This is anopenaccess article undertheCCBY-NC-SAThis México. of de Autónoma responsibility Nacional the Universidad under Reviewing Peer Manuscript accepted: Corrected manuscript received: Manuscript received: http://dx.doi. org/10.18268/BSGM2021v73n3a301220 A301220. (3), 73 Mexicana, Pacific,USA: Geológica Sociedad la de Boletín (, ) from the Eastern , Berglundus bretoni n. gen., n. sp. Vega, F. J., 2021,Anewcyclodorippid Nyborg, T., Garassino, A.,Nyborg, B., How tocitethisarticle: [email protected] * Corresponding author:(F. J. Vega) Coyoacán, CdMx,04510Mexico. Universitaria, Ciudad México, de Autónoma 4 97078, USA. 3 92350, U.S.A. CA Linda, University,Loma Linda Loma es, 1,2 Torrey Noroeste delPacífico,EUA cangrejo Un nuevo ciclodorípido, Cyclodorippidae) fromtheNortheasternPacific,USA crab, A newcyclodorippid Berglundus bretonin.gen., n. sp. (Decapoda, license(https://creativecommons.org/licenses/by-nc-sa/4.0/) Instituto de Geología, Universidad Nacional Universidad Geología, de Instituto 40 W 7t Aeu, lh, Oregon Aloha, Avenue, 179th SW 4400 Department of Earth and Biological Scienc and Earth of Department Nyborg 1 , Alessandro Garassino Boletín de la Sociedad Geológica Mexicana Geológica Sociedad la de Boletín - Berglundus bretoni Washington State, USA. rippoidea, Miocene,, Keywords: Crustacea, Cyclodo- unchanged. the Cyclodorippidae distribution of range and the palaeogeographic the stratigraphic America, keeping herein cyclodorippids inNorth genusof is State described. Itrepresents thefourth Washington of Astoria Formation the (early to middle Miocene) from 1892) Ortmann, bretoni A newcyclodorippid crab, ABSTRACT 1 , Brant n.gen., n. sp. (Cyclodorippidae Nyborg 2 , Francisco J. n. gen., n. sp. (Decapoda, Cyclodorippidae) Berglundus Vega /73(3)A301220/2021 3,* de Washington, es descrito aquí. Este (Mioceno temprano amedio)delEstado Ortmann, 1892) de la Formación Astoria bretoni ciclodorípido, cangrejo Un nuevo RESUMEN Unidos de América. Unidos deAmérica. onomía, Washington, Estados Cyclodorippoidea, Mioceno, tax Palabras clave: Crustacea, intactos. ypaleobiogeográfico tigráfico Norteamérica, manteniendo elalcance estra es elcuarto género deciclodorípidos en n.gen.,sp. (Cyclodorippidae Berglundus 1 - -

A new cyclodorippid crab from the Northeastern Pacific, USA ABSTRACT INTRODUCTION / A new cyclodorippid crab from the Northeastern Pacific, USA GEOLOGICAL SETTING 2. Geologicalsetting 1. Introduction 2 2 fCyclodorippoidae (Cyclodorippinae and of 46 within species themextant. fossilrecord The genera, most of 231 includes 2007 Ng, and Colgan Sharkey, Lai, Ahyong, According toGuinot euneo early to middle Miocene, terrestrial sequence of Walsh 1987; Logan, 1989; 1979, Wells, 1972; 1968, Mckee, Willapa Hills area ( Washingtonsouthwestern inthe Montesano and in and Naselle, of town the near Creek, Salmon River,Naselle and the along outcrops east, the to Knapptononthewest andAltoona townsite of old the between especially River, Columbia the Washington,southwestern withoutcrops along AstoriaFormationThe iswidelydistributed in unchanged. the Cyclodorippidae remains distribution of range and palaeogeographic the stratigraphic from North America. Based upon thisnewreport, cyclodorippids representing the fourth genus of (USA), State Washington of Miocene) middle to glundus Washington State (USA). (Priabonian) of toma antiqua reported (2001) Finally,Schweitzer 2020). (for complete list see Nyborg andGarassino, Japan and (USA) Oregon and State,Washington Alaska, species from the Miocene to Pliocene of distributionspread palaeogeographic including 7 the family witha wide the best-known genus of del Fuego (Argentina). Tierra of 2011 Griffin, and dío Miocene the from Miotymolus quadratus Feldmann (USA). Texas of (Albian) Early the from 1983 Bishop, et al (Schweitzer respectively species 10 with genera 4 only to restricted scarce, very is Xeinostominae) / / Boletín de la Sociedad Geológica Mexicana Geológica Sociedad la de Boletín Boletín de la Sociedad Geológica Mexicana Geológica Sociedad la de Boletín ujc f hsnt stedsrpino of description the is note this of Subject ., 2017). Bishop (1983) described (1983) Bishop 2017). ., n. gen. from the Astoria Formation (early Schweitzer, 2001 from the late et al , 97. hs nt s at f a of part is unit This 1987). ., Feldmann, Schweitzer,Casa Feldmann, e.g ., Weaver,and Wolfe., 1937; Tymolus et al (03 te Section the (2013) . et al Stimpson, 1858 is 1858 Stimpson, http://dx.doi.org/10.18268/BSGM2021v73n3a301220 . (2011) reported Hillius youngi Xeinos Ber /73(3)A3012202021 - - - - /73(3)A3012202021 the unit below these basalt flows,basalt belowthese the unit fossil placing the of Formation Washington State appear to represent that part of Astoria the within Fossils 1987). the early-middle (Walsh Miocene boundary near dated been have which Group, Basalt River Columbia the of Basalt Wanapun the and Washington Basalt Grande Ronde the of with interfingers locally Formation Astoria The 1987). et al lithologic characteristics ( faunal contents,upon and position, stratigraphic Oregon based with the Astoria Formation of correlated tentatively and State, Washington of siltstone, portion exposedthe southwestern-most in conglomerate, sandstone, and and marine beds of have also been described from this locality,this from described been also have crabs ity andsurrounding outcrops. Two other fossil local the of details locality,faunal providingthis wahkiakumensis crab a fossil describe to first the were (1992) Goedert and Berglund Formation. Astoria area the correlating the outcrops at Pigeon Point of withthe map detailed a gave (1968) Mckee thatthe Pigeon contains Point area. Wolfe and Quandrangle RiverGrays the of area the map to State (USA) (Figure 1). Weaver (1937) was the first WashingtonAltoona, of town River,the bia near theColum Pigeon Point, alongthenorthshore of stone exposures ona west-facing road cut along described herein,were collected from sandy-silt 1822, found inbothStates (Nyborg, 2002). with onlyonegenus, State,Washington and Oregon of Formation ria fossilin from decapod crustaceanfaunas theAsto contemporaneous,be markedtherea is difference to WashingtonState in those and Oregon in tion FormaAstoria believedthe to thatrocksassigned have workers past and environments depositional a similarityin decapod crustaceanfaunassuggest and Foraminifera (Rau, 1948). Although the fossil 1987), Addicott, and Moore 1931; (Etherington, molluscs 1972), distributionthe biostratigraphical of Mckee, and (Wolfe composition isotope Miocene. Age control isalso baseduponmineral crabs described herein mostlikelywithinthe early l fthe specimens, except for one, as All of ., 1978; Wolfe and Mckee, 1972; Walsh1972; Mckee, Wolfeand 1978; ., (Berglund and Goedert, 1992) from Mursia e.g ., Rau, 1948; Snaveley1948; Rau, ., Leach inDesmarest, Anisospinos et al et al ., ., - - - - - 3. Systematicpalaeontology Figure 1 eto Ccooipia hog Li Sharkey, Lai, Ahyong, Cyclodorippoida Section to-orbital width;W4,posteriorwidth. fron W3, width; frontal W2, carapace; of width Abbreviations this locality. from described crab fossil other only the is 2015 Quadrangle, map. graphic Knappton the topo Washingtonseries minute 7.5 State-Oregon of R9W T10N, NE1/4, NW1/4, the in USA, State,Washington (Wolfe and Mckee, 1972) near the Naselle, town of FormationAstoria the from derived been have to interpreted Creek Salmon of gravels stream the and Vega, 2016. Macrocheira columbiaensis Lophomastix altoonaensis vex in both direction; rostrum tridentate, densely tridentate, rostrum direction; both in vex Diagnosis Superfamily Cyclodorippoidea Ortmann,1892 Mapdrawing showinglocality (X)where theapproximate Subfamily Cyclodorippinae Ortmann,1892 One additional specimen wascollected from Family Cyclodorippidae Ortmann, 1892 : Carapace subcircular and gently con Munida branti Colgan andNg,Colgan 2007 Genus L, egh f crpc; W1, carapace; of length L1, : Berglundus Nyborg and Vega, 2008 and Nyborg,Nyborg,Garassino Nyborg,Nyborg,Garassino Boletín de la Sociedad Geológica Mexicana Mexicana Mexicana Geológica Geológica Sociedad Sociedad la la de de Boletín Boletín Nyborg andGarassino, http://dx.doi.org/10.18268/BSGM2021v73n3a301220 nov. - - - Berglundus bretoni Berglundus carapace in outline andfor the presence orabsence difference the dis by other each are from tinguished which 1897, Bouvier, and 1892 Ortmann, Cyclodorippidae families, Discussion monotypy. Type species Gender: specimens. masculine. studied the donated who Etymology minute smoothposterolateral with granules; margins. fringed margin anterolateral cles; tuber 14 with inflated strongly and defined well regions directed; forward teeth triangular sharp, granular, axiallydepressed, endingintotwo small, having subsquare carapace and absence of orbits. of absence and havingcarapace subsquare we exclude their belonging to the Cymonomidae well-defined and inflated carapace regions. Indeed, orbits, and subcircular carapace, the presence of can be assigned to the Cyclodorippidae for the the Cyclodorippoidea, they in thesystematics of not preserve ventral parts, do useful diagnostic characters specimens studied the Although (USA). Texasfrom the Early Cretaceous (Albian) of genus sole one just including Quadratoplanidae, family new the Franțescudescribed Finally (2014) Phyllotymolinidae,three including extant genera. orbits. of Later,Tavaresfamily new the described (1998) n.gen.,sp.was collected. atr os egud deceased, Berglund, Ross after : /73(3)A301220/2021 Tvrs 19: 6) eotd two reported 264) (1993: Tavares : / 73(3)A301220/2021 : Berglundus bretoni n. gen., n. sp. by 3 3 - -

GEOLOGICAL SETTING / A new cyclodorippid crab from the Northeastern Pacific, USA SYSTEMATIC PALAEONTOLOGY A new cyclodorippid crab from the Northeastern Pacific, USA SYSTEMATIC PALAEONTOLOGY 4 4 imens from theQuadratoplanidae. carapace regionsthe studied spec also distinguish developed moderately and flat the and carapace specimens. the Finally, therectangular outline of distinguish the Phyllotymolinidae from the studied The carapace regions scarcely defined and inflated the Cyclodorippidae. of representatives the among unique are which 14 tubercles, with regions carapaces inflated the new genus Heard, 1984;Tavares, 1993). and (Goeke shape rostralvariable highly havinga 1899, and 1880, the subfamily, of such as genera extant some by proven as shape, the Cyclodorippinae, although the distinct rostral is uniqueamongthe specimens fossilgenera of The rostrum. fromthe studiedspecimens in two small, sharp, triangular teeth distinguishes ending rostrum, tridentateHowever, grooves.the and Garassino, 2020,andwell marked by deep Nyborg and Garassino, 2020 or tubercles insomefossilspecies, like with regions carapace inflated and granulate the inflated carapace regions withouttubercles. and from respectively); tubercles two and one with regions scarcely tuberculate (except protogastric and and cardiac inflated not regions carapace and fromstudied specimens the distinguish tubercles 14 with inflated strongly sharp, triangular teeth and the carapace regions and Tymolus (fossil); 2011 Griffin, and Casadío weitzer, (fossil); 1983 Bishop, genera: three includes Cyclodorippinae / / Boletín de la Sociedad Geológica Mexicana Geológica Sociedad la de Boletín Boletín de la Sociedad Geológica Mexicana Geológica Sociedad la de Boletín ae uo Schweitzer upon Based In conclusion, we justify the description of the of description the justify we conclusion, In share studied specimens with The tridentateThe rostrum, endingintwo small, Stimpson,1858(extant andfossil).

hp f h otu fthe studied the rostrum of shape of Berglundus bretoni Miotymolus Deilocerus Berglundus (Figures 2and3) A. Milne-Edwards and Bouvier,and Milne-Edwards A. Tavares, 1993, include species Coricodus having a bifid rostrum and rostrum bifid a having for tridentate rostrum and Miotymolus Hillius n.gen.,sp. Tymolus having a wide front A. Milne-Edwards, et al http://dx.doi.org/10.18268/BSGM2021v73n3a301220 edan Sch Feldmann, T. collinsi having a bifid havinga . (2017) the T. bottemilleri Nyborg Tymolus Hillius /73(3)A3012202021 - - /73(3)A3012202021 2.7, 97181, Measurements ria Formation. Geological age State, USA). Washington Seattle, Washington, of University Collections, Paleontology Invertebrate Culture, Natural and History BurkeIP = The Museum of (UWBM 97182-97193 IP UWBM paratypes and Type material Cenozoic fossil crabs. important his contribution to the knowledge of Etymology Diagnosis spines by a deep U-shaped notch (see Figure 2E: Figure (see notch U-shaped deep a by spines adjacent from separated inflated, rounded, large, notchFigureinner-orbital(see 2); spine 2E: spines U-shaped shallow by spines medial from arated rounded, triangular, sharp, laterally directed, sep side, either on spines of pair 3); spine 2E: Figure sharp, triangular directedspines forward (see densely tridentate, granular rostrum smooth; surface of tulose on tubercles andfrontal region, remainder deep by pus almost granular, surface carapace delineated grooves; inflated, width; strongly carapace maximum regions percent 96 length about longitudinally, and transversely vaulted Description: lected at PigeonPoint. the other col specimens regions match that of carapace howeverdorsal preserved, partially only is 97193 IP UWBM Specimen Naselle. of town the near Creek, Salmon along float as concretion a from collected was 1). 97193 IP UWBM (FigureParatype State Washington Altoona, of town the River,near Columbia the of shore north the lected from concretions along Pigeon Point, along col were 97182-97192 IP UWBM paratypes and Occurrence partially preserved. paratypeonly specimens = 7.1.Remainder of mately 97186, IP UWBM Paratype 4.4. = 10.9, approximately W4 12.8, L1 , axiallydepressed,two endingin small, = 5.8. Paratype UWBM IP 97185, IP UWBM Paratype 5.8. = =10.2, : Asfor thegenus. W1 : after Gerard Breton (1944-2019) for Te ooye WM P 97181 IP UWBM holotype The : Carapace subcircular, weakly = 12.8, = : early to middle Miocene, Asto Miocene, middle to early : Hltp UB I 97181, IP UWBM Holotype : : (in mm) Holotype UWBM IP UWBM Holotype mm) (in : W1 W1 =10.2, W2 =11.1, = 2.5, = W2 W2 W3 =1.1, L1 1.8, = = 5.0, = = approxi W3 L1 W4 W3 = = ------placement of thefourteen carapacetubercles. B)Holotype,UWBMIP97181, right-lateral viewof carapace.Arrow indicates outer-orbital Figure 2 Holotype, UWBM IP 97181,close-up view of rostrum withnumbers indicating the threespinesoftherostrum. Scale barequals 5mm. spine. C)Holotype, UWBM IP 97181,close-up viewof rostrum. D)Paratype UWBMIP 97185,general view of carapace dorsalsurface. E)

Berglundus bretoni Berglundus n.gen.,sp.,A)Holotype,UWBM IP97181,general viewof carapacedorsalsurface. Numbers 1-14represent Boletín de la Sociedad Geológica Mexicana Mexicana Mexicana Geológica Geológica Sociedad Sociedad la la de de Boletín Boletín http://dx.doi.org/10.18268/BSGM2021v73n3a301220 /73(3)A301220/2021 / 73(3)A301220/2021 5 5

SYSTEMATIC PALAEONTOLOGY/ A new cyclodorippid crab from the Northeastern Pacific, USA CONCLUSION A new cyclodorippid crab from the Northeastern Pacific, USA CONCLUSION Figure 3 6 6 portion largest with longitudinally inflated tuber inflated longitudinally with largest portion posteriorly grooveaxial and groove;anterior shallowby divided and deep sinuous, by separated elongate, transversely region, hepatic rectangular small, forward; curving weakly axially, directed small tubercles, subhepatic ridges with arow of grooves; deep by separated inflated, regions bital concave,subor medially and subhepatic rimmed; margin posterior posteriorly; tapering rounded, epibranchial region; smooth posterolateral margin hepatic region, secondonealong lateral margin of sub spines, one situated along lateral margin of rounded two with margin anterolateral granules; strongly convex nearly vertical, fringed margin with minute anterolateral width; carapace maximum percent 35 occupying width fronto-orbital jected; outer-orbital spine broadly triangular, weakly pro carapace spine 1); frontal width about 15 percent maximum / / Idealized linedrawings of Boletín de la Sociedad Geológica Mexicana Geológica Sociedad la de Boletín Boletín de la Sociedad Geológica Mexicana Geológica Sociedad la de Boletín width; orbits deep, longitudinally narrow; Berglundus bretoni Berglundus http://dx.doi.org/10.18268/BSGM2021v73n3a301220 n.gen.,sp.discussed herein. Linedrawing notdrawntoscale. /73(3)A3012202021 - - - - /73(3)A3012202021 natro agno on region; anterior granules margin on of and axi ridged with one median turbercle axially situated transversely anterior region; cardiac pentagonal anterior and lateral grooves; rounded, weakly sub and depression posterior by defined regions, axial other as much as half about only inflated region urogastric rectangular pits; gastric region deep two with metagastric depressed smooth tubercle; wider posteriorlywith anaxially medianrounded lar-shaped region, mesogastric narrow anteriorly, triangu one; anterior than larger slightly tubercle one rounded, anterior tubercle andoneposterior with region protogastric axially; directed groove, shallow,a narrow,by longitudinal half in divided longitudinally inflated, weakly triangular rounded, region protogastric spine; outer-orbital of base at poorly defined;groove extends transversely, ending depressed, transversely region epigastric tubercle; rounded small, with portion posterior small cle; - - - 4. Conclusion h Moee f Nrh n Suh mrc and Japan, that suggesting thecyclodorippids hadan America South and North of Miocene the reported from the Early Cretaceous (Albian) to and (fossil), 2011 Feldmann,Schweitzer,Griffin, and Casadío four genera the Cyclodorippidae includes fossilThe record of and appendagesnotpreserved. sternum region; metabranchial smooth tubercle; median one with region mesobranchial tubercle; median one with inflated strongly portion, terior pos triangular-shapedtubercle; median rounded, portion quadrate, strongly inflated, with one large, bydeep,narrowbranchiocardiacgroove; anterior posterior groove; cervical deep by separated rior epibranchial groove;ante deep by differentiated; parts two in divided region well not reduced, strongly region intestinal tubercle; cardiac of ally eettvso thegenus. resentatives of sole specimen,which isassmall the extant rep incomplete one upon based StateWashington of described (2001) Schweitzer forms. extant the than larger and Garassino, 2020) and assigned to Conkle 1984; Tomida, and Takeda 1952; (Imiazumi, State Washington and Japan, Alaska, reported pids fromthe Mioceneof (Davie,muddyFossilsubstrates 2002). cyclodorip sandy,and soft, inhabit typically and m. 90-1200 from ranging depths at body living (Davie, Poore,2004) 2002; asmall-sized Indo-West usually the have Pacific in family the of representatives extant The Brazil. northern to States United the from and region Pacific Indo-West the in mania Japan through the Indian OceanandsouthtoTas broad range extending from geographic northern a has presently Cyclodorippidae the 1996) (1993, time (Feldmann the family mostlikely predated that the origin of amphitropical distributionthe Miocene and in Xeinostoma Tymolus Xeinostoma antiqua Hillius tbig 12 (osl n extant) and (fossil 1920 Stebbing, Stimpson, 1858 (fossil and extant), and (fossil 1858 Stimpson, et al Bishop, 1983 (fossil), ., 2011). Based upon Tavaresupon Based 2011). ., Boletín de la Sociedad Geológica Mexicana Mexicana Mexicana Geológica Geológica Sociedad Sociedad la la de de Boletín Boletín from the late Eocene et al http://dx.doi.org/10.18268/BSGM2021v73n3a301220 , 06 Nyborg 2006; ., Tymolus Miotymolus are - - - - - References Acknowledgements making thismanuscript ready for publication. Stuttgart, in for and suggestions their comments Germany) Naturkunde, für Museum (Staatliches G.Schweigert and Japan) Gifu, Akeyo,Mizunami, Karasawa (Mizunami Fossil Museum, Yamanouchi, from the Pigeon Point locality. We also thankH. and V. additional specimens Nyborg for collection of described hereining specimens from Pigeon Point Weto thankR.Berglund,wish deceased, for donat ai, ... 20, rsae: : Crustacea: 2002, P.J.F., Davie, R.M., Feldmann, C.E., Schweitzer, L.N., Conkle, les classification, la Sur 1897, E.-L., Bouvier, Bishop, G.A., 1983, Fossil decapod from species new A J.L.,1992, Goedert R.E., Berglund, Ahyong, S., Lai, J.C.Y., Sharkey, D., Colgan, D.J., Ng, Brachyura, inWells, A.,Houston,W.W.K. Eucarida (Part 2), Decapoda-, Fossil Mizunami the Museum, 33,47-57. of Bulletin Alaska: of and areport on Blodgett, R.B., 2006, A new species of (1897), 54-70. 1896 (9), 8 Paris, de philomathique Société la de Bulletin Dorippidés: des famille origines et la distribution desCrabes de la https://doi.org/10.5962/bhl.part.28997 San 27-55. History,20, Natural the of Society Diego of Transactions Texas: central of the Lower Cretaceous, Glen Rose Limestone and Natural History, 9,1-11. Burke Museum Contributions inAnthropology Washington: in Formation Astoria Miocene of https://doi. org/10.1016/j.ympev.2007.03.022 576-86. 45(2), Phylogenetics Evolution, and Molecular RNA: ribosomal Podotremata based on smallsubunit nuclear of status The Decapoda): (Crustacea: crabs P.K.L., the brachyuran 2007,Phylogenetics of Cancer /73(3)A301220/2021 / 73(3)A301220/2021 Dcpd: rcyr) rm the from Brachyura) (Decapoda: Metacarcinus from the Miocene Tymolus 7 7 -

A new cyclodorippid crab from the Northeastern Pacific, USA ACKNOWLEDGEMENTS/ REFERENCES A new cyclodorippid crab from the Northeastern Pacific, USA REFERENCES 8 8 Desmarest, A. G., 1822, Les crustacés proprement maui R, 92 A icn crab, Miocene A 1952, R., Imiazumi, 2013, P., Castro, M., Tavares, D., Guinot, Goeke, G.D., Heard, R.W. Jr, 1984, A review the of the from decapods Fossil 2014, O., Franțescu, S., Casadío, C.E., Schweitzer, R.M., Feldmann, fauna and Stratigraphy T.J., 1931, Etherington, / / Boletín de la Sociedad Geológica Mexicana Geológica Sociedad la de Boletín Boletín de la Sociedad Geológica Mexicana Geológica Sociedad la de Boletín Levrault, Paris, 67-142. F.-G.géologiques. et zoologique rapports les sous fossiles crustacés des naturelle Histoire dits, inBrogniart, A., Desmarest, A.-G. (eds.), 19.3B, CSIRO Publishing, Australia. 641p. Australia, Vol.(eds), Zoologicalcatalogue of aaotlgcl oit o aa, (new Japan, series), 7,201-204. of Society Palaeontological the Transactionsand Proceedings of Formation kamadai https://doi.org/10.11646/ 1-414. zootaxa.3665.1.1 3665 Zootaxa, (1), taxa: podotreme ranked Brachyura), with newnominafor higher- brachyuran crabs(Crustacea, Decapoda, of and openings the sexual structuressupplementary onthe phylogeny of Significance grr.0704.06 https://doi.org/10.18785/ 351-355. 7(4), Clythrocerus https://doi. Mexico withnoteson of Gulf the eastern genus 221-239. org/10.1127/0077-7749/2014/0427 Geologie 273(3), für Jahrbuch und Paläontologie, Abhandlungen, Neues Texas: Tarrant County,Cretaceous (late Albian) of Carnegie https://doi. 91-123. of org/10.2992/007.079.0202 79(2), Annals Museum, implications: Paleobiogeographic Tierra Argentina: Fuego, from del Brachyura) (Thalassinidea; Decapoda Miocene New 2011, M., Griffin, 20(5), 31-142. California, of University the Department of Geology the of of Bulletin Washington: southwest the Astoria Miocene of of Clythrocerus n.sp.

stimpsoni f te oa Coal-field: Joban the of (Brachyura: Dorippidae) in (Brachyura:Dorippidae) from the Numanouchi : Gulf ResearchReports, Gulf : http://dx.doi.org/10.18268/BSGM2021v73n3a301220 Tymolus /73(3)A3012202021 /73(3)A3012202021

Milne-Edwards, A., 1880, Études préliminaires sur the Chehalis R.L., 1987, Logan, Geologic map of Nyborg, T., Garassino, A.,2020,New Neogene squat fossil New Nyborg,T.,2015, A., Garassino, Nyborg, T.G., 2002,Fossildecapod crustaceans Moore,J.,E. W.O., Addicott, Miocene The 1987, Les 1899, E.-L., Bouvier, A., Milne-Edwards, omnig Blei o h Msu o Comparative Harvard, Zoology U.S.N., 8(1),1-68. of Museum the Bartlett, of Bulletin J.R.Commanding: Commander and U.S.N., Sigsbee, D. C. Lieut-Commander “Blake”, Steamer Survey U.Coast S.the by and in the Caribbean Sea, 1877, 1878, 1879, Mexico, Alexander Agassiz, of in of the Gulf Supervision the under Dredging of Results the on Reports in Partie, 1ère Crustacés, les plate, scale1:100,000. 1 p., 17 87-8, Report File Open Resources Earth and Geology of Division Washington River and Westport quadrangles, Washington: njgpa/2020/0901 296(1-2), 167-174. https://doi.org/10.1127/ Geologie und Paläontologie, Abhandlungen, Neues Pacific: Jahrbuch für eastern the from (Decapoda, Brachyura, Cyclodorippidae) species of 95-101. Palaeodiversity, 8, Pacific: Eastern the from Munididae) Anomura: (Crustacea: lobsters State Kent University, Kent, Ohio, 224p. thesis, Master’s Unpublished U.S.A.Formation,Oregon, Washingtonand from the early tomiddleMioceneAstoria 1664, 13p. Bulletin, Survey U.S.Geological California: usefulness incorrelations from Alaskato Washington andOregon and their of Pillarian and Newportian (molluscan) stages Comparative Zoology, 27,1-127. of Museum the of Memoirs “Blake.”: Steamer SurveyU.S. Coast ‘79, the by ‘78, 1877, Sea, Caribbean the in and Mexico, Alexander Agassiz, of in of the Gulf Supervision the under Dredging of Results the on Reports in Oxystomes, et Dromiacés Tymolus tmsn 1858 Stimpson, a, .. 14, oaiiea rm the from Foraminifera 1948, W.W., Rau, Poore, G.C.B., 2004, Marine Decapod Crustacea des Dekapoden-Krebse Die 1892, A., Ortmann, fossil new Three 2008, F.J., Vega T., Nyborg Nyborg, T., Nyborg, B., Garassino, A.,Vega, F.J., cwizr CE, edan RM, Karasawa, R.M., Feldmann, C.E., Schweitzer, Tertiarythe to Additions 2001, Schweitzer,C.E., 1978, J.E., Pearl, A.R., Niem, P.D. Jr.,Snavely, pce fspecies of Mexicana, 5(2), 123-135. Paleontología Pacific: Eastern (Brachyura, ) from the North fossil of occurrences New 2016, https://doi.org/10.17161/to.v0i0.6659 1-6. 97, Online, 1, Treatise Cyclodorippoida: Volume Revised, R, Chapter 8R: Systematic descriptions: Section Part 2017, H., https://doi. df]2.0.co;2 521-537. org/10.1651/0278-0372(2001)021[0521:attt 21(2), Biology, of Northwest Journal America: Pacific North the of of fauna decapod A, A111-A120. 1977: U. S. Geological Survey Bulletin, 1457- nomenclaturebyU.S. the Survey,Geological Formations, Wright, W.B. (eds.), Changesinstratigraphic Pyshy N.F.,Sohl, in County,Washington, and Clallam the Hoko Makah, include River, to defined – Miocene) Lower to Eocene (Upper TwinGroup River 774-782. Paleontology,22(6), of Journal Washington: Miocene AstoriaFormation southwestern in Australia. 574p. Publishing, CSIRO Australia. Southern of https://doi. 532-588. 6, Biologie org/10.5962/bhl.title.15948 Thiere, und der Geographie für Systematik, Abtheilung Jahrbücher, Oxystomata: Zoologische und Dromiidae, , V.Abtheilungen Museums Die Strassburger https://doi. sol]2.0.co;2 361-369. Crustacean org/10.1651/0278-0372(2008)028[0361:tnf 28(2), Cenozoic of Journal Biology, the Washington: of from Blepharipodidae) Lophomastix Boletín de la Sociedad Geológica Mexicana Mexicana Mexicana Geológica Geológica Sociedad Sociedad la la de de Boletín Boletín http://dx.doi.org/10.18268/BSGM2021v73n3a301220 (Decapoda: Macrocheira aea M, oia S, 94 To e Fossil New Two 1984, S., Tomida, M., Takeda, Prodromus 1857-1860, W., Stimpson, Crustacea African T.R.R.,Stebbing,South 1920, el, .. 17, elgc a o h Cape the of map Geologic 1979, R.E., Wells, of stratigraphy Tertiary 1937, C.E., Weaver, Walsh, T.J., Korosec, M.A.,Phillips, W.M., nouvelle Plyllotymolinidae, 1998, M., Tavares, des systématique Révision 1996, M., Tavares, les Decapoda: Crustacea 1993, M., Tavares, cecs f Piaepi, Crustacea Philadelphia, Maioidea, 9(3),216-221. of Sciences Natural the Academy of Proceedings of Descripsit: et observavit Ducibus, Rodgers Federata Republica et JohanneCadwaladaromissa, Ringgold a Septentrionalem, evertebratorum, PacificumOceanum ad Expeditione in quae animalium descriptionis https://doi.org/10.5962/bhl.part.22316 231-272. 17(4), Museum, African South of the Annals Africa): South in Investigations Marine the for Crustacea, A. S. of X (Part iapitetNsle ie ae, Pacific area, Washington River Disappointment-Naselle of University Publications inGeology, 4,1-266. Oregon: Washingtonwestern and northwestern 1:250,000. Resources Geologic MapGM-34,scale Earth and Geology of Division Washington quadrant: southwest – Washington of map H.W.,Schasse,Geologic R.L., 1987, Logan, 109-122. 20(1), Zoosystema, Decapoda): (Crustacea, famille de Brachyoures Podotremata 233-295. naturelle,d’Histoirenational série,4 18(1-2), (Crustacea, Muséum du BulletinBrachyura): Decapoda, américains Cyclodorippidae naturelle, 156,253-313. d’Histoire national Muséum du Mémoires 10: volume MUSORSTOM, Campagnes Cymonomus l’Indo-Ouest-Pacifique à l’exclusion du genre Cyclodorippidae et Cymonomidae de the MizunamiFossil Museum,11,39-48. of Bulletin Japan: Central group, Mizunami the Tymolidae from the Miocene Crabs of /73(3)A301220/2021 / 73(3)A301220/2021 , in Crosnier, A. (ed.), Résultats des 9 9

A new cyclodorippid crab from the Northeastern Pacific, USA REFERENCES A new cyclodorippid crab from the Northeastern Pacific, USA REFERENCES 10 10 el, .. 18, elgc a o h Cape the of map Geologic 1989, R.E., Wells, Wolfe, E.W., McKee, E.H., 1968, Geology of the Wolfe,of E.W., Geology 1968, McKee,E.H., / / Boletín de la Sociedad Geológica Mexicana Geológica Sociedad la de Boletín Boletín de la Sociedad Geológica Mexicana Geológica Sociedad la de Boletín 1: 62,500. Scale 1832, IMAP Series, Numbered USGS and WahkiakumCounties, Washington. Pacificarea,River Naselle - Disappointment Open-File Report, 79-389. County,Survery U.S.Washington:Geologic ry Rvr udage Whikm and Wahkiakum quadrangle, River Grays http://dx.doi.org/10.18268/BSGM2021v73n3a301220 /73(3)A3012202021 /73(3)A3012202021 Wolfe, E.W., McKee, E.H., 1972, Sedimentary Sedimentary 1972, E.H., McKee, E.W., Wolfe, org/10.3133/b133 https://doi. 1-70. 1335, River Bulletin, Geologic Survery Grays S. U. the Washington: of quadrangle, rocks igneous and Geologic MapGM-4. Resources, Earth and Resources Geology of Natural Division of Department State Washington Washington: Counties, Pacific