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Pseudoscorpionida, Garypidae and Pseudogarypidae) Muchmore, W. B. 1981 . Cavernicolous species of Larca, Aracheloarca and Pseudogarypus with note s on the genera (Pseudoscorpionida, Garypidae and Pseudogarypidae) . J. Arachnol ., 9 :47-60 . CAVERNICOLOUS SPECIES OF LARCA, ARCHEOLARCA , AND PSEUDOGAR YPUS WITH NOTES ON THE GENERA, (PSEUDOSCORPIONIDA, GARYPIDAE AND PSEUDOGARYPIDAE ) William B. Muchmore Department of Biolog y University of Rochester Rochester, New York 1462 7 Abstract Six new species are described from caves in the western United States, as follows : Larca laceyi and Pseudogarypus orpheus from California, Archeolarca welbourni, A . cavicola, and Pseudogarypus hy- pogeus from Arizona, and Archeolarca guadalupensis from Texas . Larca granulata (Banks) is redes- cribed and other species of the genera are discussed. INTRODUCTION Over the past several years I have received for study several cavernicolous pseudoscorpions from western United States which deserve notice and description . Among these are representatives of Larca Chamberlin and Pseudograypus Ellingsen sent by Lawrence A. Lacey, of Larca sent by Andrew G . Grubs, and ofArcheolarca Hoff and Clawson and Pseudograypus sent by W. Calvin Welbourn . In addition to describing th e new forms I take this opportunity to report some observations made on these gener a during the past 20 years . FAMILY GARYPIDAE HANSEN Subfamily Garypinae Simon Genus Larca Chamberlin For a recent review of this genus see Benedict and Malcolm 1977 : 114-118 . Larca chamberlini Benedict and Malcolm A single male referable to this species was found in Dirty Fissure, Calaveras County , California, 26 May 1977, by A .G. Grubbs. This is of the same size as the specimens from 48 THE JOURNAL OF ARACHNOLOG Y Oregon and northern California described by Benedict and Malcolm (1977) but ha s somewhat more robust palps and legs. It is undoubtedly an epigean form, only accidenta l in the cave . It should be noted that this specimen has four setae in the flagellum on each chelicera , not three as mentioned by Benedict and Malcolm (1977 :116) as characteristic of L. chamberlini. If the reduced number of setae is really characteristic of all the more north - ern forms, then this one may turn out to be a distinct species . Larca laceyi, new specie s Figs. 1-6 Materials .—Holotype male (WM 4652 .01003) and eight paratypes (3 male, 4 female, 1 tritonymph) collected in Music Hall Cave, Calaveras County, California, 20 January 1973 , by Lawrence A . Lacey. Types are deposited in the Florida State Collection of Arthro- pods, Gainesville . Diagnosis.—Generally similar to Larca granulata (Banks) and L. chamberlini Benedict and Malcolm with only two trichobothria on the movable chelal finger; but, much large r than those species, with carapace more than 0 .55 mm and palpal femur more than 0 .85 mm in length. Description.—ADULTS : Males and females similar but females slightly larger . With the characters of the genus (see Benedict and Malcolm 1977:114). Well sclerotized and color- ed, palps reddish brown, other parts light brown . Carapace trapezoidal, with posterior width greater than length, surface covered with low granules and with a transverse furro w two-thirds distance from anterior margin; anterior margin nearly straight, slightly rough- ened near middle ; with four strongly corneate eyes ; about 40 prominent, curved setae, o f which six are at anterior and 6-8 at posterior margin . Coxal area typical, widest across 4t h coxae . Tergites 2-8 completely divided, 1 and 9 partly divided, 10 and 11 entire, surface s covered with low granules ; sternites 4-8 completely divided, others entire, surfaces reticu- lated to scaly; pleural membranes longitudinally plicate . Tergal chaetotaxy about 5 :6:10 :10:12 :12 :12:12 :11 :T6T:9:2 ; setae prominent and curved as on carapace . Sternal chaetotaxy of male about 20 : [3-3] :(0)20(0):(0)6(0):9 :9 :9 :8 :8 :7 :6:2 ; all setae small an d delicate ; setae on male genital opercula and internal genitalia as shown in Fig. 1 ; setae on female genital opercula as in Fig. 2; female internal genitalia marked by three well sclerotized cribriform plates in a transverse row and a smaller one behind (Fig . 2). Chelicera small, about 0 .35 as long as carapace ; four setae on hand, es quite long (two setae in position es on three specimens) ; fixed finger with two tiny subterminal denticle s and four small teeth, movable finger with a subterminal roughened area ; galea of female very long (about 2/3 as long as movable finger), straight, and with three small rami near tip ; galea of male short and with variable serrations near tip ; serrula exterior of about 1 5 blades; flagellum of four sparsely denticulate setae . Palp long and moderately slender (Fig .3);femur 1 .52-1 .59 and chela 1 .65-1 .74 times as long as carapace . Palpal trochanter 1 .75-1 .95, femur 4 .85-5 .5, tibia 3 .55-3 .95, and chela (without pedicel) 3.8-4.45 times as long as broad ; hand (without pedicel) 2.35-2 .8 times as long as deep ; movable finger 0.89-0.94 as long as hand . Surfaces of palp strongly granulate except distal halves of fingers ; setae mostly prominent, arcuate. Trichobothria as shown in Fig . 4; fixed finger with eight, movable finger with only two, presumbly sb and b. Fixed finger with 33-38 contiguous marginal teeth, retroconical distally an d becoming flattened toward proximal end of row; movable finger with 35-38 similar teeth . Venom ducts in each finger inconspicuous, nodus ramosus 0 .2 length of finger from tip. MUCHMORE—NEW CAVERNICOLOUS PSEUDOSCORPIONS 49 Legs long and slender (Figs. 5 and 6) ; leg IV with entire femur 3 .9-4.5 and tibia 4.7-5 .1 times as long as deep. Proximal segments heavily granulate, becoming scaly distally . Setae of proximal segments large, arcuate, becoming smaller and straight distally ; subterminal tarsal setae simple ; no noticeable tactile setae present . Arolia twice as long as claws . TRITONYMPH : Much like the adults but paler, smaller, and with less attenuate appendages and fewer setae . Palpal femur 5.05, tibia 3.6, and chela (without pedicel) 4 .3 times as long as broad . Movable chelal finger with two trichobothria as in the adult, fixe d finger with only seven, apparently ist missing. Chelicera with four setae on hand ; flagellum of four setae ; galea long and trifid at tip as in female . Measurements (mm).—Figures given first for the holotype male, followed in paren- theses by ranges for the adult paratypes . Body length 2.11(2.05-2 .39). Carapace length 0.56(0.56-0.62). Chelicera 0 .21(0.19-0 .22) long . Palpal trochanter 0 .33(0.33-0.355) by Figs . 1 - 6 .-Larca laceyi, new species : 1. male genital opercula and internal genitalia ; 2, female genital opercula ; 3, dorsal view of right palp ; 4, lateral view of left chela ; 5, leg I ; 6, leg IV. 50 THE JOURNAL OF ARACHNOLOG Y 0.18(0.17-0.20) ; femur 0.89(0.865-0.96) by 0 .17(0.16-0.19) ; tibia 0 .71(0.69-0.76) by 0.19(0.185-0.21); 'chela (without pedicel) 0 .96(0.97-1 .06) by 0 .23(0.22-0.27); hand (without pedicel) 0.51(0.50-0.57) by 0.19(0.185-0.235) ; pedicel about 0.07 long; mov- able finger 0 .47(0 .47-0.525) long . Leg IV: entire femur 0.59(0.59-0 .64) by 0.13(0.14-0.155); tibia 0.445(0.445-0.47) by 0.095(0.09-0.095). Tritonymph: Body length 1 .65. Carapace length 0 .52. Palpal femur 0.71 by 0.14; tibia 0.555 by 0 .155 ; chela (without pedicel) 0 .86 by 0 .20 ; hand (without pedicel) 0.46 by ?; pedicel 0 .06 long; movable finger 0 .41 long. Etymology .—The species is named for Lawrence Lacey who collected all the speci- mens. Remarks—Larca laceyi is definitely modified for cave life and is probably a troglobite . It may be compared with the local epigean form, L. chamberlini (see above and Benedict and Malcolm 1977) : it is considerably larger than L. chamberlini; it has six setae at the anterior margins of the carapace, rather than eight ; the palps are longer with reference to the carapace ; and the palpal and pedal segments are more slender . Like L. chamberlini, this species has only four setae on the cheliceral hand, whic h probably indicates a close relationship between the two . Also found in Music Hall Cave is a new species of Pseudogarypus, described below. Larca granulata (Banks) Garypus granulatus Banks, 1891 :163 . Larca granulata; Chamberlin 1930 :616, Hoff 1949 :447, Hoff and Bolsterli 1956 :163, Hoff 1958 :15 , Nelson 1975 :282 . The type locality for this species is Ithaca, Tompkins County, New York . It has also been reported from Illinois, Tennessee, and Michigan . New Records.—NEW YORK: Albany, Cattaraugus, Genesee, Monroe, Schuyler, and Wyoming counties . NEW HAMPSHIRE: Sullivan County. PENNSYLVANIA : Lycomin g County. WEST VIRGINIA : Greenbrier, Mercer, and Pocohontas counties . VIRGINIA: Giles County . NORTH CAROLINA : Jackson, Macon, and Transylvania counties . KAN- SAS: Lincoln County. Inasmuch as no complete description has ever been published for this species, the type species of the genus, it seems appropriate to present such a description here. It is based mainly on the three syntypes (1 male, 2 females) in the Museum of Comparative Zoology at Harvard University, of which a female (WM1216 .01001) is selected as the lectotype ; also considered are a number of other specimens from upstate New York, within 100 miles from the type locality. Descriptions.—Males and females are quite similar except for a few sexually dimorphi c characters and the slightly larger size of the latter .
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