003), Wu and Lee (2005), and Hsieh et al. (2006). Newly recorded species were further confirmed by Dr. Yen-Chen Lee, a Mollusca specialist and postdoctoral researcher in the Biodiversity Research Center, Academia Sinica. Fourteen new recorded species were found to be native to based on the Ecological Metadata Language (EML). Taiwan but previously unreported on Gueishan Island. The scientific names of all mol- ZooKeys 204: 47–52 (2012) A peer-reviewed open-access journal ZooKeys 198: 103–109 (2012) A peer-reviewed open-access journal ZooKeys 152: 87–91 (2011) A peer-reviewed open-access journal ZooKeys 205: 5–18 (2012) A peer-reviewed open-access journal Nature Conservation 4: 1–12 (2013) A peer-reviewed open-access journal PhytoKeys 12: 59–67 (2012) A peer-reviewed open-access journal lusks were checked against the Taiwan Malacofauna Database and World Register of Antarctic, Sub-Antarctic and cold temperate echinoid database 47 A dataset from bottom trawl survey around Taiwan 103 Project description: DNA barcodes of bird species in the National Museum of Natural History... 87 MOSCHweb — a matrix-based interactive key to the genera of the Palaearctic Tachinidae... 5 Antarctic macrobenthic communities: A compilation of circumpolar information 1 Florabank1: a grid-based database on vascular plant distribution... 59 Marine Species (http://www.marinespecies.org/). doi: 10.3897/zookeys.204.3134 DAtA PAPer doi: 10.3897/zookeys.198.3032 DATA PAPer doi: 10.3897/zookeys.152.2473 DAtA PAPer doi: 10.3897/zookeys.205.3409 OnlIne IDenTIfICATIOn Key doi: 10.3897/natureconservation.4.4499 DAtA pAper doi: 10.3897/phytokeys.12.2849 DAtA PAPer

www.zookeys.org Launched to accelerate biodiversity research www.zookeys.org Launched to accelerate biodiversity research www.zookeys.org Launched to accelerate biodiversity research www.zookeys.org Launched to accelerate biodiversity research http://www.pensoft.net/natureconservation Launched to accelerate biodiversity conservation www.phytokeys.com Launched to accelerate biodiversity research Data resources

The data underpinning the analysis reported in this paper are deposited at GBIF, the Global Biodiversity Information Facility, http://taibif.org.tw/ipt/resource. do?r=gueishan_island

Datasets

Dataset description: This dataset incorporates seven publications (Hayasaka and Tan 1934, Kuroda 1938, Kuroda 1941, Jung and Lai 1999, Wu 2002, National Museum of Marine Biology and Aquarium 2003, Hwang and Lee 2003) associated with the Antarctic, Sub-Antarctic and cold temperate A dataset from bottom trawl survey around Taiwan Project Description: DNA Barcodes of Bird Species MOSCHweb — a matrix-based interactive key to the Antarctic macrobenthic communities: Florabank1: a grid-based database on vascular plant molluscan fauna of Gueishan Island and field investigation results. The dataset includes sampling date, taxonomy information, GPS location, elevation, type of habitat, name of collector, method of collection, and literature record. Based on the literature published during the period between 1934 and 2003, 112 species from 61 genera and 37 families echinoid database in the National Museum of Natural History, genera of the Palaearctic Tachinidae (Insecta, Diptera) A compilation of circumpolar information distribution in the northern part of Belgium of Mollusca were recorded on Gueishan Island. Of the 34 species from 28 genera and 23 families identified during our 2011-2012 field investigation, fourteen species were 1 1 1 new records on Gueishan Island. In total, our literature survey and field investigation Smithsonian Institution, USA (Flanders and the Brussels Capital region) documents 126 species from 71 genera and 45 families of Mollusca on Gueishan Island. Kwang-Tsao Shao ,Jack Lin , Chung-Han Wu , The fourteen newly recorded species are: Liolophura japonica (Lischke, 1873), Lottia lu- 1 1 chuana (Pilsbry, 1901), Siphonaria laciniosa (Linnaeus, 1758), Nerita costata Gmelin, 1 1 1 1 Hsin-Ming Yeh , Tun-Yuan Cheng 1,2,† 3,‡ 2,§ 1 2 3 4 1791, Nerita rumphii Recluz,1841, and Littoraria undulata (Gray, 1839), which were Benjamin Pierrat , Thomas Saucède , Alain Festeau , Bruno David Pierfilippo Cerretti , Hans-Peter Tschorsnig , Massimo Lopresti , Julian Gutt , Barnes David , Susanne J. Lockhart , Anton van de Putte sampled from the marine environment; Assiminea sp. and Laevapex nipponica (Kuroda, 1,2,§ 1947), which were discovered in a freshwater environment, Tail Lake; and Solenom- 1 2 3 1 Filippo Di Giovanni 1,2 3 1 phala taiwanensis (Habe, 1942), Diplommatina suganikeiensis (Pilsbry & Hirase, 1905), 1 Biodiversity Research Center, Academia Sinica, No. 128, Sec. 2, Academia Rd., Nankang, Taipei, TAIWAN, David E. Schindel , Mark Y. Stoeckle , Chris M. Milensky , Michael Trizna , Wouter Van Landuyt , Leo Vanhecke , Dimitri Brosens Carychium hachijoensis Pilsbry, 1902, Zaptyx crassilamellata Kuroda, 1941, Allopeas pyr- 1 UMR CNRS 6282 Biogéosciences, Université de Bourgogne, 6 boulevard Gabriel, 21000, Dijon, France 3 3 3,4 1 Alfred Wegener Institute Helmholtz Centre for Polar and Marine Research, Columbusstr., 27568, Bremerhaven, gula (Schmacker & Boettger, 1891), and Succinea erythrophana Ancey, 1883 discovered R.O.C. Brian K. Schmidt , Christina A. Gebhard , Gary R. Graves in the terrestrial environment. This dataset provide basic information for the island’s 1 Dipartimento di Biologia e Biotecnologie “Charles Darwin”, Università di Roma “La Sapienza”, Piazzale Germany 2 British Antarctic Survey, High Cross, Madingley Road, CB3 0ET, Cambridge, United Kingdom biodiversity and biogeography. This dataset will be maintained by the Malacology Lab, Corresponding author: Benjamin Pierrat ([email protected]) 1 Research Institute for Nature and Forest, Kliniekstraat 25, 1070, Brussels, Belgium 2 Flo.Wer, Bouchout Biodiversity Research Center, Academia Sinica. A. Moro 5, 00185, Rome, Italy 2 Centro Nazionale per lo Studio e la Conservazione della Biodiversità Fo- 3 NOAA Southwest Fisheries Science Center, La Jolla Shore Drive, CA 92037, La Jolla, United States 4 ANTA- Object name: Darwin Core Archive Molluscan fauna of Gueishan Island, Taiwan Corresponding author: Kwang-Tsao Shao ([email protected]); Jack Lin ([email protected]) 1 Consortium for the Barcode of Life, MRC-105, National Museum of Natural History, Smithsonian In- Domain, Nieuwelaan 38, 1070, Meise, Belgium 3 NBGB (National Botanic Garden of Belgium), Bouchout restale – Corpo Forestale dello Stato, Via Carlo Ederle 16/A, 37100, Verona, Italy 3 Staatliches Museum für BIF, Vautierstraat 29, B-1000 Brussels, Belgium Academic editor: stitution, P. O. Box 37012, Washington, D.C. 20013-7012 USA 2 Program for the Human Environment, Domain, Nieuwelaan 38, 1860, Meise, Belgium V. Chavan | Received 27 March 2012 | Accepted 14 June 2012 | Published 25 June 2012 Naturkunde, Rosenstein 1, 70191 Stuttgart, Germany Academic editor: V. Chavan | Received 29 February 2012 | Accepted 22 May 2012 | Published 30 May 2012 Rockefeller University, New York, USA 3 Department of Vertebrate Zoology, MRC-116, National Museum of Corresponding author: Julian Gutt ([email protected]) Corresponding author: Wouter Van Landuyt Citation: Pierrat B, Saucède T, Festeau A, David B (2012) Antarctic, Sub-Antarctic and cold temperate echinoid database. 4 ([email protected]) Citation: Natural History, Smithsonian Institution, P. O. Box 37012, Washington, D.C. 20013-7012 USA Center for Corresponding authors: Pierfilippo Cerretti ([email protected]), Hans-Peter Tschorsnig (hanspeter. ZooKeys 204: 47–52. doi: 10.3897/zookeys.204.3134 Shao K-T, Lin J, Wu C-H, Yeh H-M, Cheng T-Y (2012) A dataset from bottom trawl survey around Taiwan. 12 Chih-Wei Huang et al. / ZooKeys 261: 1–13 (2013) Macroecology, Evolution and Climate, University of Copenhagen, DK-2100 Copenhagen, Denmark [email protected]) Academic editor: L. Penev | Received 12 December 2012 | Accepted 12 February 2013 | Published 19 February 2013 ZooKeys 198: 103–109. doi: 10.3897/zookeys.198.3032 Academic editor: Vishwas Chavan | Received 6 February 2012 | Accepted 15 May 2012 | Published 16 May 2012 Character encoding: UTF-8 Format name: Darwin Core Archive format Corresponding author: M. Y. Stoeckle ([email protected]) Citation: Format version: 1.0 Gutt J, David B, Lockhart SJ, van de Putte A (2013) Antarctic macrobenthic communities: A compilation of Citation: Distribution: http://taibif.org.tw/ipt/archive.do?r=gueishan_island Academic editor: Lyubomir Penev | Received 22 May 2012 | Accepted 26 June 2012 | Published 4 July 2012 Van Landuyt W, Vanhecke L, Brosens D (2012) Florabank1: a grid-based database on vascular plant Publication date of data: 2012-12-21 circumpolar information. Nature Conservation 4: 1–12. doi: 10.3897/natureconservation.4.4499 Language: English Abstract distribution in the northern part of Belgium (Flanders and the Brussels Capital region). PhytoKeys 12: 59–67. doi: Licenses of use: This work is licensed under a Creative Commons CCZero 1.0 License Abstract Academic editor: L. Penev | Received 29 November 2011 | Accepted 2 December 2011 | Published 8 December 2011 Citation: Cerretti P, Tschorsnig H-P, Lopresti M, Di Giovanni F (2012) MOSCHweb — a matrix-based interactive http://creativecommons.org/publicdomain/zero/1.0/legalcode This database includes spatial data of Antarctic, Sub-Antarctic and cold temperate echinoid distribution 10.3897/phytokeys.12.2849 Metadata language: English Date of metadata creation: 2012-09-21 Bottom trawl fishery is one of the most important coastal fisheries in Taiwan both in production and key to the genera of the Palaearctic Tachinidae (Insecta, Diptera). ZooKeys 205: 5–18. doi: 10.3897/zookeys.205.3409 Hierarchy level: Dataset (Echinodermata: Echinoidea) collected during many oceanographic campaigns led in the Southern Hemi- Citation: Schindel DE, Stoeckle MY, Milensky C, Trizna M, Schmidt B, Gebhard C, Graves G (2011) Project description: economic values. However, its annual production started to decline due to overfishing since the 1980s. Its sphere from 1872 to 2010. The dataset lists occurrence data of echinoid distribution south of 35°S lati- DNA barcodes of bird species in the National Museum of Natural History, Smithsonian Institution, USA. ZooKeys 152: Abstract Acknowledgements bycatch problem also damages the fishery resource seriously. Thus, the government banned the bottom Comprehensive information on Antarctic macrobenthic community structure is publicly available since tude, together with information on taxonomy (from species to genus level), sampling sources (cruise ID, 87–92. doi: 10.3897/zookeys.152.2473 Abstract We would like to thank Yilan County Government and Northeast and Yilan Coast fishery within 3 nautical miles along the shoreline in 1989. To evaluate the effectiveness of this policy, a National Scenic Area Administration, Tourism Bureau, MOTC for approval of field sampling dates, ship names) and sampling sites (geographic coordinates and depth). Echinoid occurrence Abstract the 1960s. It stems from trawl, dredge, grab, and corer samples as well as from direct and camera ob- investigation on Gueishan Island. Fieldwork was made possible by the assistance of Florabank1 is a database that contains distributional data on the wild flora (indigenous species, archeo- Toucheng Fishermen’s Association, all members of the Malacology Laboratory (Biodi- data were compiled from the Antarctic Echinoid Database (David et al. 2005a), which integrates records four year survey was conducted from 2000–2003, in the waters around Taiwan and Penghu (Pescadore) servations (Table 1–2). The quality of this information varies considerably; it consists of pure descrip- versity Research Center, Academia Sinica) and all members of the Laboratory of Grass We provide a general overview of features and technical specifications of an original interactive key web phytes and naturalised aliens) of Flanders and the Brussels Capital Region. It holds about 3 million records Lizard and Fish (Department of Life Science, National Taiwan Normal University). Islands, one region each year respectively. All fish specimens collected from trawling were brought back tions, figures for presence (absence) and abundance of some key taxa or proxies for such parameters, We would like to thank Dr. Duncan Wright as well as two anonymous reviewers for from oceanographic cruises led in the Southern Ocean until 2003. This database has been upgraded to application for the identification of Palaearctic Tachinidae genera. The full list of terminal taxa included helpful comments which greatly improved the manuscript. to lab for identification, individual number count and body weight measurement. These raw data have Abstract e.g. sea-floor cover. Some data sets even cover a defined and complete proportion of the macrobenthos of vascular plants, dating from 1800 till present. Furthermore, it includes ecological data on vascular plant take into account data from oceanographic cruises led after 2003. The dataset now reaches a total of 6160 in the key, which is the most updated list of genera currently recorded for the Palaearctic Region, is given. been integrated and established in Taiwan Fish Database (http://fishdb.sinica.edu.tw). They have also The Division of Birds, National Museum of Natural History, Smithsonian Institution in Washington, with further analyses on diversity and zoogeography. As a consequence the acquisition of data from ap- species, redlist category information, Ellenberg values, legal status, global distribution, seed bank etc. The references occurrence data that have been checked for systematics reliability and consistency. It constitutes today the We also briefly discuss the need for dealing with detailed and standardized taxa descriptions as a base to DC, has obtained and released DNA barcodes for 2,808 frozen tissue samples. Of the 1,403 species proximately 90 different campaigns assembled here was not standardised. Nevertheless, it was possible database is an initiative of “Flo.Wer” (www.plantenwerkgroep.be), the Research Institute for Nature and Chen WD, Fu IF (2007) A New Species of Gibbula (Gastropoda: Trochidae) from Taiwan. most complete database on Antarctic and Sub-Antarctic echinoids. been published through TaiBIF (http://taibif.tw), FishBase and GBIF (website see below). This dataset Bulletin of Malacology 31: 10–16. keep matrix-based interactive tools easily updated, by proposing a standardized protocol. to classify this broad variety of known macrobenthic assemblages to the best of expert knowledge (Gutt Forest (INBO: ) and the National Botanic Garden of Belgium (www.br.fgov.be). Florabank Hayasaka I, Tan K (1934) Three species of Mollusca in Taiwan. Transactions of the National contains 631 fish species and 3,529 records, making it the most complete demersal fish fauna and their represented by these samples, 1,147 species have not been barcoded previously. This data release increases www.inbo.be History Society of Formosa 24(133): 259–263. [In Japanese] 2007; Fig. 1). This overview does not replace statistically sound community and diversity analyses. Hsieh BC, Hwang CC, Wu SP (2006) Landsnails of Taiwan. Taiwan Forestry Bureau, Council temporal and spatial distributional data on the soft marine habitat in Taiwan. the number of bird species with standard barcodes by 91%. These records meet the data standard of the aims at centralizing botanical distribution data gathered by both professional and amateur botanists and of Agriculture, Executive Yuan, Taiwan, Taiwan, 277 pp. [In Chinese] Keywords Keywords However, it shows from where which kind of information is available and it acts as an example of the Hwang JS, Lee CS (2003) Investigation of marine organisms and tourism resource of sub- Consortium for the Barcode of Life and they have the reserved keyword BARCODE in GenBank. The to make these data available to the benefit of nature conservation, policy and scientific research. marine hot spring of Gueishan Island. Northeast and Yilan Coast National Scenic Area feasibility and power of such data collections. The data set provides unique georeferenced biological Administration, Tourism Bureau, MOTC, Yilan, Taiwan, 146 pp. [In Chines] Southern Ocean, echinoids, Antarctic species, Sub-Antarctic species, cold temperate species Interactive key, identification tool, web application, data matrix, morphology, description protocol, Dip- The occurrence data contained in Florabank1 are extracted from checklists, literature and herbarium Juang WS, Chang SS, Chen JC (2011) Exploration the history of volcanic activities at Kueis- Keywords data are now available on GenBank and the Barcode of Life Data Systems. basic information for the planning of future coordinated research activities, e.g. under the umbrella of hantao based upon the transition of its geological and geomorphic landscape. Bulletin of tera, Tachinidae, Palaearctic Region the Central Geological Survey 24: 155–188. [In Chinese with English abstract] specimen information. Of survey lists, the locality name (verbatimLocality), species name, observation Jung BS, Lai KY (1999) Notes on mollusca collection on Gueishan Island. The Pei-Yo 25: Bottom Trawl, Taiwan, IPT, Darwin Core, Fishery the biology program “Antarctic Thresholds - Ecosystem Resilience and Adaptation” (AnT-ERA) of the 20–22. [In Chinese] Keywords date and IFBL square code, the grid system used for plant mapping in Belgium (Van Rompaey 1943), Kuroda T (1938) Geographic distribution of land snails in Taiwan. Taiwan Tigaku Kizi 9(4): Scientific Committee on Antarctic Research (SCAR) and especially for actual conservation issues, e.g. 99–108. [In Japanese] Data published through Project details GBIF: http://fishbase.tw:8080/ipt/resource.do?r=bottom_ DNA barcoding, GenBank, BOLD, genomics the planning of Marine Protected Areas (MPAs) by the Commission for the Conservation of Antarctic is recorded. For records dating from the period 1972–2004 all pertinent botanical journals dealing with trawl_survey Introduction Marine Living Resources (CCAMLR). Belgian flora were systematically screened. Analysis of herbarium specimens in the collection of the Na- Project title: Macroecology of Southern Ocean echinoids: Distribution, Biogeogra- tional Botanic Garden of Belgium, the University of Ghent and the University of Liège provided inter- phy and Ecological Niche Modelling. Introduction With more than 1,500 valid genera worldwide (cf. O’Hara 2011), the Diptera fam- Keywords esting distribution knowledge concerning rare species, this information is also included in Florabank1.

Molluscan fauna of Gueishan Island, Taiwan 13 Macrobenthic communities, trawls, dredges, grabs, corers, direct observations (scuba-diving, sea-bed The data recorded before 1972 is available through the Belgian GBIF node (http://data.gbif.org/ Personnel: Pierrat Benjamin (collection identifier, data collector, data manager, Taxonomic coverage ily Tachinidae represents a good model to which “alternative” diagnostic tools to the Kuroda T (1941) A catalogue of Molluscan shells from Taiwan (Formosa), with descriptions video, sea-bed photography) datasets/resource/10969/), not through FLORABANK1, to avoid duplication of information. A of new species. Memoirs of the Faculty of Science and Agriculture, Taihoku Imperial data publisher), Saucede Thomas (collection identifier, data collector), Alain Festeau The Division of Birds, National Museum of Natural History of the Smithsonian In- traditional dichotomous keys can be applied. We here propose an original interac- University 22(4): 65–216. dedicated portal providing access to all published Belgian IFBL records at this moment is available at: Lai KY (1990) Shells. Vacation Publishers, Taipei, Taiwan, 200 pp. [In Chinese] (computer specialist), David Bruno (collection identifier, general manager). “Fishes of the World” (Nelson 2006) was used as a taxonomic reference for this work. stitution (USNM), has released approximately 2800 DNA barcode data records into tive/multi-entry key web application, MOSCHweb (“mosch” derives from the Ital- Data published through GBIF: http://ipt.biodiversity.aq/resource.do?r=macrobenthos Lai KY (1998) Shells II. Vacation Publishers, Taipei, Taiwan, 196 pp. [In Chinese] Lee CY, Chen CL (2010) A New Siphonalia in the Family Buccinidae from Northeast Taiwan. http://projects.biodiversity.be/ifbl Bulletin of Malacology 34: 45–48. General taxonomic coverage description: The coverage (Figure 1) of this dataset in- the public domain through GenBank and the Barcode of Life Data Systems (BOLD). Lee YC, Chen WD (2003) Land snails. Chin-Chin Publications, Taipei, Taiwan, 287 pp. All data in Florabank1 is georeferenced. Every record holds the decimal centroid coordinates of the [In Chinese] cludes Class Actinopterygii (90%), Class Chondrichthyes (9%) and Class Myxini (1%). National Museum of Marine Biology and Aquarium (2003) Investigation of aquatic biota of IFBL square containing the observation. The uncertainty radius is the smallest circle possible covering the Head Lake and Tail Lake on Gueishan Island. Northeast and Yilan Coast National Scenic Area Administration, Tourism Bureau, MOTC, Yilan, Taiwan, 97 pp. [In Chinese] Lee YC, Wu WL (1998) A new Trochid (Gastropoda: Trochidae) from the Kue-Shan Island, Copyright Benjamin Pierrat et al. This is an open access article distributed under the terms of the Creative Commons Attribution License 3.0 Copyright Kwang-Tsao Shao et al. This is an open access article distributed under the terms of the Creative Commons Attribution License 3.0 Copyright David E. Schindel et al. This is an open access article distributed under the terms of the Creative Commons Attribution License, which Copyright Pierfilippo Cerretti et al. This is an open access article distributed under the terms of the Creative Commons Attribution License 3.0 Copyright Julian Gutt et al. This is an open access article distributed under the terms of the Creative Commons Attribution License 3.0 (CC-BY), Copyright Wouter Van Landuyt et al. This is an open access article distributed under the terms of the Creative Commons Attribution License 3.0 NE of Taiwan. Bulletin of Malacology 22: 57–60. Pace GL (1973) The freshwater snails of Taiwan (Formosa). Malacological Review Supplement (CC-BY), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited. (CC-BY), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited. permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited. (CC-BY), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited. which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited. (CC-BY), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited. 1: 1–118. Wu WL, Lee YC (2005) The Taiwan common mollusks in color. Taiwan Forestry Bureau, Council of Agriculture, Executive Yuan, Taipei, Taiwan, 294 pp. [In Chinese] Wu YH (2002) Handbook of Ecotours at Gueishan Island. Morning Star Publishers, Tai- chung, 285 pp. [In Chinese]

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A peer-reviewed open-access journal A peer-reviewed open-access journal A peer-reviewed open-access journal A peer-reviewed open-access journal A peer-reviewed open-access journal A peer-reviewed open-access journal A peer-reviewed open-access journal A peer-reviewed open-access journal COMPARATIVE Biodiversity

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