J. E NT OM OI.. Soc. BllIT C O L U ~I B IA 79 (] 982), D EC. 31, 1982 37

Plank. 1-1 . F. 1922. The blackh eaded firc\I'orm of cranberry on thc Pacific coas t. U.S. Dept. Agr. Bull. No . ] 032: 1-41. PowelL J. A. 196-1 . Bi ological and taxonomic studies on tortricine . w ith references of the species in C ali fornia. Bcrkcle\' and Los An gclcs Uni, . Calif. press . 3 17 pp. Rainc, J. 1966. Life histon ' of Dasljstollla w lieel/tllll (I-Ibn. ) (Lcpidoptera: Oeeophoridae). a new pest of bluebcrries in British Colu mbia . Can . Ent. 98:.'3.'3 1-334.

A LIST OF PLANT -FEEDING LEIDOPTERA INTRODUCED INTO BRITISH COLUMBIA DAVID R. GILLES PIE AND B ET H I. GILLES PI E Rescarch and Plant Quarantine Station, Agriculture Canada, 880 1 East Saani ch Road. Sidne)', British Columbia. and 2523 Meado\dand Dri\·e. Vi ctoria, British Col umbia. Respecti ve'" ABSTRACT Each of 48 species of pl ant-feed ing known to be foreign to North America and to occur in British Columbia is briefh summarized \I ith its histon ' and importance. T hose feeding on stored products ~ r e not indicated. .

INTRODUCTION T he criteria used here to incl ude spccics in the The foll owin g are species- based summari es of list were rather strict. O n,," those species are incl ud­ the history and importance of exoti c Lepidoptera in ed for w hich definite records were located , giving British Columbia . W e hope that this paper \I'ill exotic origin and date of entry or first capture. sti mulate further im'esti ga ti on of some of these There are probably man\' introduced species of species; particularly useful would be information on Lepidoptera not included here for which no such the adaptation of nati ve parasites and predators to records ex ist. the exotic spccies as hosts. Thc species are treated al phabetical'" in order TABLE 1. Abbreviati ons of PrO\'inces and States us­ of their generic names . Approved North Ameri can ed in text . English common names (Benoit 197.5, Sutherl and CANADA 1978) are included \I· here a,·a il able. Where no such Place Abbrcviation common names are a\'ailable, we ha\'e used British Alberta Alta . common names. as in the reference material. British Columbia H.C. Species of Lepidoptera feeding on stored pro­ Manitoba Man. ducts are exluded from the list because they arc vir­ New Brunswick :\ .13. tuallv cosmopolitan, often independent of cl imate Newfoundland NOd . and can be regarded as being native to man's stored NorthWest Territories i\.W .T. products. wherever such products occur. Nova Scotia N.S. Limitations of time and resources imposed so me Ontario O nt. restrictions on the scopc of these summarics . They Prince Edward Island P.E. I. are not complete bibliographes of each spccics. but Quebec Que. rather brief notcs on the histor,·. distribution. host Saskatchewan Sask. plants and pes t status or potential in J\ orth Am eri ca Yukon Yuk. and elsewhere. Standard abbreviati ons of C anadia n UNITED ST ATES provinces and postal abbreviations of the Un ited Arizona AZ States (Table 1) are used throughout. California CA It is noteworthy that in the t\\"o successful Connecticut CT eradications of introduced Lepidoptera in B.C. , en­ District of Columbia DC tomologists were awarc of the presence of the in­ Idaho lD sects before the pests had had a chance to spread Maine ME from their origin al entry poirits. I n both cases pro­ Maryland MD mpt and perhaps radical action preH'nted the Massachussetts MA establishment of breeding populations. It is also Mi chigan MI noteworthy that 19 of the 48 introduced Montana MT Lepidoptera are of the famil,' Tortricidae. This New Jerse\ NJ disproportionate representation might indicate an, New York NY as yet unknown, biological feature predisposing the Ohio OH Tortricidae to being imported , or to establishing Oregon OR once imported . Pennsylva ni a PA 38 J. E:-:TO \I 0 I.. SOC . BIl IT. CC lI .D IIl I" 79 ( 1982). DI-:c . 3 1. ]()82

Virginia VA Allllr.,ia lil/ ea/el/a Zeller. Ceicchiidae. the peach \Vashi ngton \\'A t wig borer . First ohsen'ed as a pe,t in :\Y in IS72 b\' G lo\'Cr Acleris (, fI/lIl1rimlf/ (Zl'l"'r) . Tortriciriae. a (lS7-1 ) hut descrihed 12 \'Ca rs earl ier In ' Clelllcns stra \I' berI,\, lea fr ai ler. (Li nter IS82). It is fo und in the U.S. \\'hcrc\'l,r First r ~po r ted in North A lI1 eriea ill I !)72 as a pest peach('s arc gro\\'n (j ones 19305) . It has been known of straw bern in Richmond. B.C. (Cram lD7,'3 ). It as a pest in B.C. since 1908 \\' here it feed., on tender has nO\l' be~n recorded on stra\\' lJl'rn·. ra.'pbe rn·. shoots a nel frlli t of peaches. aprieots. pluln and blackhern' alld Spirea in the Lo\\'er F ra.se r VallC'\ or prune (\ladsen & Arrand 19(05). It is a lso found in B.C. (Cillcspi,' I!)SI ). It has not been recorded fmm ant. (Dustan 6: Da\'ic!so n 19(3) but as of 1933 it elsc\I·here in North Alllerica. had not heen reported as being cOlllllll'rc ialh' im­ It is found in E urope. Chi ll a. and Japan. and is portant the re (Hoss & I'lI tnlan 1933). und 11 0 reccllt apparenth' nat i\'e to [ ump,' (Bradle\ ('I 01. 197,'3 ). It references \I'(' re found relating to its stat li S the re . is a Illinor pest of stra\\'berry in England (Alford It \I' as proba bh' a nati\'c of Asia. the o ri ginal 1~)7,s. Im6: Vernon 1!J7!: Petherbridgl' 1921)) alld hom,' of peaches. (nuhman !!l21» but is now found IIungan' (Balazs & Bodor 196!)). I'o lelllilia in t,'mpcrate zoncs of lI10st continents being par­ palllsll'i.\ (1.. ). the marsh cinquefoil. " the main ticllia rh ' \\'cll established in thc ~I cdi t l'l'ranean \I'ild hos t plant in England (VerIlon ID71 1. (Talhot'lk 1!l77). It is a serio", pest of peach a nd a prieot fru ih (Ltlho uk 191i: F ri lli 1!J77) a nd a A d"ri" I'orieg{l//(/ (Denis & Schi rf,'rlll ld ll'r). m inor pcs t of almolld and plum (Taillolik W" ). Tort ricidac First recorded in North America frolll a sin!!le A llif//Ils plf/gif/ /a (I .. ). C('oll1et ridac . specimen collected at Vi ctoria. B.C. ill 1D20 Deliberate\' introdllced into the J..:ettiL' Valle\·. (Blac km o rl' I D23 ) . Fo rbes ( I D2 :3) recordl'd B.C. in lOG, 'as a hiological control agent against specimens frulI! NY. In IDSO and IDSI it \I' as lIoted the \I'ced lIypc r i('ulII Iwrio/'(/lu lIl L. (51. Johns\\'ort on bllll'berr\' in the Lower Fraser Vall('\·. B.C. or Klamath \I'ced) (Harris el al. 19(9). It \\'as not (Gil les pie 19S1). In \H'ste rn North Anll'ric,; it 110\1' rl'cO\'('rl'd in 19G8 (il arris & Pcse hken I!JI I) or occurs in B.C., \VA. on and C A (Po\\'dl 106·1). sll bsl'll'l('nth·. Food plants in No rth America arc spL'cies of Hosa. It is natiH' to \\'cstern and Central Ellro pe Pnillll" . MaillS. Cralacg ll ' alld \ 'acrillilllll (1'0\\'(,11 f('cdin !! on IIYJl criClllu spp. (Johallsson ]()62). 196-1 ), Distriblltion ahroad is [llrope. Nortl, -\l','s t :\ r('lIil'" I'fl r/WU/ (Scopoli ). Tortricidac. Africa , Asia \Iinor. Central Asia and C hina Fir,t rccorded ill North America in H).37 frolll (Razo\l'Ski l!l()f» ). Apparcnth' a rl,lati\'('h ' eonlinOIl Vall('I "IH'r. B.C. and SH·al nore. ant. (F rt ','man and pol\'pha!,(olls spl'cics in Ellrope (Ilradk-\ cl ai, 19.51» . it \I'as rc ported fe~d i ng on A/llelalld/i{'/' sp. W (3 ) . at \ 'anCOU \'e r ( ~la c :-':a\ ' 1!l.')-1 ). It is not of pest .·\ ('(111'.' l'!I/illll/{1 (1Iullller). Phaloniidae. pale jllnipl'r .... i!...!lIific allL'L· in the Lowe r F ra :-. cr \ 'allp:: Do!!anlar W t' !J\\,O rlll . and Reirne (19,8 b) fOlll'ld it tn bc uncommoll Oil First I'('cordl'ci in l'asterll U.S. in lil7i1 makl'r apple. 19(2). Hccnrded at Field . H.C: som"tin )(' h,.fof(' It is found in Ellropc. Asia \ Iinor. South Russia. Wf).5 (Prentice el al. HJ 6.S). I t has I)('('n "b,,'rw'd Eastern Siberia to J apan (Brad lc\' cl al. I !J7.'3 ). h)od fepelin g Oil junipers and ()tl1('r o rna mental .shrlll), in plants are a \\'ide rangc of deciduous trees and the LO\l','r Fraser Valle\' a nd on \ 'anCOU\'l'r Island. ,hrllbs. and occasional II' conifcrs (Hradle\' ('I 01. H.C. Elsc",here in . :urth Anll'rica it has hee ll 197.'3 ). In Europe it is CO.-IIl1lon in fruit orch;lrds and recorded frolll :\.S. to Alta. alld in the northeasterll is a pest of uppil' (Bradl,'\' ('I al. !!l73: Bo nc" I !)76: U.S., and 01 1 (Freeman 19f)7: Baker I!J(2) feedillg Krako\\'lak I !l7·1) . on juniper e" pecialh· jllll i pcl'lls CO/l/l11l1l1is. It is a Palaearcti c sl)('cies (Bradl,,\' ('I 01. 1 D,:3) record,'d a., .\r('lr i"., ro.' Wl/l8 (L. ). Tortricidae. the European fceding on juniper in the' Old \\'orlel. leaf rolle r. Present in castern U.S. b\· ISBn (Comstoc k & Sl­ "\ lIIpllip'lrtl Imgflpillll., L.. :\octuidae . the mOIl Sl'. ingerland IS90). First fO llnd in Canada in 1919 A possihle "introduced " accordin!! to simultan('llII.sh- in Victoria. B.C. (Black more ID2 1) Ma\\' ( I!),C;) ",Iw gi\'Cs its range in :\orth Alllcrica as and I S. (G ihson 192-1 ). Found in Vanc:o u\'Cr in Canada and southern I'A . Appa rl'nth- till' onh' HJ2:3 (Trcherne H12.3 ). Common on apple recorc! of this species in B.C . is on apricot. pllllll and (Do!!illliar and Beirne I B'Rb). bhll,bl'IT\' and rose a bout W3i1 . probahly in the Fraser \ ' alle\' raspberr\' !Cillespie HJS I) in thc LO\\'L'r FraS('r (Lle\l el lyn-Jones HJ38). It has hel'll rl'corded frr)!u Valle\ and in the Okanagan \ 'alle\' as a pe., t on ap­ alfalfa in NY (\\'], eder 1975) a nd tobaeco in :\ .S. ple ( ~I a d s en ('( al. 1!)77). Its distribution d se\d1('re (Specht I H/2). in 1\orth Aml'rica is N.II .. Ont .. CT. :\Y. 1\J . \\·A. A Palaea rclic specics (1\l aw W7G). reporll'd a.S a alld on (Frceman I!J,sS: Chaplnan & Lienk 19,1 ). minor pest of stra\\'hern' in Ellrope (Aifnrcl Im-1 ) It ha.' Iwen recorded as a pest of fil bcrts in the U.S . and hern' crops in the USSH (M usich W7G) . (Al i;\Jiaz" l' HJIi) and cllrrant in N .S. (\\'hitclll'acl J . E:,\TO~IOI . Soc Bil l"!" COI .l·\l Il I.-I 79 ( 1982). Dlx . :3 1. 1982 :39

1(26). and has abo been recorded from pri"ct recordcd frolll e1 se" 'here in :'s'orth An1t'rica (I lodges (Chapman & Lienk WII ). 1 97~ ) . Occu rs ,,·here deciduous trees and bush frllits It is a I'alae.lrclic species (\tt'nick 192,: Ilotit!es an: gn)\\"Il in Ellrope. the principal food plants be· W7~ ) ,,·he re it has bet'n rccorded as a pest of roses ing apple. pear. l1

(Andiso n 1937). In B.C. A native of E urope where it is a pest of la rches B,C , near Vi ctoria in 1034 of common and ornamental (Dawson 1971a). it is a minor pest junipers on the so uthern half of Vancouver I sland, cent mainland, and in the southern in­ Coieop/wro serralella (L.), Col eophoridae, eigar and the adja n 1937; Bauman & Sugden 1969), In casebea rer. terior (Andiso s also found in the East and Unintentionall y imported into Ont. on orchard North Am erica it i s (Nordin & Appleb\' 1969) and stock about 1885 (B. Wright, Curator of Zoology, Midwest State Canada (Bauman and Sugden l!J68) N .S. Museum, 1747 Summer Street, Halifax, southeastern crn Ont. and Que. (Prentice e/ ai. Canada, B3H 3A6, pers. comm.). [n Canada it has including south s been recorded a s a serious pest of been coll ected in N.B., N.S . and southeastern Onto 1965), It ha tern North Am erica (Nordin & Ap­ (P rentice el ai. 1965), where it is common on Fruit juniper in eas trees. It is principallv a pest of apples in the nor­ pleby 1(69). a Palaearctic s pccies found in Europe and thern U.S. but food plants also include cherry. It is (S taudinger & Rebel 1 901 ) fceding on hawthorn, plu m, quince and pear (Baker 1 972). It Siberia ss & Lott 1922). arrived in B.C. recentl y, probabl y in the 1960's (B. juniper (Wei Wright, vide Slipra, pers. comm.). lls/iom,U/ (llaworth), Tortricidae, the It infests apples and allied plants in E urope Dill/ill {l1Ig apricot or vine moth. (McDunnough 1(57) and occasionall y causes serious small st r eported in No rth Am erica at Victoria, injury to fruit trees in central Europe (Thiem 19~ 3). Fir B,C. in 1924 (Clarke 1930) and latcr at Nil es, CA, 1932 (Keifer 1(33) . It has no t b eco me generally Croesia /wimia"a (L.), Tortri cidae. in in North America, a nd it is not known if First recorded in No rth Am erica from the common lations still ex ist in CA (Powell 1(64), Lower Fraser Vall ey, B.C. , in 1977, by Doganlar popu It is native of Europc and is distributed through and Beirne (1978a) on unsprayed apple and pear e. Asi a, and North Africa (Bradley e / ai. trces. They suggested that it could become a se rious Europ It has been recorded from fruit and hops pest of apple and other fruit trees in B.C. 1973), , from stored apples (Adkin 1(31) and The natural range is north and central Europe (Massee 1946) oak (Matthey 19(7) in England, and Asia Minor (Bradley el a i. 1973) where it is a fro m minor pest (Evenhuis el ai. 1973; Miczulski and /ll/Il(/I/{/ (Treitsehke), Tortricidae, green Koslinska 1976). Food plants in Europe arc El'i,wlill ce leafminer. Cara/aeqlls. H osa. Hllblls. MaillS. I'lJrIIs. and spru rded in North America in MA in 1907 Cydonia spp. (Bradlev e/ ai. 1973). First r eco (Kearfoot 1907), Rcported in B,C. from a single record on an undetermined date before 1965 (Pren­ ,,"'

E/iella zinckcllella. (Treitschke). Pnalidae, lima­ the peach growing districts of the U.S. (Chapman & bean pod borer. Li enk 1971). According to Handford (1955) it has been It was brought into B.C. at Su mmerl and, in recorded from B.C. although the original record 1956. in infested fruit from \V A for canning could not hc located . Thcre are no subsequcnt (Touzcau & Neilson 1957). It was suceess full ~ ' records of this species in B. C. The onh' other record eradicated b,' 19.58 by heavy spra"ing, fumigation for Canada is one from Oxbow, Sask. (lleinrich of the cannen ' and removal of the orchard adjacent 1956) probabl y in 1955 (Mac:\a,' 1961). According to the canner)' (Touzeau & Neilso n 1958). to Pierce (1917) it was in thc U.S. by 1917, and was It is found in South Am crica , Europc, Asia, and at that time vi rtuall,· a cosmopolitan pest of peas Australi a (Anon 1951-19.52). It is a serious pest of and beans. peach , appl e and other fruits where\'Cr it occurs It is w ideh ' distributed in the world , found in (C hapman & Lienk 1971). the tropical, s ~btropi ca l and warmer regions of the temperate zones of Asia, E urope, Africa , Central lIedia 'lIIhife,.,,"lI (H a\\'orth), Tortricidae. the green and Sou th Am erica and Aust ralia (Hei nrich 1956; budworm. Naito 1961). It is a pest of peas and beans and feeds First recorded in North Am erica in 1913 from on other leguminous plants in th ese areas (Naito N.S. (Sanders & Dustan 1919) . Found in B.C. at 1961). Hatzic in 1 9 1~ and at Vernon in 1919 (Treherne 1921). It was found . but not common on apple, in FlIrrhypara IlOr/lI/a/a L. Pnalidae. the Lower Fraser Vall e,'. B.C. in 1978 and ]979 bv Fi rst reco rded in North Ameri ca in 1907 one of the authors and b,' Doganlar and B e irn~ (Munroe 1976) . It no\\' has a wide range in the nor­ (1978b). It was reported from Onto in 197,3 theast, ex tending from Nnd. to Ont. and (C reelman 1973). It has al so been recorded fro m south\\·ard. First recorded in B. C. in 1977 (Lazorko P.E. !. and wideh ' in the northeastern U.S. (C hap­ 1977). Little is kn own about food plants in No rth man & Li enk 1971). It has been recorded as a pest Am erica but potential hos t plants cu lti va ted in of applc in "'.S. (MacNay & Creelman 19.58) . ga rdens in B.C. are Stachy' recta. Alel/tha sp .. It is a common species in Europe and Asia Minor Ca/ystegia .\'epillm and Rifle.l· sp. (Lazorko 1977). (Gibson 1923: Chapman & Li enk 1971 ) where it It is native to Europe and temperate cast Asia feeds generall,' on Rosaceac including apple, (Lazorko 1977). European food plants include net­ hawthorn and species of Pi'll II liS and Sorhll .\' (Chap, tIe (Urt ica diocia). w hite horehound (Marrtlhillll/ man & Lienk 197 1). vulgare) . wou ndwort (Stach!), sp.). mint (Mentha lIelllithea aes/iw';(1 (Hubner). Geomctridae. the sp. ) cu rrant (/lihe.\' sp.) and other plants (Munroe common emerald. 1976). First reported in North Am erica in 1978 from E utromllia fllemerophi/(lJ parimlll (C lcrck). Glv­ New Westminster, Burnabv and Langley, B.C. phipt e r~ ' g id a e , apple- and-thron skeletonizer. (Doganlar and Beirne, 1979), it has not been First noted in North Am eri ca in NY in 1917 (Felt reported from e!se\\·here in No rth Ameri ca (Bolte 1917) as a pest of apple. It subsequcntl y spread to and Munroe 1979) . N .S. (Leonard 1923) and was found in Ont. in 1929 It is found in mos t of E urope and in Eastern Asia (Caesar 1929). Complaints were first recorded in (Doganl ar and Beirne 1979) and has bcen reported B.C. in 1937 from the Victoria region where it from a wide varietv of wood" trees and shru bs bccanlc a pest on unsprayed fr uit trees CT'\dnn (Doganlar and B e i~n c 1979, Bolte and Munroe 194 1a). It was \\'ell established on the mainland of 1979). B.C. by 194.5 (Twinn 194.5). It is a pest of the coast Lasl'eyresia "igricwU/ (Stephens). Tortricidae. the and occasionall y onc in the interior of B.C. on apple pea moth . (N ei lso n 1957). 'Its i'\o rth Ameri can distribution is in First found in North Arm'ri ca in eastern Canada the apple growing areas of the northeastern U.S. in 1893 (Fulton 194 7). Injury to peas \\'a.s reported and so utheastern Canada, and in the west from in Onto and Que. in 1897 (Fl etcher 1899) and it is B.C. to OR , !D. and CO (Heppner H178). also wid espread in the Maritimes. First notes as a It is a European speeics (lIeppner 1978). It has problem in B.C. in 1933 \\·hen a hean' infes tation been reported feeding on apple. pear and So rblls in occurred on the Sumas Prairie (F ul ton 19n) . It has Fin land with severe damage in some vear (Ekholm been suggested that it migrated to Sumas from the 1958). Also attacking apple and pear in Siberia north\\'estern U.S. (Fulton 1947) . It was res ponsible (Vorzheua 19.58) and doing conspicious damage to for the elimi nation of dried- pea and pea seed appl e in Austria (Bohm 1962). culture in B. C. from 1935 onwards (Beirne 197 1). It Gral'ho/i/ha lll oies/a (Busek). Tortricidae. the orien­ has become established in most of the pea growing tal fruit moth . areas in North Ameri ca (Fulton 1947). Introduced to North Ameri ca in 1912 or 19 13 at This European species (Beirne 1971) has been Washington, DC (Peterson & Haeussler 1!)26) . First reported as a common and important pes t of peas fou nd in Ont. in 1925 where it is a serious pest of grown for drying in England (S mith 193 1; Gould et peach , is injurious to quince and some vari eties of al. 1962). It infests cultivated peas in Finland and pear and feeds on apples, plums and cherries larvae have also been found on wil d tufted "etch (Dustan & Davidson 1973). It is now found in all (Vicia cracca) (E kh olm 1961) . 42 J . E~TO ,\I() I. . Soc. BIIIT. C()I.I'\IIlI.~ 79 ( 191':2). DEC. :3 1. 1982

l.a'l'eyres;a p()/l/orudla (1.. ). Tortricidae. codling in 1!)22 at Abbotsford. 1l ,C. frolll lI'hi te pine (Fe'lt moth. 1922). i'\ow found in the Lower Fraser Vallel' and Thc exact date of introduction to i'\nrth America Vancollvcr Island (ila nis 10.58). It has also' been is unknown (Chapman & I,i,'nk 1071) hut it \I'a,s recorded from Ont. (Harris 19.58) and Qllt'. (I'ren­ probahh- introduced from Ellrol)(~ \I'ith car'" col ­ tice (' / 01. HJ6.5 ) and \n' (Fllrniss & Carolin 1077). onists. Putnam (190:3) suggcsts that it lIlight IHlI 'l' Illi ning needles of pine. been introduced to Ont. in 16:3.'5. Fletchcr ( II':!)~ ) It is an Iinilllportant pest of Scots pine in Europe report ed damagl' to apple crops in Canada and (lianis 1%1': ), Hetllllne ( 1880) reported it as wide'spr('ad ()I 'lT tilt' IV hole Dominion. First recordcd in H.C. in luno on Ol'erol'h lera iJrrt/l/flla ( L. ). G('onll'tridae. thc Vancoul'L'r Isla nd in IHO,) in the int,'rior fruit gW\I'­ lI'inter Inoth. ing districts (Mc Leod 19'5 [). f\cports fro nl the tinll' First rccorded in North Alll l'rica in :\.S. in l!J~!) indicate that a long and losing hattie was fOll ,ldlt in lI'here it n"I\' hal'e been present for as long as :30 H.C .. \I'ith the pest heing intmdlJ('l'd. eradicated. I'cars (S nIith 19,')0). It has since extendcd its ran,ge and rc- introduced at "'Irious poinh "H'r a Ilt'riod of into l\. B, and 1' . E,I . (P renticc ('/ ai, I !)(i:3: Forbe.s 1'/ I'ears (Anon 19 12: Anon 1!l21: Tn,herne 1!l 1 ~ : Brit­ II I . I !J fi~), First reported in H. C. in l!J7fi froill \ 'an­ tain 19[~ ). In the B.C. Depl. of Ag riclIitml' Annual COU\ 'c r Island. but the olltbrcak startcd earlier Report fo r 19 1-1 (Anon 1D1.5 ) there was optimislJl (Gillcspie 1'1 Ill. 1,) 71'), Found at Po rtland , OR in that the codling moth would Ill' e radicated frolll I !)78 ",hcrl' it appcars to ha lT been localll' establish­ B.C. the follo\l'ing I'ear. a hope whieh i, still l'\­ ed from at least I!J.5 R (Ferguson 1971'). AI ~o found at pressed from time to timel It nO\l' ()('Cllrs \I'lll'rl'H'r Vancoul'l'r. \\'A ancl on San Juan Island (Ferguson apples are grown in Canada and the U.S. (C hap­ !H78). In the \laritilllcs and B.C. it is a scriol ls man & Li enk UJ7I ), dl'foliator of forest. urban and orchard trees and Probabh- nativc to southern Eurasia it is nO\l a shrubs (Gill espie & Finlal'son l!ll':!: [lIlbre(' 19(7). pest of app(,' whercI"'" thl'I' arc grown \I'ith the l'X­ It is natil'(' and \I'icl espread acmss northern ceptio n (as of UJ7I ) of J a pan and parts of mainland Africa and temperate Ellrasia (Ferguson I !l7R ). I J1 Asia (Chapman & l,ienk I !)71 ). Eurasia it is all important bllt cI'clical defoliator (Sechser 1970) of fnl it and decid ll ollS forest trees /.().r()8/ege "/;('/ ;ml ;,, (L.). !'lTalidae. till' beet (WI'lie H)()I ) and is rccorded as attacking the fruit \I'e!J\I'orm. itself (Briggs 19.57). This species nHII' hal'(' heen introduced into l'flllI/l'lII;" (,,'m.wllla I lubne r. T o rtrieidae. I\orth America from Asia (Beirne I !l(1 ). h,rl)J's First found in :'\ orth America at Saanich. H.C. (1!l2:3) suggcsted that the first introdllction \I 'as pro­ in !9fj~ ([I'ans I>loG). It has spread to the Low('r babh- in eastern North Aillerica. Hecorded in \ Ian, Fraser Valicl' \I' he re it was common on applc in in 100:3 (Anon ID0:3 ). it has hecn rccordcd as callsin,g 1977 (Doganlar and Beirne 197Ilb). In B.C. it feeds damage some\l'herc in the Prairie prol'inces l'l en' on a I'arietl' of hos ts including ()IIl'rCIiS. '\/0 111 8. ~ ' e ar since I II 17 (Beirne I nil ). esp('cial'" in rape. IIIIIIIS and A('('r ' 1'1'. (EI'ans 1970) and hlll('herl'l' Oa, and Sli gar beets (e .g. Cibson 1921: \ lcDonald and raspbl'rn' iu the LO\l'cr Fraser Valley (G ill es pip 190:3: Beirne 1971 ). First kno\ln danlage in Ont, in 1981 ). 19:36 (Caesar 1!),37) and in B.C. in HUI': (Twinn It is a P"lal'arclic spl'l'ies (Bradle\' c / 01. 1m3) 1939). It is considered a pest in the dry interior of prorllincnt in the European o rchard pest control B.C. as wcll as in sOlltlll'rn \ lan .. Sask. and Alta. literatme (e.g, :\ordlandcr [!)I7: Kole\' and Balcl'­ Food plants indude a larg<' I'ariet\' of plants in ­ ski I Dl8). cluding forage and trJIck crops lI'itli a preference fo r broad Ical'Cd plants. (illll I1rOt' 1076). \Ill nroc ( I DI(;) P(IIIt1elll;8 hel'fllw/{/ (Denis & SchifferrIlJIll er). Tort­ gil'cs its present I\orth Amcrican distril)JJtion as ricid ac. "common in northeast, ranging across Canada and First rccorded in North Amcrica in the I,o\l'cr U. S. frolll 1':.5. and \ 'IA to tlie sOlI tlwrIl :-':.\\'.T. Fraser Vallel', B,C. in 1978, but mal' hal'(' I)['en in­ and Yuk .. and B.C. and AZ ." Ikcent'" it has caused troduced n;J1ch ea rli er (Doganla'r and Beirne complete loss of sugar beel. alfalfa ;Illcl I'egetable­ 197!l b). Lan'al' \lcn.' collected from apple . crabap­ crops in somc areas in :\orth AIlll'rica (Munroe pic. pear. piIlnl and species of PI'IIIIIIS. Cro/OCgI18 . 1970). LOllicero. lllliJllS. \'11('('illi1l111 and Spirco (Doganlar It is a pest in tlie !'alaea rctic rcgion. lI'ideh' and Beirne 1979b). d istributed particularh' in drier areas (" Ilinroe It is fOllnd in Europe, the i'did-l'ast. Siberia, 1976). It was rcported attacking rape in Bul garia in Korea and Japan feeding on \'arioll.S trees and 1977 (Po pOI' W71l ) and is a seriollS pest of past JJl'C shrIJbs including species of ;\IlIllIs . 1'1'1111 li S. Pljl'll.\ . grasses and legullles. as lI'dl as peas. cereaLs. beets Tilia . Salix. L OII;c!'/'({. Be/lt/o. Rihcs. \'o('eillilllll and other erops in the USSR (11e10l' £'/ 01 . I !Jill). and JIIyri('({ (Bradlcl' ('/ al. 197:3). In Europe it al so occasionall\' damages apple fruit (Adkin 1 92~ ). ()ClleroS/Olll(l 1';1/;(/1.;,,1/(1 Zeller. Yponolnelltici

Ladner. fl.C. (Doganlar & Beirne ][),Dh) on !lo.\"([ arc l(rO\\'n aod more Or less se\'ere damage ill man\' spp. It is a European specit·s not recorded pn" 'ioush' \·ears. especialh' in southerh- districts (Beirne 1971 ). frOll1 :\orlh America. . Ilareou rt ( I !)()'2) stated that it can cause seH'rc It is ,,·ideh· distrihuted and conllnon ill Europe economic damage to cahbage ancl related c rops and parts of the \1 iddle East. It is recorded as throughollt !\'orth Amel·ica. feeding 0 11 a " 'ide range of ,,·ooth· o rnamental I t is nearl\- a \\"orld"'ide pcest of c rll c iferolls plants. fruit trees. and decidllolls forest trees plants (I'fadt 19, I). (Dogaular & Beirne I H'Da). 1'11I1ell" x'floslell" (L. ). )\·pononlt'lltidae. diamond­ back moth. I'erir! m /l w W ll ci(l (H ubner). "octuidae. "ariegated Introduced to "orth America from Eu ropc' cllt\\·orm. sometime beforc the middle of the 19th eelltlln' \la,' hm'e been introduced into North Aillerica (\letealf & flint 1%2). no\\" oceurring " 'll en'H'r its (fleirne 19(1 ). If so. this prohahh happened earh' host plants arc grO\\"I1. Food plants include all (Beirne ][)71 ). for it is no,,' distributed throughout Cruciferae. a/so sCJl ne ornalllental and grcenhouse til(' U.S. and southern Canada (Ilo li and W6R: plants sl lch as s\\"eet akss uJll. stocks. cand\·tllf!. and Beirne 19(1). In 1900 the first seriolJ) outbreak \\"as "'allflo\\"er (Mt,tcalf & Flint J(62). Hepo'rted frolll recorded from B.C. (Cibso n 191'2). In Canada it is B.C. in 1900 (Anon 19(0) ,,·here is \\"as ,'en' perhaps most important as a pest in B.C.. P.E.I.. destructi \'e to cruciferoll.' crops in part\ of Va,{­ sO llth\\"cskrn Onto and N.S. in that sequence cO llver Island. It ,,'as reported as a pest in (Beirne 1m 1). It has a " 'ide range of food plants Okanagan Val le\' in ID I-I (Buhman IDI.'5). It is a \\"ith su rne preference fo r l(arden crop.' hilt \\"i11 feed sporadic pest in :\orth Aillcrica rcaching high on the fol iage. buds and fmit of trees and \·ines. numbers in limited areas in -'OlIle \"Cars (Da\·id.\on & tohacco. ornamentals and greenhou.\e plants (\"'t­ Peairs ID66: Bt'irne 1971). calf & Flint HJ62). It is comlllon in Ellrope oecasiona lh· occurring It is found in most of the culti\'all'd areas of the in \'ast numbers and damaging cruciferous plants " 'orld (\letcalf & Flint U)()'2: \lorris WRO ) hilt is (Anon Ul:1()) bllt these outbreaks are usualh' confin­ rclath'ch' rare in the old \\"orld (Forbes H) .'i.J ). ed to small a reas. This 1' lII"opean species is 'found in Bibolini (1970). rcported an outbreak in Itah' in As ia. Africa. Australia and :\e,,' Zealand as " 'ell as 19G6 on lettuce and tomatoe\ ill l(reenhouses .. Ameriea (S mith ID:3I ). I'hIJ",rilll(le6D llolland ( 196il) suggests that it "as prohahh- in­ (Baker 1972) fr om France. SC'\'ere defoliation of troduced from Australia. It has been recorded shade and fru it t·rees occllrred in the Ne,,' England damaging potatoes in \ 'A in field and storal(e States and despite attt·mpts to pre\"Cnt i., spread it (Spencer & Strong 192.5). It \\"as fC!lmd damHl!in.l( no\\" oecms in :\Y. " J. FA . :-'11 and southeast Que. potatoes in Duncan. B.C. in HJ5R (Arnott & Anand (Raker 19(2) and is still spreading. In eastern "orth H)60) and at t\\"o IIn,sl)l'cified points in J!)(jfj in AnH'rica it fecds on most hard"'oods and S(Jll le in­ potatoes for the chipping indl lSt lT (Creelman 1!)(jEi ). termixed conifers (Fmniss & Carol in HJIi). Aecep­ 1\:" r('corcis ,,'ert' fount! of it ,"h"'quenth beinl( prc­ table food plants inchldc oak. ,,·illo\\"s. poplars. sent in R.C. most birches. larch. linden and apple. First It is a pe;t of potato('s in AIl(iers and other threatenecl \\"estern Canada in 1911 ,,·hell it \\"as in­ .\l edil<"rranean countries (Ilolland I U(iil). I mills tercl'pted at \'anC(III\·l'r. fl .C. (Anon 1 DI9a) . ( ID(;.J) states that it is a " 'ide,preael Ill'st of stored DiseO\'eret! in 10,1> in the J..: itsi lano a rea of \'an­ potatoes. more rareh- affectinl( the field crop and coun'r e\·identl\· fr om eg.l!s that arri\'ed on a canoe parlicul arh ' in ,,'a nner countrie.\ (joncs & Jones from Qlle. (Anon 1()79a). It set'lllS to ha\"(' been 19(;.J). It has bcen reported as a pest from 11Ial1\' eradicat<:d frolll B.C. b\· the search alld destruction areas e.g. Rhod('sia ( ~Iitchell W7ill. l'e1"ll (Arestel!l;i of egg masses and I,,· spra\'ing \\" hen permissio n" as 1977) and Ne\\" Zealand (Foot W7D). on potatoes. .l(h·en b\' land()\\"Ilcrs (T. Finlanon. Prof Emerita. Simon Frasc'r LiniH·r.sit\". Burnal,,·. B.C.. V(iA IS6. and in El"ypt (S haheen W"I. 011 tomat","s. pers. cOJll m. ). It is a potentiall\- .~e riou s pe., t ,,·here

Pieri., ral'''{, ( L.). PierichH". till' i III ]JIJrtecl introduced and has .J.')O knO\\"Il host plants (Anon cabbal(E'\\"orm. Im!)a). It \\"as recorded in CA in 1970 and 1973 First introduced to :\orth America ahout lil tiO at (Furniss & Carolill ID77 ) ancl an act i\'e inlestation Que. (Chittenden 190.5: Pfadt 19(1 ). Spread rapid­ " 'as found there in HJ7H (Anon lD7Hb). HC'cent I~ through U.S. and Canada reachinl( CA b\· lili>:3 reports indicate that this species is no'" present in (Pfadt ID7 I). Obsen'ed in c'usll'rn B. C. in' !I"UI>. \\'A (Anon U)ilO ). reached the Pacific coast in 1901. and " 'as founcl on It is found in temperate regions. ("x[c'od ing Vancouver Island in 1902 (Trelwrne 191.5). In through(Jl lt central and southern Emope. northern Canada an important pest that calI S('S SOllie damage Africa. central ancl sOllthcrn As ia. including Japan ill most \'ears ,,·here\·er cabbal(t, and relateel plal;t, and Cedon (Gerarcli & CriJllIll 1<)79). It is a major pest of forest and o rchards in Yugosla\·ia. Francc . 44 J. E:-JTOMo l .. Soc. BnlT. COLU~ I B I A 79 (1982) , D EC . 31. 1982

Ru mani a and Japan where population peaks occur from O nt. eastward, as well as B.C. (Torge rsen & at irregul ar intcrvals (Marx 1973). Coppel 1965, Baker 1972). Distribution in central Canada is limited by the severe w inter climate RecurlJllria nl:m el/a Ilubner, Gel cchiidae. the lesser (Prentice et al. 19(5). Hosts include a large varietv bud moth. of pines (Baker 19(2) especially ornamental and Recorded in eastern U. S. as earl v as 177A shelterbclt types (Harris & Ross 1975) . It is a serious (Gilliatt 1934). It was recorded in 1915 from the pest of pines in plantations in Ont. (Prentice el 01. Annapolis Valley, N.S. (Sanders 1915) and fr o l1J 19(5), ornamental pincs in res idential areas and Ont. (Gilliatt 1934), and was present in mall\ pine.s in Christmas tree farms (n yan & Medle!' districts in the eastern half of the U.S. (Sa nders 1970). 1915) . Food plants include apple, plum, apri cot. It is a European species injuri ous to PilillS pear, quince, peach, cherry and hawthorn (G illi att sy/ves lris a nd other nati ve pines (B usck 19 14). a nd 1934). It has been recorded as a pes t of apples in has been recorded damaging pine in England , Ger­ N.S. and H.C. (MacNa\' and C reelman 1958) many and Denmark (Crvstal 1937) but is not a although a first record date was not fo und fo r H. C. serious pest in its native rangc (Furniss & Caroli n Native to E urope (Sanders 1915) where it has 1977). been recorded as a pest of apple and other fruit trees in various areas e.g. Bulgaria (N ikolm'a 19(2). the -""ilOlwla ocel/alla (Denis & Schiffermuller), Tortri­ Soviet Union (Matnashvili 1975) and Swit zerl and cidae, the cye-spotted budmoth . (Baggiolini 19(7). First reported fro m No rth America in 184 1 from MA (Harris 184 1). Was very co mmon in 1912 in Vancouver (Wilso n 1912) and Victoria, R.C. (Brit­ Rlwpubola 1I11eV(IIU/ (H ubner). Tortri cidae, the black-headed fireworm . tain 1912) but rare in the Okanagan Vallcy (Brit­ A form known as the black-headed fi rcworm tain IBI2). It is now widely d istributed in B.C .. as was probabl v introduced to W A and O R betwecn well as Ont. , Que., N.B.. N.S. , P .E.I. (MacNa!' & 1912 and 191 5 from cuttings of cranberry from /vl A C reelm an 1958) and w herever deciduous trees are (Plank 1922). It had then been a pest of eranberry in grown in the northern U.S. (Chapma n & Licnk NJ, MA , and WI for a long timc (Pl ank 1922) and 197 1; Baker 19(2) a nd a pest of fruit trees in R.C. was al so present in NY and CA . First reported in (Gerber el al. 1980) a nd throughout its range B.C. in 1954 as attacking cranberry on Lulu Island (LcRoux & Reimer 1959; Prentice el 01. 1965; Baker (Anon 1954), it is now a ve ry se rious pes t of 1972). It is also a pest on blueberrv in the Lower cranbern' in B.C. (C ram & Ncil son IB78) a nd is Fraser Vall ey, B.C. (G ill espie 1981). Food pl a nts di s tribut ~ d throughout the cranberry growing areas incl ude apple, whi ch is the most consistent food of the Lower Fraser Valley (Neil so n 19( 9). It at­ (C hapman & Lienk 1971 ) other orchard trees , tacks cranberry wherever the crop is grown in hawthorn, larch, laurel , oak (Baker 1972) and North America incl uding N .B. and 1\ .S. (Maxwell mount ain as h (Prentice el al. 19(5). & Pickett 1957) , thc eas tcrn U.S. (Franklin 1950). It is a European species (Porter 1924) common WA , and OR (Breakey 19AO, Pl ank 1922). in apple growing districts. Also reported from A form known as the holl y budmoth , (Kearfoof s Pakistan, C hina, Korea, and Japan but not as vet R . lIaevana ilicifnliana ) was recordcd in R.C. in from the so uthern hemisphere (Chapman & Lienk 1923 and is now present in W A and O R (Swenson 1971 ). 1958) . T he holl y bud moth and the black-headed Sliipllotia salici, (L.), Lymantriidae, the satin moth. fireworm are thought to be Ve rY cl osely related First recorded almost simultaneouslv about 1920 (Breakey 19(0). The hoi'" bud moth is an important from the East coast of No rth Ameri ca at Boston, pest in OR holl y plantings (Swenson 1958). MA (B urgess 1921) and the West coast at New Rhopnbola nae v (IIl(/ is recorded feeding on fruit Westminster, B.C. (G lendenning 1924, 1929). trees, holl y and species of C rataeg lls and \fa('ci llilllll Found in the Maritimes in 1930 and Que. in 1938 in Europe (Meyrick 1927; Lucchesse 194 1). (Reeks & Smith 1956). It became a seri ous pest of poplar and cottonwood on both coasts of North RhYlICiollili buolillnll (Schiffermuller), Tortricidae. Am erica (G lendcnning 1929; Reeks & Smith 1956) the E uropean pine shoot moth . and by 1956 in the interior of B.C. (Condras hoH First recorded in No rth Am erica in 1913 at 1956).' In B.C. it is now found on Vancouver Island . Great Neck, NY (Husck 19 14 ). First recorded in along the coast and in to the southern interi or B.C. at a nurserv in Victori a in 1927 (Downes 1928) (Dawson 197Ib). It has also been recorded from but eradicated by destroying pines (Mathers 1940). Nfld ., N.S., N. B., eastern Que. (Prentice el 01 . Recorded at Vancouver, B.C. in 1938 (Mathers 19(2) and the New England States , W A ancl on 1940) it is now established in the Lower Fraser (Baker 1972) . It is an occasionally seri ous pest of Valley on southern Vancouver Island and the Gulf poplars, trembling aspen, and cottonwood in North Island; (Prentice el al. 1965: Harris & Ross 1975). Am erica (Dawson 1971b). It ma!' also feed on Transport of infested nursery stock res ulted in rapid willow, oak, crabapple and Saskatoon berry spread of the moth which is now found in WA , on, (Amelal/ chier) (Lejeune & Sil ver 1961 ; Dawso n CA, through the northeastern U.S., and in Canada 197Ib). J. ENTOMol.. Soc: BHIT COI.L ;.\ I BL~ 79 (1982). DEC: 31. 1982 4,5

It occurs \\'ideh' through Europe and Asia (Le­ rhellalcl'illlll elalills (I. ) and n ('s lll('a sp. (Carl jeune & Silver I 9U I; Dawson 1971b) and is a 19(8). sporadic pest of poplar and willow throughout its range (e.g . DeMcirlcirc 1966; Kailidis 1964; Molis Tyri(l ja('ol}(lI!lIe, Arctiidae, the c innabar moth. 1070). I ntentionalh' introduced to the Lower Fraser Valley, B.C. in 196,5 as a biological control agent Thyme/iells lincoln (Ochsenheimer). f-1esperiid ae. against the weed SCl/ccio iaro/;aea L. (tans\' European skipper. ra",'\\,ort) (Wilkinson I B6:5). The first introduction to Introduced to North Ameri ca a t London, Ont. B,C. did not establish but later introductions have sometime before 1910 (Saunders 19 16). In eastern (Harris el al. 19 7.5 ). It has also been introduced to North America it is now found in Que .. P.E.I. the Atlantic prodnces (Ilarris pi al. U)7.5) and the (McNeil 1978), Ont. , N.B , ML OH, NY, PA, MD, western U.S. w here it causes extensive defoliation :\J, CT (Burns 1966), Nfld. (Jac kson 19(8), N .S. (Hawkes 1968) of the weed it was introduced to and Man. (Preston and Westwood 1981). It \\'as control. first found in B.C. at Terrace in 1960 (MacNa\' It has been intentionalh' spread from its native 1965). This colonization ma\' ha\'e arisen from a Europe (M\Tes & Campbell 19(6) to New Zealand new introduction from abroad (Beirne 197 1). (Cameron 1935) and Australia (Schmidl 1972), for Preferred food plant in ;\orth America is timo thy, the same purposc . Phlelllll pralellse, but several other grasses are readil v eaten (Burns 19(6). It has reached local ACKNOWLEDGEMENTS population densities in eastern Canada high enough W e thank Dr. B. P. Beirne, Profcssor of Pest to become an econom ic pest on ha\' (Pengellv 1961: Managcment and Dr. A. L. Turnbull. Professor, Arthur 1966; McNeil 19(8). noth of the Dept. of Biological Sciences, Simon A widely distributed Palaearctie butterfly Fraser Universit)" Burnab)', B.C. for allowing us (Bergmann 1952-1955), it is restricted in Europe to access to their literature files. \Ve also thank Dr. headlands, forest borders, river banks, roadsides, Beirne for helpful advice and cditorial cOlllments. pasturelands and other places not IIsed agricultural­ W e thank Prof. Thelma Finlayson, Prof. Emerita, h · and is rare and of no economic importance (Carl Dept. of Biological Sciences, Simon Fraser Universi­ 1968). Food plants in Europe include Holclls ty, Burnaby, B.C. for access to her libran' and en­ Iwwllls L .. AgroPlj/'01l r epells (L.), BrachYJJor/illlll couragement during pre paration of t h is syl va liclIllI (Huds.), Dactljlis girJlllel'ala L .. AI'- manuscript.

LIST OF REFERENCES Adkin, R. 1924. Apple fruit attackcd b\· Tortrix ir r l'aralla. Entomologist .S7: 188-189. Adkin, R . 1D31 . Ba/ocies al/guislioral/a Haw. bred from stored apples. E ntomologist 64: 214. Alford, D . V . 1974. Lepidopterous caterpillars damaging strawbern' blossom. PI. Path 23: 48-49. Alford, D. V. 197,5. Specific feed ing preferences of to rtricid lan'ae on fl owcring st ra\\bern' plants. PI. Path . 24: ,54-.58. Alford_ D. V. 1976. Somc observations on thc cffeel of pests on Slnl\\'lw!'n' \'ields. Ann, appl. BioI. 84: 440-444.

Ali Naizee, M. T . 1977. Bionomics and life histon' of a filbert leaf roll er Arrhi"" rosa II us (Lepidoptera: Tor­ tricidae). Ann. entom ol. Soc. Am . 70: 391-401. Andison, H. 1937. The juniper webworm (Dichomeris lIlargillr l/a Fabr.). Proc. entomol. Soc. Brit. Col. 33: 3-5. Anon. 1900. Heport of the Ent. and Bot. Ann . Rept Expt'al Fms: Append. Rept \1 in. Agr . Expt'al Fms. Ses­ sional papers XXXV (6): 16, p 238. Anon. 1903. Ann. Rept Expt'al Fms. ; Append. I\ ept \1in. Agr. Expt'al Fms. Sess ional Papers XXVIII (6): 16 I' 16,5-167, 184-186. Anon. 1906. Report of the E nt. and Bot. Ann. Rept Expt'al Fms.: Append. Hept Min. Agr. Expt'al Fms. Sessional Papers XLI (6): 16, p 67. Anon. 19 12. Ent. Soc. Ont.. Ann. Rept 43: 1,5 . Anon. 191,5. B.C. D ept. Agr. 9th Ann . Rept, UJI4. PI' H6.5 , RH2-9.3. Anon. 1921. B.C. Dept. Agr" 1.5th Ann. Rept Hl20, pp Q3, Q20, Q27. Anon. 1927. CIPR. ,5: 39. Anon. 1936. The Diamond-back moth. \1in. Agr. Fish , Britain. Ach-ison Leaflet No. 19,5. Anon. 19,51-19,52 . Distribution maps of insect pests. Ser. A Nos 8 and 9 . Commonwealth lnst. of Entomol. 19,51-1 9,52. 46 J. E'TOMOL. Soc . BHI T. COL.U ~ lIl I A 79 (l982). DEc. 3 1, 1982

Anon . 1954 . B. C . Dept. Agr. 49th Ann. Rept. 1 3 ~ pp. Anon. 1968. Outbreaks and new records. PI. Prot. Bu ll. FAO 16 no ..5 pp 92-94: No . 6 pp 11 4- 11 5. Anon. 1979a. Informati on package on the !'(\'PS\' moth in the Kitsilano Area. Prepared jointh ' by B. C. Min. Agr. and Can Dept. Agr. 6 pp. An on, 1979b. Coop. PI. Pest Re pt. 1: 720. Anon. 1980. Coop. PI. Pest Rcpt. 5: 68D . Arestegui, P. A. 1977 [Pest of potato in Andahuadas- Apurimac]. He\'ista Pen'ana de Entomologia. 19: 97-98. Abstract in R.A .E. Ser. A 1979.67: 31. Arnott, D . A. and J. C . Arrand. 1960. C IPR . 38: 12. Arthur, A. P . 1966. T he present status of the introduced ski pper, Th yme/i clls li ll eo/a (Ochs.) (Lepidoptera: Hes peri idae) in North America and poss ibl e methods of control. C an. Ent. 98: 622-628. Bacon, O. G . 1960. Control of potato tuberworm in potatoes. J . eeo n. Ent. .5 3: 868-87 1. Baggiol ini , M. 1967. The apple moth. Il ecllrvaria /ell ('alel/a , a little kno\\n pest of fruit trces in Switzerl and. Agr. romande. 6: 111 - 114. Abstract in H.A.E. Se r. A 1970: .58: 211. Ba ker, YV. L. 1972 Eastern Forest . U.S. Dept of Agr. For. Se r. Pub #1175 .642 pp. Balaz, K. B. and J. Bodor. 1969. Larenrnorphologie an E rdbeere lebender Wicklerarten.lThe larval mor­ phology of Tortri cids living on stra\\·herries]. Acta. phytopath . Acad. Sci. Hung. 4( 1): 77-79. Abstract in R.A.E. Ser. A 1970: 345. Bauman, N. G. and B. A. Sugden . HJ 68. Juniper wehwon n in Brit ish Col umhia. Can. For. Ser .. For. Ins. D is. Su n 'e\', For. Pes t Leanet 8: 3 pp. Beirne. B. P. 1971. Pest insects of annual crop plants in Canada. I. Lepidopkra. Mem. Ent:. Soc. Can. #78: 124 pp. Belosel'ska\'a, Z. G . 1963. Argyreslhis ('ol/il/gel/a in the Lcningrad region. Zash. Hast. 8(2): .5 1. Abst ra ct in R.A.E. Ser. A lD64. 52 : 323. Belo\', U. K. , Yu. B. Shurovenkov and N. A. Tara n. 1978. [Care - the meadow llIoth!] Zash. Hast. 3: 11 -13. Abstract in R.A.E. Ser. A 197D. 67: 14. Benoit. P. (Comp.) 1975. Noms fr ancais d'insecls au Canada. Agriculture Quehec. Pub QA38-R4-30. Bergmann, A. 1951-19.55 . Die Grasssc hmetterl inge Mitteldeutscltlands I·V. Urania Verlag. Jena. in Can. E nt. 100: 800. Bethune. C. J. S. 1869 . Larvae infesting the parsni p. Can. Ent. 2: I -~. Bethune, C . J. S. 1886. Remedies fo r nox iolls insects. Ent. Soc. O nt .. Ann. Rept 17 : 55-64. Bibolini , C. 1970. Bioloi,'Y and damage in the Pisa d istrict of Periilrollla sall cia Il h (I.ep . ;\octuidac) wit h refe rence to its geographic distribution. world dietology and relati \'e control lII easures. Fru., tula E n­ tomologica 10: 98 pp. Abstract in R.A.E. Se r. A 1D75. 63: 13R. Blackmore, E. J-l . 1921. B.C. Provo Mus. i\at. IIist. Hept 1!)20: 22-32. Bl ackmore. E . H . 1923. Ent. Hept. Provo Mus. i\at. /-list. B.C. 1922: IR-35 . Rohm , H. 1962. Auffallende Schadcn an Apfcl baumen. [Noticeabl e damage to apple trees]. pnanzenarzt 15: 28-29 . Abstract in fLA.E. Se r. A 19(i2. 50: 302. Bo lte, K. B. and E . Munroe. 1979. J-/ clIlilhco oesli varia in British Columbia: C haracters of a palaearctic genus and species not previousI\' known from North Am erica (Lepidoptera: Geomelridae: Geo metrinae). Can. Ent. 11 1:11 21-1126. Boness, M. 1976. Trials to control the fru it tree leaf ro ll er AU'hips podano \\ith pheromones. Zeit. Ange. E nt. 82: 104 - 107. Bradley. J. D ., \\I. G. Treme\\'an and A. Sm ith. 1973. British Tnrtricoid '.Ioths. Coch,'lidae and Tor­ tricidae: Tnrtricinae. The Hav Societ,·. London. 251 pp. Breakey, E. P. 1960. The bl aekheaded fire\Vorm of cranbern'-a pest of the e\'ergreen hucklebern' in western Vlashington . J. econ. Ent ..53: IODI- 109D . Bri ggs, J. B. 1957. So me featllres of the iJiol og\' of the winter moth on top fruih. J . II ort. Sci. 32: 108- 12.5 BfA 33: 282. #3628 Brittain. W. H. 1912 Report from the Okanagan Dist rict: Insects pest of the \','ar in Okanagan. Proc . en­ tomol Soc. Brit. Col. 2: 13. Brittain. W. II. 1914 . Report from the Okanagan Dist ri ct. Insect pes ts of the \'Ca r in the Okanagan. proc. entollloi. Soc. Br it . Co\. 4: 1-1 - 19. Brittain. W . J-l . and C. B. Gooderham 1916. Popillar and Practical Entomologv: an insect cnem\' of the parsni p. Can. Ent. 48: 37-,11 . J. ENTml0L. Soc. BlllT COI.U~ "l I A 79 (1982), D EC. 31, 1982 47

Burgess, A. C. 1921. The satin moth, an introduced enem" of poplars and wil lows. U.S. Dept. Agr. Circ. 167. Burns, J. M. 1966. Expanding distribution and evolutionary potential of Thljlll eliclI .\·lill eola. an introduced skipper, with special reference to its appearance in British Columbia. Can. E nt. 98: 859-866. Busck. A. 1914. A destructi ve pin e-moth introduccd from Europe. E ll eiria hllo/i(lll(J Sch iff. J. econ. Ent. 7: 340-34 1. Caesar, L. 1929. A new orchard pest in Ontario. Ent. Soc. Ont. , Ann. Rept 60: 140-141. Caesar. L. 1937. Insects of the season in Hl36 in Ontario. C IPR . 15: 33. Cameron, E. 1935. A studs' of the natural control of ragwort (Se ll ecia iacobaea L. ). J. Ecol. 23: 26.5-322. Carl , K. P. 1968. Thymeli CII s lill eo la (Lepidoptera: Ilesperiidae) and its parasites in Europe. Can. Ent. 100: 785-801. Chapman. P. J. and S. E. Lienk. 1971. Tortricid fauna of apple in New York. Special Pub. N.Y. State Agr. Exp. Sta .. Geneva. Cornell Uni\·. 122 pp. Chittenden, H. F. 1905. The imported cabbage worm (Polliia rapae Linn. ). USDA Rur . Ent. Cir. No. 60. Clark, J. F. G. 1930. Entomology ill Rept. Provo Mus. Nat. Hist. B.C. 1929: 12-16. Cochran, S. G. 1974. The birch casebearer in Newfoundland. Masters thesis. McGill Uni\·ersit\. P.Q. , Canada. 126 pp. C it ed in Raske, A.G. 1978. Condrashoff, S. F. 1956 . Advance of the satin moth, Stilpnolia salicis (L. ) in to the interior of British Col­ umbia. J . entomol. Soc. Brit. Col. 53: 26-27. Cornstock, J. H . and M. V. Slingerland, 1980. A leafroll er on currant. Cacoecia rosana. Order Lepidoptera: fam il y Tortricidae. Cornell Univ. Agr. Exp. Sta. Bull. 23: 119- 121. Coshan, P. F . 1974 . The biolo!.,,,· of Coleophora serra leila (L.) (Lepidoptera: Coleophoridae). Trans. Roy. Ent. Soc. 126: 169- 188. C ram, W . T. 1973. Occurrence of the strawberry tortrix. Acleris c01llariana (Zeller). a new pest in British Columbia. J. entomol. Soc. Brit. Col. 70: 17. Cram , W. T. and C. L. Neilson . 1978. Major insect and mite pests of bern ' crops in B.C. B.C. Min . Agr. Pub 78-12: 24 pp. Cram, W. T. and N. V. Tonks. 1959. Note on the occurrence in British Columbia of the omnivo rous leaf tier, Cllephasia IOllgana (Haw. ) (Lepidoptera: Tortricidae), as a pest of strawberry. Can. Ent. 9 1: 155-156. Creelman, I. S. (cd.) 1966. CIPR. 44: 12. C reelman. l. S. (ed. ) 1973. CAIPR . 51: 32. Crys tal , R. N. 1937 . Insects of the British sodlands. Frederi c Warne & Co. Ltd .. London. 338 pp. Daviault. L. and R. Ducharme. 1966. Life histon' and hahits of the green spruce leaf min er, Epill olia 1l(JllQllQ (Treitsc hke) (Lepidoptera: Tortricidae). Can. Ent. 98: 693-699. Davidson, R. H. and L. M. Peairs. 1966. Insect Pests of farm. garden and orchard . John \Vile\' & Sons, Inc. , New York. 675 pp. Dawson, A. F. 1971a. Larch casebearer in British Columbia. For. Ins. Dis. Sur. Pest Leafl et. No. 34, 3 pp. Dawson, A. F . 197 1b. Satin moth in British Columbia. Dept. Environ. For. Ins. Dis. Sur. Pest Leanet. No. 38, 3 pp. DeMeirleire, H. 1966. Un ennemi des peupli ers: Ie bomb\'s du saule. Ph\·toma 18: 48-49. Doganlar, M. and B. P. Beirne. 1978a . Croesia holllliana (Lepidoptera: Tortricidae), a fruit tree leaf roller new to No rth America established in British Columbia, Canada. Can. Ent. 11 0: 639-640 . Doganlar, M. and B. P. Beirne. 1978b. Fruit tree leaf-roll ers (Lepidoptera) and parasites (Hnllenoptera) introduced in the Vancoll\'er district, British Columbia, Canada. J. entomol. Soc. Brit. Col. 'i5: 23-24. Dognalar, M. and B. P. Beirne. 1979a. Pal'{/ia cljllosiJalel/a (Lepidoptera: Tortricidae). an European leafroller new to North America. Can. Ent. 111: 869. Doganlar, M. and B. P. Beirne. 1979b. Panrielllis heparana (Lepidoptera: Tortricidae) a Tortricid nel\' to North Am erica established in British Columbia, Canada. Can. Ent. 111: 970. Doganlar, M. and B. P. Beirne. 1979c. lIemithea aesli varia. a geometrid new to North Am erica, establ ish­ ed in British Columbia (Lepidoptera: Ceometridae: Gcometrinac). Can. Ent. III: 1121. Dow nes, W. 1928. Notes on economic insects on Vancouver Island in 1928. Proc. entomol. Soc. Brit. Col. 25: 16-18. Dustan, A. G. 1932. Vegetable insects and their co ntrol. Dom . of Can. Dept. of Agr. BIlII. 1\0. 161: pg 51. 48 j . E1\ Tm tOL. S(Jc. BIUT . C() t .l"~ t B L~ 79 (1982) , D EC . 31, 19S2

Dustan, C. C. and T . R. Davidson 1973. Discases, insects and mites of stone fruits. C.D.A. Pub .. 3rd rc\·ision. Edland, T. 19().5. [Apple fruit moth (1\. (,o l/ il/gel/a) . Bionomics and control - a prdi tni na r\' report]. C art­ ncr\Tket 55 pt. 16-17 pp 430--136. Abstract in B.A. E. SC I' . A lflfi7 . 5.5: 1-1 6. Edwards. vV . D. and D. C. 'vl otc. 1!J36 . Omni\'()ro lls lea f tic I' , CI/cphosio /ol/gol/o Ila\\·. A rel ati\'eh' ne\\' pes t of strawberries, iris and other crops in Ore!!:on. j . ccon. [nt. 29: 111 8- 11 2:3. Ehnst rom, B. , B. Bejer-Petersen , K. LO\'ll\'ni emi a nd S. TII ('rIll \T. 1!J7-1. Insect pests in forest of the :\ordie cou ntries HJ67 to 1971. Ann. [nt. Fcnn. -10: 37--17. Abstract in I\ .A. I:: . Ser . A Im.s. 63: 1-140 . E kholm. S. HJ58. A I/lhaph i/a poria,/{J as a pest in Finla nd. :\otul. e nt. 38: 87-!J0 . E kholm, S. HJ6 1. The pea moth in Finland and its control. II lataloli s jakot'luilll 15: 228-23-1. Abstract in I\ .A.E . Ser. A W()3. 51: 6-1 . Embree, D . G . 1967. Effects of the \\'inter IIl oth on ((ro\\'th a nd nlO rtalih' of red oak in :\()\'a Scotia. For. Sci. 13: 2!).S-29!). . . Evans. D . 1966. First North American record of POI/r/emis ri/)('o lla (Illlbncr) at Vi ctoria. B. C. Hi-ll1on. Pro!!:. Hept Can. Dept. For. 22: 3.

E \' ans. D . 1970. L ife histo J'\' and imill atul'! ' .s tages of /'ol/dl'mi., ( ' (' /'(1 .1'01/0 (Lepidoptera : T ortricidae). Can. Ent . ItJ2: 15!l7- 1603. Evenhu is. \·on. II . H .. D . \\I. deJong and II. j . Vilig. 19.3 . Oil' I'lippell dcr in den f\:iederlanden am Apfel \'orko m menden B1 att\\'ickl er (Lepidoptera: Tori rieidae). Z ange\\'. Ent. 73: ,'351-365. Felt , E . D. 19 10 . j un iper W e bworm . f\: .Y. Sta te ~1 u s . Bu ll. 1-1 7: 35-36. Fel t. E. P. 19 17. Th" apple and thorn skell'lonizer. Ent. Soc. Ont.. Ann. Hept -1 8: -1-1--17. Felt, E. P. 1922 . OCllerosl()lIla pilliariella Zl'i l. . another introdllced insect. J . ('con. Ent. 1.5 - -1 32--133. Ferguso n. D. C. 1975. Pests not kno\\'1l to ocelli' in the Uni ted States or of lim ited di.st ri lliitioll. \\'inter moth (Opera/lillera /}/'/I/}/fIla (L. )) . Coop. c('o n. Ill S. Hept .'3 (-1 8- .,) 2): CiH7-69·1. Fletcher, J. ISB-1 . Heport or the Ent. and not. - frllil s. Ann. Bept on the ExpLd Fnls .. Append 10 the Hepl or the Min. or Agr. on E xpt'al FillS . IDS - 19(i. Fletcher, j . IS97. Heport of the Ent. and Bot. Ann. Hepl Expt'al F illS .: Append. i1 ept Min. A(( r . Expt'al F ms. Sessional papers XX XII (fi ): 8a p 201 -202. Fletcher, j . IS!)!J. Inj urio lls insects in 1899. 29th Ann. i1 cpt l' nt omo!' Soc. Onto pp 7.'i-H7. Foot. II I. A. 1979. l3ionomies of the potato tuber moth . l'ilihori lllaea "!i('rcli/ella (Lepidoptera: Gelechiidac) at Pllkckohc. N.Z. j . Zoo!. R: 023-630. Abstract in B.A.F:. Ser. A 19S0. 6R: 507. Forhes. n. A. ('I at. 196-1 . Ma ri times Pro\, in c( 's - forest insect conditions. Pages 21-32 ill Ann. Rept For. Ins. a nd Dis. Sur\, . 1963. Call. Dept. For .. Ottawa. Forbe." \ V. T . 1\1 . 1923. The Lcpidopll'ra of Nc\\' York ancl neighhouring states . prilTlili\'e fo rms. Mi crolcpidoptera. P\Taloids, nOlllh\·ces. Cornell Uni\·. Age. Exp. Sta. \km . 6H: 729 pp.

Forbes. W . T. M . 192-1. '1'\\'0 minor pests from ElII'ope (Lepidoptera) (Plutclli dat'). ] . N .Y. cnto rn ol. Soc. 32: 173. Forbes. \\' . T. \ 1. 19.'5-1. Lepidopll'ra of Ne\\' Ynrk and nl'ighhoring slates. Noctllidae. Part III . Cornell Univ. Agr. Exp. Sta. M em . 329: -1 33 pp. Frankli n. II. j . 1950. Cra nberr\' illsects in \1assachusdts. 1'-. lass. A!!: J'. E xpt Sta. nlill . -1-1.') , parts 2-7 : 8R PI'. F reem an. T. N. 19SH . Thc Archipinae or North America (Lepidoptera: Torlrieidac). Can . En!. Suppl. I. 89 pp. Freeman, T. 1\. 19R7 . Annota ted ke\'s to some 0/careti c leaf-mining Lepidoptera on conifers. Can. Ellt. 99: 419-435. Frilli , F . 1!J77. Guicled control aga insl the pests of peach . Informatore Fitopatolo!!:ico 27 : 21-22. Fulton. II. G . 1947. Natllral control or the Europea n pca Ill oth . LaSpClj!'l'sio IIigrimlla on SUlllas Prairie, B.C. (Lepidoptera : OI" lIthr(,ll licla(') . Pmc. enlomol. Soc. Brit. Col. -1 .3 : 25:27. Furniss, H. I. and V. M . C aroli n. ID7 7. W estern Forest Insec t.s. U.S . Dept. Agr . For. Ser\, . I-lisc. Pub. 1,3.39: 6.5-1 PI'.

Gerardi, M . I I. a nd j. K. Grimlll . 1970. The histoJ'\·. biolog\·. damage and control of the g\'pS\' moth P OI'­ Ihe/ria dispol' (L. ). Associated Un i\'prsity PI'!'sses. Inc .. Cranbur\'. Ne\\' j ersc\' and London. E ngland. 233 Pl'. Gerber, H. S. c/ a/. 1980. Reco!!:n ition and life histol'\' or the major insect a nd mite pests of ornamental shrubs and shade trees in British Coilimbia. Brit. Col. Dept. Agr. Pllh. 80-,5. Gibson, A. 19 12. Cutworllls and armyworms. Can. Dept. Agr .. Ent. D i\·. Bu ll . 3 . J. EI\TO.\IOL. Soc. BRIT . CO J. U ~IIlIA 79 (1982), DE c. 3 1, 1982 -19

Gibson, A. 192 1. Common garden insects and their control. Can. Dept Agr.. F.nt. Di\·. Cir :\0. 9. G ibson. A. 1923. The specifi c name of the green buch·orm. Can. Ent. 55: 2-1 3. Gibson, A. 1924. The occurence of the tortricid Cacoecia rosalia L. in Canada. J. econ . Ent. 17 : 51·54. Gillespie. D . R. 1981. Introduced and native leafrollers (Lepidoptera: Tortricidae) on berry crops in the Lower Fraser Valley, B.C. Doctoral Thes is. Simon Fraser Uni\-eTsit\" . Burnabv, B.C. 179 pp. Gillespie. D . R. and T. Finlavson. 198 1. Final-instar larvae of nati\'e Hmlcnopterous and D ipterous paraistes of Operophlera spp. (Lepidoptera: G eometridae) in British Columbia . Can. Enl. 113- -15·55. Gillespie. D. R. , T . Finlayson. :\. V. To nks and D . A. Hoss. 1978. Occure nce of the \\'inter moth. Operophlera brlllnaia (Lepidoptera; Geometridae) on Southern VanCOU\'cr Island. British ColulIl' bia. Can. Ent. 110 : 223·224. Gilliatt, F. C. 193-1. Notes on the lesser budmoth, Recll rvaria 'wileI/o Hbn . Sci Agr. J./ : 466·-176. G lendenning, H. 1924 . The satin moth in British Columbia. Dept. Agr. Pamphlet no .5 0. Glendenning, R. 1929. Host adaptation in the European satin moth. Proe. entomol. Soc. Brit. Col. 26: 34·38. Glendenning, H. 1944. the parsnip \\"eb\\'orm (Depressaria " ewe/ialla) and its control in British Columbia (Lepidoptera: Oecophoridae). Proc. cntomol. Soc. Brit. Col. 41: 26·2R. Glover, T. I R74. Report of the Entomologist . Ann. Rept U.S. Dept Agr. for I S72. p 11 2· 11 3. ill Jones. S. I. 1935 . Gould. H . J ., T. J . Lego\\'ski and E. C. Atkins. 1962. Sun'e\'s of pea moth damage on dn' har\'esting peas in East An gli a. 1957· 1959. PI. Path. ll : 1·6. Grar. J. E. 1917. The potato tuber moth. U.S. Dept. Agr. Bull. -127: 56 pp. ill Bacon. O.G . 1960 . Handford , R. H. 19055. Insects of the season in British Columbia 1955. p 17. Compiled h\· C. L. Neil son, Provincial Entomologist. Vernon. B.C. Harcourt. D. C. 1962. Design of a sampling plan for studies on thc population dmamics of the imported cabbage \\·orm. Pieri.'; rapae (L.) (Lcpidoptera: Pi cridae). Can. Ent. 9-1: 1l-l9-8.59 . Ha rris. J . W . E . and D . A. Ross . 197.5. European pine shoot moth. For. Pest Leanet :\0. 18: 3 pp. Harris. P. 1958. Life- histon ' and natural control in British Columbia of Ocn""0810,,,a piniariel/a Zell. (Lepidoptera: Yponomeqtidae) a needle m incr on white pine. Can. Ent. 90: 627·63 1. Harris. P. , A. T . S. W iki nso n. M. E . :\ean', L. S. Thompson and D. Finnamore. 197.5. Establishment in Canada of the cinnabar moth. Tyria ja robaeae (Lepidoptera: Aretiidae) for controlling the weed Senecio ;acobaea. Can. Ent. 107: 9 13-9 17. . Ha rris, P., D . Peschken and J . 1--lilro\·. 1969. The status of biological control of the \\·eed. lIYJJeri('",n per· jonnailim in British Columbia. Can. Enl. lOl: I· IS. Harris. P. and D. P. Peschk en. 197 1. lIyperirllm peljoralll'" L .. St. John's W ort (H\·peri caceae). in Biological control programmes against in sects and weecls in Canada 1U.59- 1968. Technical com· muni cation No. -I . C. I.B.C., Trinidad. pp 89·9-1 . Harris, T . W. 18-11. A report on the insects of \1assehusetts injurioLls to \·egetation. -1059 pp. C it cd from C hapman. P. J. a nd S. E. Lienk 197 1. Hawkes. R. B. 1968. The cinnabar moth. T'fT ia ;acobaeae. for the control of the tans\' ragwort. J. econ. Ent. 61: 499-50 I. Heinrich, C . 1923. Revision of the Korth Ameri ca moths of th e subfamih' Eueosminae of the fam ih ' Olethrcutidae. U.S. I\:at. ~vlus. Bull. 123. . . Heinrich. C. 1956. American moths of the subfamil\' Ph\'eitine. U.S. Nat. \lus. Bull. 207: 69. 90. 209. Heppner, J . B. 197R. Ellimaw/a pariana (Clerck) (Lepidoptcra: Choreutidae). the <:oITeet nallle of the apple-and-thorn skeletonizer. J . entomol. SoC'. Brit. Col. 75: -10·-11. Hodges, R. W ., in Dominick. R. B. , el a/ .. 197-1. The "Ioths of Am eri ca North of I\ lexico, Fase. 6. 2, Celechioidae; O ecophoridac (in Part). E. \V. Classey Ltd. and H.B.D. Publications Inc. , London. Holland, \V. J. 1968. The moth book. a popula r guide to a knowledge of the moths of North Am erica. Dover Publication .. Inc .. Nc\\' York. 479 pp. I nllllS , A. D. 1964. A general textbook of entomol o).,1Y . :,\ inth edition rC\'ised b\· O . \\'. Richards and R. D . Davis. Methuen and Co. Ltd .. London. SIl6 pp. J::ekson. 13. S. 1978. Records of the European skipper in Newfoundland. Can. Fld. :\at. 92; 200. Johansson, S. 1962. Insects associated w ith H ypericllm I. 2. Lepidoptera. Diptera. Ih'mcnoptera, Homoptera and general remarks. Opusc. ent. 27: 175- 192 . 50 J . E:-.JTO .\lOI.. Soc. RHI T C() I.l ~ I B I II 70 ( lHR2), D E( . 31. 1082

Jones, F . C. W . and 1\ 1. Jo nes. 196-1 . Pests of fi eld c rops. Ed\\'ard Arnold (Publishers) Ud .. London . -106 pp. Jones. S. I. 1935. Obsen 'ation o f the ha bits and seasonal life histo n ' of Allorsia /il/ca /e//a in C alifo rnia. J. econ. Ent. 28: 1002-10 11. Ka ilidis. D . S. 196-1. An epidemic or S/i/pllo /io so licis L . (Le pidoptera: LI'mantriidac) during 106-1 on poplars in \1accd onia. [in Crcek]. Dasik a Ilro n . no 7.'3 -7-11 2 pp. Abstract in H. A.E. SCI'. A !H67 . .')5: 158. Kearfnot, W . D . 1907. Ne\\' \Iicro l.e pidoptera. C an. Ent. :JD: 79. Keifer, H . 11. 19:33 . California \Iierolcpidopll'ra . V!. Calif. Dept Ag r. \1 0. Bllil . 22: :3.5 1-.'3 £i.'5. Kiziroglu, I. ID76 . On the kno\\'ledge o f the lil ac leaf mine r ll1oth. Craci/aria .I·I/rillgc//a F. (Lepid .. C racil ariidae). I. Dist ribution and Jllorpholo,,'\,. Zeit. Fur. Angc. Ellt. RO: -107--1 26. Ahst ract in I1 .A.E . St'!'. A 1977. 6.'5: 57R . Kolel·. K. and A. 13al c\'ski . 1978. In<:reased infestation IJI' leafroll c rs 011 fr ui t tn·t·s. Ha st. Zash . 26: 27-:32. Kra ko\\'la k. A. 197-1 . [O ccurence of Tortricidae (Lep.) in orchards o f the Pozna lJ region (Poland)]. Pol. Pismo cntol1lol. -1-1 : 8:35 -8:38. KrueL W . 1866. SpontaneollS llluSS occurence of the ro\\'an mot h (Argl/u's /" ia l'O lljllge//a) in apples (in C er­ m am' in 196.5). Nac hrfll. D t. PnSciultzdie nst Berl. 20 (1\.S. ) pt -1 11:3- 11 -1. Lazorko. VV . 1977. Ellrr/Il/l)a/'(/ " "r/II/a/a l ,. (urticata L.) on the Pac ific coast. (Lc p : !'::ralidae). J . entol1lol. Soc. Brit. Col. 7-1 : 3 1. Lejeu ne . 11. R. and C . T . Sil l't,r. 196 1. Parasitcs a ndlll'j)crparasites of the sa ti n lIloth. S/i/pllo /ill solicis l.i n­ naells (Ll'fnantriidae) in British Columbia. Can . Ent. 9.'3: -156--107 . Leondard. 1\1. D. 1923. The present sta t llS and distri b u tion lI f the apple and thorn skele tonizer . E nt. Soc. Ont. An n. l1 ept .'1-1: 18-20 . Le l1 oux, E. J. and C. Heimer . 19.59. V ariat ion iJ el\I'l'en safll ples o f imIwltli re stage's. and of mortalit i( 's from SO lli e fac tors. of the el'e-spotted b udll1 oth. Spi/VIl() /Oocd/lll1a (D. & S. ) (L('p .: Olethreutidae). and the pistol casebeare r, Co/cophol'(l se rra /el/a (L.) (Lep.: Coieo phorirhl!'). on apple in Quebec. Can. Ent. !J I: ~ 28--1-1 9 . Lindquist , O. H . a nd L. S. MacLeod . 1967. A biological stllch' of Epillo/ia sO /(/Ildrialla (Lepidoptera: Olcthreutidae). a leaf roller on birc h in O nta rio. Can. Ent. 99: 1110- 111 -1. Lintne r, J. A. 1882. Entoll1ologist o f Ne\\' Yo rk . F irst An n ual Re port of I nj lIrio lls a nd Oth('r I nseets of:\ . Y. P 1.52- 1.55 . ill Jones. S. L. 193.5 . Ll e \\' e ll ~ ·n -Jones. J. R. S. 1!J3R. Some fo od planh of Lel'i

""auhe\'. \ V. 1067 . The natmal histor\' of three oak feeuing To rtric:i cb: PI 1/e/IO /11 III a /I'chca ll a (L.), Palldelllis ('crasana (H b. ) and Balor/cs allgllsliuralla (I-Ia\\·.). Entomologist 100: 11 .5- 120. Maw, ~v1. C. 1976. An annotated list of insects associated with Canada Thist le C i rs ill lll {1fTI'll.\'{' in Canada. Can. Ent. 108: 2.35-2 ~~ . Ma xwell , C. v\' . and A. D. Pickett. 1957. Insects affecting the ('ranbern' Can. Dept. Agr. Pub. 8 10, the e ranb e rr ~ ' P 2.5-28. McDonald. S. 1963. C hemical cont rol of the beet webworm on sugar beets in southern Albcrta. J. ccon . Ent . .56: 2~8-25 1. McDllnnough, J . H. 19.57. S\'Iloll\'m ic and bi ological notes on Coleophoridae (Lepidopte ra). Amer. \1us. Novit. No 1827: pp 1 -~ . Abstract in R.A.E. Ser. A 19.59.47: 11 7. McLeod, J. II . 195 1. Biological control investi gati ons in British Columbia. Proe. entomol. Soc. Brit. Col. 47: 27-36. McNeil. J. N. 1978. The num ber of larval stages of Tilljlll e/i('lIS /ill('u/a (Lcpidoptera: I! esperiid ae) in eastern Canada. Can. Ent. 110: 1203- 129.5. Metcalf, C. I. and W. P. Flint 1062. Dcstructive a nd useful insects, their habits and control. \1cCraw- Hill Book Co. , Ne\\' York . 1087 pp. Me\Tick, E . 1927 . A rcvised ha ndbook of British Lepidoptera. W atkins Doncaster, 36 Strand , London, W.C. 672 pp. Miczulski, B. and M. Koslinska. 1976. [Parasites of leafroll crs (Tortricidae) occurring in apple orcha rds in so me regions of Poland]. Pol. Pismo Ent. 46: 165- 178. \1iller. C. E. 1076. Native parasites of Coleophora larieel/a (Lepidoptera: Coleophoridae) in British Col­ umbia. Masters Thes is. Simon Fraser University, Burnab\', B.C .. Canada. 75 pp. Minder. I. F. 19.59 . Leaf-roll ers injurious to fruit crops in the fl ood-plain of thc Oka ri\'Cr. He\·. Ent. UHSS. 38: 98- 11 0. Abstract in R.A.E. Ser. A 106 1. 40: 341. Mitchell, B. I. 1978. The biological control of potato tuber moth, Phlhorima('a operclliel/a (Zell er) in Rhodesia . Rhodesia Agr. J. 7.5 : 5.5-.58. Abstract in R.A.E . Scr. A 1978.67: 8.5. Molis, S. 1970. A ma.'s outbrea k of the satin moth (S lilpllolia sa lids L.) on poplar in the south of the Lithua­ nian Hepublic (Soviet Union) in 1967. Acta Entomolngica Lituanica I: 1 8.3 - 1 8~ . Abstract in R.A.E. Ser. A 1972. 60: 968. Molnar, A. C. , J. W. E. Ha rris a nd D . A. Hoss. 1966. Can. For. Scr. Dept of the Em·i r. Annual Hept For. Ins. Dis Sur. , 1966, Otta\\'a, Information Canada 1967. 1~ 2 pp. Morris, H. F. 1980. Butterflics and moths of Ne\\'foundl a nd and Labrador. the macrolepidoptera. Res. Branch. Agr. Can. Publ. 169. Munroc, E. in Dominick, H. B., cl 01 .. 1076, The moths of Amcrica north of Mex ico , Fac. 13.2A. Pyraloidea (in part). E . W. Classey and R.D .D. Publi cati ons Inc., London. Murdoch. R. F. ivl. 1967. The biology of the lilac Ieafminer, Craci/laria sl/ril/f!,('/Ia Fabr. (Lcp .: C racillariidae). Quaesl. enl. 3; 24 7-281. \ 'l usich, E. N . 1976. [Cutworms as pests of bern ' crops]. Zash. Rast . 4: ~7 . Abstract in n .A .E. SC I' . A W77. 6.5: 722. l\1\'crs, J . H . and B. J . Campbell. 1976. Predation b\' carpc nter ants: a deterrent to the sprcad of cinnabar moth . Proe. entomol Soc. Brit. Col. 7.3: 7-n. Naito, A. 1961. Studies on the distributi on and abundance of the lima bean pod borer, fliel/a ::. illckel/ cl/a Treitschke, and the sm'bean pod borer Crapho/il/w [ L cg lllllilliwraj glljci,livorel/a ivl atsulllllra IV . On the distribution of the two specics in thc world and their host plants. Kont\'ll 29: 39-.55. 1\akamllra, M. 1974. A ecological st udy of fpil/ olia su lalldrial/ a (Li nnaeus) (Lepidoptera: O lethreutidae) a leaf roller on Belllia enl/al/i Cham. (Betul acac) in the Shi ga\'ama IPB area (Japan). Jap. J. Ecol. 24: 186- 193. Abstract in Ent. Abstr. 7: 76. Neacu, P. a nd I. Dragia . 1973. Donnes ecologiq ucs sur certains Lepidopte ra et Diptera mineurs dans la foret de Petresti , district Vrancae Tra\'call x du Muscl lm d' lIisto ire Nat urell e. "Crigo re Antipa" 13. 195-202. Ne ilson, C. L. 19.57. Tree fruit in sects: Ha ndbook of the main economic in sects of B.C. Part ~ ..52 PI'. Neilso n, C. L. 1969 . Berry and grape insects. Handbook of the main economic insects of B.C. Part 3. B.C. Dept. of Agr. PI' .51-.52. Nikolova. V. 1962. Injurious leafroll ers (Lep. Tortricidac) a nd Gelcchiids (Lep. C elech iidac) on fru it trees and their parasites. Izv. nallch. Inst. Zash. Rast. 3: 0 1-1 Ill. Abstract in H.A.E. Ser. A WG3. 5 1: 554. 1\ordin. C. L . and J. F:. Appleb\·. 19G9. Bionomics of the juniper \\·cb\\·orm. Ann. cntomol. Soc. Am. 62: 287-202. 52 j. EKTml OL Soc. BIHT. CO I. U~ I HIA 79 (1982), D EC. 31, 1982

Nordlander, C. 1977. Observations on the insects fauna in apple trees in connection wi th tests on insec­ ticides for integrated control. Vaxtskyddsnotiser 4 1: 39-48. Abstract in R. A. E. Ser. A 1978. 66: 20.54. Pengell y, D. I-J. 196 1. ThlJl1l eliCIIs lill eola (Ochs) (Lepidoptc ra: IIesperi idac) a pest of ha!' and pasture grasses in southern Ontario. Proc. entomol. Soc. Ont. 91: 180- 197. Peterso n, A. a nd C. j . Haeussler. 1926. The oriental fruit moth. U.S. Dept. Agr. Dept. C ir. 395. 27 pp. Petherbridge, F . R. 1920. The life histon' of the strawberr)' tortrix Oxygrapha f"O l1lariOli a Zell er. Ann. appl. BioI. 7: 6- 10. pfadt, R. E. 1971. Fundamentals of appli ed entomolog\·. Macmillan Pub. Co. , Inc. , New York and Toron­ to. 693 pp. Pierce. W. D . 1917. A manual of dange rous insects likel\' to be introduced in thc United States through im­ portations. U.S. Dept of Agr. 256 pp. Plank, H . F. 1922. The hl ackh eaded fireworm of c ranbern' on the Pacific coast. U.S. Dept. Agr. Bull. No . 1032: 1-4l. Popov, P. 1978. Pests of a rape varict)· and their control. Rast. Zash . 26: 2~ - 26. Abstract in B.A.£, Ser. A 1979. 67: 402. Porter. B. A. 1924. The bud moth . U. S. Dept. Agr. Bull. No. 1273. Cited in LeHoux. E. J. and C. Reim er. 1959. Powell , j . A. 1964. Biological a nd taxonomic stud ics on Tortricine 1ll oths, with reference to the species in California. Un iv. Calif. Press. , Berkle,' and Los Angeles. 3 17 pp. Prentice, R. vI. e/ al. 1062. Forest Lepidoptera of Canada recorded h,' the forcst insect survey. Vol. 2. N\'cteolidae, ;'\otodontidae, Noctui dae, Liparidae. For. Ent. and Path. Br. Can. Dept. For. Bull. 128 . pp. Prentice, R. M. e / al. 1963. Forest Lepidoptera of Canada recordcd 1)\ the forest insect survey. Vol. 3. Las icampidae, Th"atiridae, Drepanidac, Ceometridac. For. Ent. and Path. Br. Can. Dept. For. Pub. 10 13. 543 pp. Prentice, R. M. e/ al. 1965. Forest Lepidoptera of Canada recorded h)' the fo rest insect sun·ey. Vol. ~. \1 icrolepidoptera. Dept. For., Ottawa, Can. , Puh. 1142. 84 0 pp. Pres ton, W. B. a nd A. H. Westwood. 198 1. The European ski pper, ThYl1l elirll s lill('ola (Lepidoptera: Hesperiidae), in Manitoba and northwes tern Ontario. Can. Ent. 11 3: 11 23 - 1I 24. Putnam , W. L. 1963. The codling moth, Corporapsa Jlol1loll ella (L.) (Lepidoptera: Tortricidac): a re\'iew with special reference to Ontario. Proc. entomol. Soc. Ont. (1962) 22-60. Haine, J. 1966. Life history of Dasys/oma salicelhull (Ilbn. ) (Lepidoptera: Oecophoridae), a ness pest of blueberries in British Columbia. Can. Ent. 98: 33 1-334. Rasle, A. C. 1978 . Parasites of birch cascbearer larvae in Newfoundl a nd (Lep.: Colcophoridae), En­ tomophaga 23: 103-108. Razowski , J. 1966. World fa una of the Tortricini (Lcp .. Tortricidae). Krako\\', Pola nd. Reeks, W . A. and C. C. Sm ith. 1956. The satin moth, Slilp"olia salicis (L.) in the tvla ri time provinces and observati ons on its control by parasites and spraying. Can. Ent. 88: 565-579 . Reichert, A. 1932. Hosensc hadlinge (Hose pests) d ie Kranke Pflanze, IX , nos. 5-4. 75-78. Dresden. Rose, A. H. and O . H. Lindquist. 1977. Insects of eastern spruces, fir and hemlock. Dept Environ . Can. For. Ser.. For. T ech . Rept 23. Rose, A. H . and O . H . Lindquist. 1980. Insects of eastern larch, cedar and juniper. Dept. Environ. , Can. For. Ser., For. Tech . Rept 28. Rosenstiel , R. C., C. H. Ferguson a nd D. C. Mote. 1944. Some ecological relationshi ps of C Il ephasio IOl/gana . J. econ. Ent. 37: 814-817. Ross, W. A. and W. Putman. 1933. The economic insect fa una of Niagara peach orchards. Ann. Rept en- tomo\. Soc. Ont. 64: 36-41. Ruhman, tvl. 1915. Insect notes from the Okanagan. Proc. entomo\. Soc. Brit. Co\. 7: 7- 11. Ruhma n, M. 1928. The peach twig borer. B.C. Dept. Agr.. Hort. Branch, C irc 31 (revised ed .). Ryan, R. B. and H. D. Medley, 1970. Test release of Itoplectis quadricillgl,latus against European pine shoot moth in an isolated infestation. J. econ. E nt. 63: 1390- 1392. Salman, K. A. 1929. Notes on the immature stages a nd biology of a birch casehearer. Ann. entomol. Soc. Am. 22: 480-488. Sanders, C. E. 191 5. Budmoth in Nova Scotia. Proc. entomo\. Soc. N.S . 1: 8 ~-87. Sanders, C. E. and A. C. Dustan. 1919. The apple bud-moths and their control in Nova Scot ia. Dom. of Can. , Dept of Agr. , Ent Branch Bull. No. 16. J. ENTmlOL.. Soc, BHIT COLnllllA 79 ( 1fl82) . 01·:C. 3 1. HJ82 53

Saunders. W. E. 1916. European butterfh found at London . Ont. Otta\la NaL 30: 116. Cited in Burns. J. ~1. HlOG . Schmid!. L. 1972. Studies on the control of ra ~\\'o rt. Sel/ecio ;acoi)(lca L.. \I'ith thc cinna bar moth. Ca/limurpha ;arohaeae L. (Arctiidae: Lep idoptera). in Victori a. W eed Res. 12: .ffi-·17 . Sechser. \'on B. 1970. der parasit enkomplex des kl eint' n frostspanners (O /lcrIJ/litlef'{J bnllllota L .) (Lep .. Ceometridae) unter besonderer B e ru ck s i e htin ~u n ~ de l' Kokonparasit en. I. Z. ;,tnge\\,. Ent. 66: 1-35. Shaheen. A. II. 1977. Sun'e\' of pests attackin g tomato in Egypt \\'ith some ecological notes. Agr. Hes. Re\·. 55: .f 9-.57. Abstract in R.A .E. Scr. A 19RO. 68: .3 1.f. ' Sheppard. A. C. 1959. Caf'{Jdril/a 1lI 0rphclIs. a ne\\' record for North Alli eri ca of a E uropea n moth (Noe· tuidae) . J. Lepidop. Soc. 13: 77. Smith. C, C. 1950 . Notes on the European winte r moth (O /lcro/lh lcro bl'llmal a) in 1\o\'a Scotia. Can. Sci. Sen '. Oi\,. Ent. Fores t Insect Im ·est. Bi -mon. Prog. Hept 6: 1. Smith. K. M. 193 1. A tex tbook of agricultural cntomology . Cambridge Uni\'ersitl' Press. London. 28.5 pp. Specht, I-I. B. 1972 . C utworms of to bacco in Nm'a Scotia. l. Species co mplex and infestation. Can. F.nt. 10.f: 185.5- 186.f . Spencer, C. J. 192D . Rept cntolllo!. SOC' . OnL GO: 82-8.f. Spencer. II . and W. O. Strong. 1925. T he pota to tuben\'Orrn. Virginia Truck Expt Sta. BIIII .5.'3. C ited in Hofmaster. II . i\. and L. O . Anderson. 19.5.5. Potato tuber\\'orm control in Virginia. J. ('con. Ent . .f R: 198-201. Staudinger. P. O. a nd P. 1-1 . IIebel 190 1. Y/lSIJ /o/lh/l.\· IIwrgillclhls. p 15D in P. O . Staudinger and P. II . Rebel. C at. Lepidoptcra Palea rctic fallna. II. Friedlander and John. Berlin . Sutherland. D. \ V. S. (Chairman). 1978. C om mon names of insects a nd related organi,"}s (1978 Hc\·ision). Entomol. Soc. Am er. Spec, Publ. No. 78- 1. Swenson. K. C. 1958 . 1I 0ll y bud moth control. J. econ. Ent. .5 1: 87-90. Talhouk . A. S. 1977. Contributions to the knowledge of a lmond pest in east \ 'kditerranean countries . V . The fru it-feeding insects E llrylollla allli/gdali End ., a nd Allarsi(l /ill ('al(' /I(I Z. Zcit. ange. EnL 83: I.f S- IS4. Thiem , H . 19-13. Sac:ktragermotten un ih re Bekalllpfung illl Ha hmen der \\ 'inter- behandlung der Obstba ume. [Case-bearer ·moths a nd their eontrol in the course of \\'inter treatment of fruit trees.]. Kranke Pflanze 20 : 81-88. Abstract in B.A.E. Ser. A 194-1.32: 1-13 . Torgersen , T. H. and 1-1 . C. Coppel. HJ65. T he Eu ropean pi ne shoot Ill oth in Wisconsin . Wis. Consen 'a tion DepL For. Pest Leaflet i\o. 7 . 3 pp. Touzeau. W. and C. L. Neilson. 19.57. Plans to eradica te ori ental fruit Illoth in t h" Okanagan Valle\·. B. C. ProC. entomol. Soc, Brit. Col. ,,) 4: 23-2.f. Touzeau. W. O. and C. L. Neilson . l D58. Eradication procedures for ori cnta l frui t moth in the Okanagan V all ey of British Columbi a. Proc. entomol. Soc. Bri t. Co!. 5.5: 2.'3. Treherne, R. C. 19 14 . A review of appli ed entoillology in B.C. Proe, entomol. Soc. Brit. Co!. -I : 7-17. Treherne, R. C, 19 15. Popular and economic entomology. Notes of economic in terest frorn British Colulll­ bi a. Can. Ent. -1 7: 101 -1O.f . Treherne, H. C. 1921. A further re\'ic\\' of applied entomology in B.C. Proc. entoillol Soc. Brit. Col. 13 & 15 (Econ. series) : 135: 1-16 . Treherne. R. C. 1923, C IPl{. I: 1-7fi. l \\'inn. C. H. (ed. ) 1938. C IPH . 16: 2G7 . T\\'inn. C. H. (cd, ) 1939. C IPH . 17 : G. T winn, C . R, (cd. ) 194 1a. C IPR. IG: 1.f.5. Twinn. C. H. (ed. ) 19.f l b. C [PR . 19: 19,) . T\I'in n. C. H. (ed .) 194,5. C [PR . 23: 11 3 . Vernon. J, D . II . 1971. Obsen 'ations of the biology and control of tortricid larnle on stra\\'berrics. PI. Path . 20: 73-80. Vorzheva. L. V. 1958. On the bionoillics of some little-known .~"ilT olcpi dopt e r a and their injuriousness to fruit trees in C isba ik al ia . II c\·. Ent. URSS . 37: 820-828. Abstract in H.A.E. Ser. r\ 196 1. .f 9: IDG . W eiss, H . B. and H. B. Lott. 1922. The juniper \I·cb\l·ol'lll. Dil'hoillcris lIIargillc/llls Fabr. Enl. Ncws. 33: RO-R2. Wheeler. A. C. Jr. !D 78 . Studies on the fa ll na of alfal fa. \ ' 111. Parasitoid -host records. C an EnL 11 0: 11 17-111 9 . 54 J. ENTo~ l o L Soc . BnlT C OLL'.\ IIlIA 79 (1982). D Ec. 3 1. 1982

'Nhitehead . \V. E . 1926. Notes o n the current Iea froll er (Ca("() (' cia /"()S(l lI a L. ) in NO \'a Scoti a. Arcadia n en­ tomo\. Soc. Proe. 10: 76-7D . Wilkinson, A. T . S. 196.5 . Heleases of cinnaba r moth . lIypocri/a ja coiJa cac (L. ) (Lepidoptera: Arctiiuae) on tans\' ragwort in British Coillmbia. Proc. cntomo\. Soc. Brit. Co\. 62: 10- 13. Wilson. 19 12. Report fro m the Okanagan Distri ct: Insed pests of the \'car in the Okanagan . l'roc. entomo\. Soc. Brit. Co\. 2: 6. W vli e. H. C. HJ6 1. Insect parasites of the \\' inter llloth. OPl'ruplt/l'ra iJrllllla /a (L.) (Lepidoptera: Ceometridae) in \\'cs!t:rn Europe. Entolllophaga .5: 111 - 129.

SIPHONAPTERA FROM MAMMALS IN ALASKA. SUPPLEMENT IV. REVISED CHECK LIST FOR SOUTHEASTERN ALASKA GLENN E . H AAS'. L OYAL J O HNSON' . Ai\D R O BEHT E . W O OD '

ABSTRACT The annotated check list grc\\' to 1.5 tax a \\'ith the addition nl Opisor/ aSljS v('speralis (Jordan) a nd C lt ae/opsyl/a se/os(l Hothsl'hild. both ne\\' for Alask a: Tar­ sopsyl/a nc/odeci/lldc lI /n/a co loradellsis (Baker), nl'\\' for southeastern Al aska: and an Oropsyl/a sp. Nine species of \\'ild mammals and the domesti c ca t a re reported as hosts of neas in sO lltheaste rn Alaska for th e first til1le, increasin g the total I1lll n ber of mammali an hosts to 2.5 identified species. Se\'entv ne\\' records for 10 isla nds a nd the mainla nd arc comhi ned \\'ith published records in a d istribution tabl e. 'vl or­ phological va ri a ti ons in the posteri or ma rgin of sternum V[I of females of MOll op­ syllll.\' rilia/lls pro/illllS (jordan) a nd distinguishi ng l'haraders of three ot he r taxa are ill ustra ted.

INTRODUCTION AN NOTATED LIST The additio n of four fl eas to our first eheek list Pulicidae of mammal fl eas of southeastern Al aska (I [aas l'I 01. I . Cte,,()cepJw lides Jeli" Jelis (Bouche) 1980) raised thc total to 1.5 tax a . Especi alh­ This is the first record of fl eas in fes ti ng a cat in n ot e \\' o rth~ ' is a marmot fl ea of the genus Ompslllla. southeastern Al aska . The other three f1 eas a re kn own from British Col­ Record - l3aranof Isla nd , Sitka: two males, six umbia (Holland 1949). One of them . Ta/,.I'op.l'ljlla females (o ne gra\'id). o n cat. I .Xll . 1979. L. octodeci mdell/a/a coloradcl/ sis (Baler) is also kn own Joh nson. from Alaska west of the Yukon Territory (Holland 1963; Hopla 1965: lIaas a nd Wilson 1982). but H yst ri chopsy II idae Opisodasys vesperalis (J ordan) a nd Chae/opsylla 2. 1I11,,/riclr opsyl/o dippiei "pi",,/o Holi ancl (Figs . I setosa Rothschild a re new to the sta te of Alaska. a nd 2) New records a re presented fo r all fl eas in our first Nin e more specim ens were coll ccted. all from list except for Myodopsylla gel//ilis Jordan a nd martens ma ki ng a total of 16 fro m this mustelid and Rothschild a nd Chae/opslj l/a /Ilhe/,('/tia/ireps one from a mink. The ass ignment of southeaste rn (Bezzi). Alaska n specimens to this subspecies was facilitated !-.I am malia n hosts of th e fl eas a re listed (T a ble by dissecting male genita li a to ex pose tuhercles near I). The additional ten species res ili ted in a total of the inncr angle of sternum IX (Fig. I ). l\'ot a ll 25 identified hosts pillS an unkno\\'n species of tubercles are in fo cus but enough to show more than Micro/liS. These additions a rc thc hoan' marmot, Holl a nd (1 9.5 7) illustrated for II . d . /rll l/ca /a northern fl ying squirrel. sO llthern red-backed vo le, Holl a nd. [-Ie sho\\'ed th at tubercles a rc a bse nt in II . meado\\' \·ole. long-ta il ed vole, gray wolf, bl aek d . dippici Rot hschild. bear, ermine, wolverine, a nd domes ti c ca t. All ne\\' The aedeagus dissccted from our H . J. spilla/a a nd published loealit\· rt'cords a rc ta bulatcd 1)\· specimen is shown in Fig. 2. Aedeagi of H. dippici named isla nds a nd the mai nland (Table 2). (then known as H . gigas dippiei Rothschild) and II . oceidclI/alis (i. e . 1-1. a. sylva/i clIs Campos a nd Sta rk) fro m south of Canada were ill ustrated b\· 1.3.37 California St. . No, 7. Boule/I 'r Cit ~. \, \ '. U .S. A. H!) OO.) . Traub (1950) a nd Campos a nd Sta rk (1979). res pec­ ~ St at e of Ala ~b Oepartlllt'nt of Fi ~ h allll C a ll1t' , P.O , 13 0.\ · W ~), tively. Our specimen appears cl ose to T raub's, Sitka. A" . L.S.I\ , 99S3.S. although the scl erotizecl inner tubc curves toward .1 Stalt' of Ala." ka Departrnl'lIt uf Fi .., h am! CaIlH" 41:'1 ~ l ain St . . Hm . 20S. I\ l'iciJik<:i n, A K . L:.S.A. 8nnO i its ex it (Fi g. 2).