DINOSAURS of UTAH Frank Decourten Dinosaurs of Utah

Total Page:16

File Type:pdf, Size:1020Kb

DINOSAURS of UTAH Frank Decourten Dinosaurs of Utah Second Edition DINOSAURS OF UTAH Frank DeCourten DINOSAURS OF UTAH DINOSAURS OF UTAH Second Edition Frank DeCourten The University of Utah Press | Salt Lake City Copyright © 2013 by The University of Utah Press. All rights reserved. The Defiance House Man colophon is a registered trademark of the University of Utah Press. It is based on a four-foot-tall Ancient Puebloan pictograph (late PIII) near Glen Canyon, Utah. 17 16 15 14 13 1 2 3 4 5 Library of Congress Cataloging-in-Publication Data DeCourten, Frank. Dinosaurs of Utah / Frank DeCourten. — Second Edition. pages cm Includes bibliographical references and index. ISBN 978-1-60781-264-7 (pbk. : alk. paper) — ISBN 978-1-60781-265-4 (ebook) 1. Dinosaurs—Utah. I. Title. QE862.D5D42 2013 567.909792—dc23 2013006059 Printed and bound in Korea. CONTENTS Preface vi 01. The Mesozoic World: The Earth in Upheaval 1 02. Dawn of the Utah Mesozoic: The Age of Dinosaurs Begins 23 03. The Early and Middle Jurassic: A Time of Transition 46 04. The Late Jurassic: The Golden Age of the Sauropods 75 05. Blood Brothers: The Predators of the Morrison Formation 120 06. The Early Cretaceous: The (Un)Missing Links 143 07. The Late Cretaceous: The Beasts of the Bayous 172 08. The Curtain Falls: The Dinosaurs of the North Horn Formation 210 09. Doing It: The Allure of Paleontology 234 Appendix: Classification of Dinosaurs 243 References 255 Index 283 Utah’s Dinosaur Record. Graphic developed by the Utah Geological Survey, available online at: http://geology.utah.gov/utahgeo/dinofossil/index.htm. PREFACE For a science that embraces an immense time strengthened their staffing, and emphasized paleon- dimension, twelve years can be a long time in pale- tology in their educational programs and outreach ontology. It was about that long after the publication activities. Examples include the Utah Natural His- of the first edition of Dinosaurs of Utah that Peter tory Museum operated by the University of Utah, DeLafosse initially suggested updating that book the Prehistoric Museum at the Utah State Univer- into the edition you are now reading. Peter’s sug- sity–Eastern campus, and Dixie State College. These gestion was prompted by his perception that a great public investments in preserving and investigat- deal of new information on Utah dinosaurs had sur- ing Utah’s fabulous paleontological resources have faced over the past decade and a half, and a new edi- had a dramatic impact. More paleontologists prob- tion was needed to incorporate the results of recent ably reside and work in Utah now than at any time advances in knowledge. He couldn’t have been more in the past. correct; it is only a slight exaggeration to declare Two important developments at the federal level that we have probably learned as much about Utah have helped to stimulate paleontological investi- dinosaurs over the past twenty years or so as was gations since the original edition of this book was learned during the preceding century. The surge published. In 1996 the Grand Staircase–Escalante of new discoveries about Utah dinosaurs and the National Monument was established in the magnif- world that they inhabited has been nothing short of icent landscapes of southwest Utah. The state lands explosive. Many new species of dinosaurs have been within the monument were consolidated in 1998, identified, their patterns of distribution and evolu- and today more than 1.9 million acres are set aside tion have been established and interpreted, and new for the benefit of future generations. Part of the analytical techniques have been applied to fossils management plan for the new monument entailed to formulate neoteric reconstructions of dinosaur a detailed survey of the paleontological resources anatomy and physiology. of the region, which includes extensive tracts of What stimulated the current renaissance in Utah remote land with difficult access. These surveys dinosaur paleontology? A number of factors have included rock exposures that had not been exam- contributed to the current wave of exciting research ined carefully for fossils. Many significant discover- in paleontology in the state. First, Utah is one of ies of dinosaur fossils were made by the monument only a handful of states that maintain an Office of scientists and their university and museum col- the State Paleontologist (Dr. James Kirkland), which leagues who conducted the surveys in this large is a part of the Utah Geological Survey. This office parcel of magnificent land. In addition, subse- was created in the late 1970s and has since become quent management plans have strongly supported of pivotal importance in encouraging, coordinating, the ongoing scientific studies of the fossils that and facilitating paleontological research in Utah. have been excavated from federal lands within the In addition, over the past two decades a number of monument. Also, the United States Paleontologi- Utah public institutions have significantly improved cal Resources Protection Act was passed and signed their facilities for paleontological research, into law in 2009. This legislation strengthened the vii viii Preface legal protection of vertebrate fossils on federal land in particular were gracious in sharing new informa- and provided for the care and preservation of fos- tion on Utah dinosaurs and fossil localities. Several sils for the public good. Federal agencies such as the individuals from the U.S. Bureau of Land Manage- Bureau of Land Management have carried out this ment also were very helpful in providing informa- legal mandate and, in the process, made strong con- tion on the status of paleontological work being tributions to the advancement of knowledge in ver- carried out on the lands under their jurisdiction. tebrate paleontology in Utah. The Utah Geological Survey has been a valuable The general public has also played a signifi- source of information for this book and contributed cant role in the recent progress of dinosaur paleon- several of the graphics contained herein. Some of tology in Utah. Utah Friends of Paleontology has the original dinosaur art by Carel Brest van Kempen grown from a small nascent organization in 1998 to is used again in this book. I am grateful for Carel’s a strong statewide network of trained amateur pale- talent and gift for creating such spectacular images ontologists who have made several significant fossil of nature, past and present. Dr. Ronald Blakey, discoveries and helped professional paleontolo- president of Colorado Plateau Geosystems, Inc., gists conduct their excavations and research pro- generously permitted the use of his excellent recon- grams. Along with the many individuals associated structions of ancient landscapes. Peter DeLafosse of with Utah’s public and private institutions, Utah the University of Utah Press, in addition to suggest- Friends of Paleontology has helped ignite the recent ing a second edition, was consistently supportive explosion in dinosaur studies by providing an army and patient during the long and often delayed pro- of dedicated dinosaur enthusiasts. The extraordi- cess of revision. My old partner Tom Hill provided nary level of public support of paleontology in Utah support and sustenance during our fieldwork for the today is unduplicated at any other time since the project, as he has continuously for several decades first dinosaur bones were discovered in the state of romping about together on the back roads of more than one hundred years ago. Utah. All of these developments would render any The overall goal of this book is the same as in 1998 compilation of information on Utah dinosaurs the original edition: to tell the story of Utah’s dino- somewhat dated. This is certainly the case with the saurs against the backdrop of changing Mesozoic first edition of Dinosaurs of Utah. Accordingly, the landscapes and ecosystems. Of course, we will never current edition is an attempt to update the content know all the details of that story: our knowledge of the original by incorporating the many advances of both the dinosaurs and the world they inhab- in our knowledge of Utah dinosaurs made over ited is still fragmentary and incomplete, regard- the past two decades. Such work, like paleontology less of how much we’ve learned in the recent past. itself, is always a collaborative effort. I have bene- It is almost certain that some of the enduring mys- fited enormously from correspondence and discus- teries that this book identifies will be closer to res- sions with many people in the process of compiling olution by the time it makes its way into the hands all the new information included in this book. Many of any reader. No end of the Utah dinosaur renais- of my scientific colleagues contributed to this effort. sance that began in the 1980s is in sight: it appears Dr. Ken Carpenter of the USU–Eastern Prehistoric that we can look forward to an indefinite future of Museum, Dr. Alan Titus of the Grand Staircase– captivating new discoveries. Whatever is learned in Escalante National Monument, Dr. Randy Irmis the future will enrich our understanding of one of of the Natural History Museum of Utah, and Scott the most fascinating periods in Utah’s deep natural Williams of the Burpee Museum of Natural History legacy, the Mesozoic era. However incomplete our Preface ix current knowledge is, the story of Utah’s dinosaurs reconstructing this story will help us understand the is nonetheless an extraordinary tale of life evolv- rapidly changing world that we have inherited from ing to changing conditions for more than 180 mil- the dinosaurs and craft a way to continue evolving lion years. I hope that the lessons we learn from in harmony with the planet we inhabit. 1.1. The Mummy Cliffs near Capitol Reef National Park. Courtesy John Telford. 1.2. This butte near Hanksville is capped by hard Mesozoic sandstone, while the badlands of the lower slopes are eroded from soft shale.
Recommended publications
  • September 2019 Competitive Oil and Gas Lease Sale Monticello Field Office DOI-BLM-UT-0000-2019-0003-OTHER NEPA -Mtfo-EA
    U.S. Department of the Interior Bureau of Land Management July 2019 September 2019 Competitive Oil and Gas Lease Sale Monticello Field Office DOI-BLM-UT-0000-2019-0003-OTHER NEPA -MtFO-EA Monticello Field Office 365 North Main PO Box 7 Monticello, UT 84535 DOI-BLM-UT-0000-2019-0003_Other NEPA-MtFO-EA July 2019 Table of Contents Chapter 1 Purpose & Need .................................................................................................................... 4 1.1 Project Location and Legal Description ........................................................................................ 4 1.2 Introduction ................................................................................................................................... 4 1.3 Background ................................................................................................................................... 4 1.4 Purpose and Need ......................................................................................................................... 6 1.5 Decision to be Made ..................................................................................................................... 6 1.6 Plan Conformance Review............................................................................................................ 6 1.7 Relationship to Statutes, Regulations, Policies or Other Plans ..................................................... 9 1.8 Issues Identified .........................................................................................................................
    [Show full text]
  • A New Troodontid Theropod, Talos Sampsoni Gen. Et Sp. Nov., from the Upper Cretaceous Western Interior Basin of North America
    A New Troodontid Theropod, Talos sampsoni gen. et sp. nov., from the Upper Cretaceous Western Interior Basin of North America Lindsay E. Zanno1,2*, David J. Varricchio3, Patrick M. O’Connor4,5, Alan L. Titus6, Michael J. Knell3 1 Field Museum of Natural History, Chicago, Illinois, United States of America, 2 Biological Sciences Department, University of Wisconsin-Parkside, Kenosha, Wisconsin, United States of America, 3 Department of Earth Sciences, Montana State University, Bozeman, Montana, United States of America, 4 Department of Biomedical Sciences, Ohio University College of Osteopathic Medicine, Athens, Ohio, United States of America, 5 Ohio Center for Ecology and Evolutionary Studies, Ohio University, Athens, Ohio, United States of America, 6 Grand Staircase-Escalante National Monument, Bureau of Land Management, Kanab, Utah, United States of America Abstract Background: Troodontids are a predominantly small-bodied group of feathered theropod dinosaurs notable for their close evolutionary relationship with Avialae. Despite a diverse Asian representation with remarkable growth in recent years, the North American record of the clade remains poor, with only one controversial species—Troodon formosus—presently known from substantial skeletal remains. Methodology/Principal Findings: Here we report a gracile new troodontid theropod—Talos sampsoni gen. et sp. nov.— from the Upper Cretaceous Kaiparowits Formation, Utah, USA, representing one of the most complete troodontid skeletons described from North America to date. Histological assessment of the holotype specimen indicates that the adult body size of Talos was notably smaller than that of the contemporary genus Troodon. Phylogenetic analysis recovers Talos as a member of a derived, latest Cretaceous subclade, minimally containing Troodon, Saurornithoides, and Zanabazar.
    [Show full text]
  • A New Xinjiangchelyid Turtle from the Middle Jurassic of Xinjiang, China and the Evolution of the Basipterygoid Process in Mesozoic Turtles Rabi Et Al
    A new xinjiangchelyid turtle from the Middle Jurassic of Xinjiang, China and the evolution of the basipterygoid process in Mesozoic turtles Rabi et al. Rabi et al. BMC Evolutionary Biology 2013, 13:203 http://www.biomedcentral.com/1471-2148/13/203 Rabi et al. BMC Evolutionary Biology 2013, 13:203 http://www.biomedcentral.com/1471-2148/13/203 RESEARCH ARTICLE Open Access A new xinjiangchelyid turtle from the Middle Jurassic of Xinjiang, China and the evolution of the basipterygoid process in Mesozoic turtles Márton Rabi1,2*, Chang-Fu Zhou3, Oliver Wings4, Sun Ge3 and Walter G Joyce1,5 Abstract Background: Most turtles from the Middle and Late Jurassic of Asia are referred to the newly defined clade Xinjiangchelyidae, a group of mostly shell-based, generalized, small to mid-sized aquatic froms that are widely considered to represent the stem lineage of Cryptodira. Xinjiangchelyids provide us with great insights into the plesiomorphic anatomy of crown-cryptodires, the most diverse group of living turtles, and they are particularly relevant for understanding the origin and early divergence of the primary clades of extant turtles. Results: Exceptionally complete new xinjiangchelyid material from the ?Qigu Formation of the Turpan Basin (Xinjiang Autonomous Province, China) provides new insights into the anatomy of this group and is assigned to Xinjiangchelys wusu n. sp. A phylogenetic analysis places Xinjiangchelys wusu n. sp. in a monophyletic polytomy with other xinjiangchelyids, including Xinjiangchelys junggarensis, X. radiplicatoides, X. levensis and X. latiens. However, the analysis supports the unorthodox, though tentative placement of xinjiangchelyids and sinemydids outside of crown-group Testudines. A particularly interesting new observation is that the skull of this xinjiangchelyid retains such primitive features as a reduced interpterygoid vacuity and basipterygoid processes.
    [Show full text]
  • Theropod Teeth from the Upper Maastrichtian Hell Creek Formation “Sue” Quarry: New Morphotypes and Faunal Comparisons
    Theropod teeth from the upper Maastrichtian Hell Creek Formation “Sue” Quarry: New morphotypes and faunal comparisons TERRY A. GATES, LINDSAY E. ZANNO, and PETER J. MAKOVICKY Gates, T.A., Zanno, L.E., and Makovicky, P.J. 2015. Theropod teeth from the upper Maastrichtian Hell Creek Formation “Sue” Quarry: New morphotypes and faunal comparisons. Acta Palaeontologica Polonica 60 (1): 131–139. Isolated teeth from vertebrate microfossil localities often provide unique information on the biodiversity of ancient ecosystems that might otherwise remain unrecognized. Microfossil sampling is a particularly valuable tool for doc- umenting taxa that are poorly represented in macrofossil surveys due to small body size, fragile skeletal structure, or relatively low ecosystem abundance. Because biodiversity patterns in the late Maastrichtian of North American are the primary data for a broad array of studies regarding non-avian dinosaur extinction in the terminal Cretaceous, intensive sampling on multiple scales is critical to understanding the nature of this event. We address theropod biodiversity in the Maastrichtian by examining teeth collected from the Hell Creek Formation locality that yielded FMNH PR 2081 (the Tyrannosaurus rex specimen “Sue”). Eight morphotypes (three previously undocumented) are identified in the sample, representing Tyrannosauridae, Dromaeosauridae, Troodontidae, and Avialae. Noticeably absent are teeth attributed to the morphotypes Richardoestesia and Paronychodon. Morphometric comparison to dromaeosaurid teeth from multiple Hell Creek and Lance formations microsites reveals two unique dromaeosaurid morphotypes bearing finer distal denticles than present on teeth of similar size, and also differences in crown shape in at least one of these. These findings suggest more dromaeosaurid taxa, and a higher Maastrichtian biodiversity, than previously appreciated.
    [Show full text]
  • PALEONTOLOGICAL TECHNICAL REPORT: 6Th AVENUE and WADSWORTH BOULEVARD INTERCHANGE PHASE II ENVIRONMENTAL ASSESSMENT, CITY of LAKEWOOD, JEFFERSON COUNTY, COLORADO
    PALEONTOLOGICAL TECHNICAL REPORT: 6th AVENUE AND WADSWORTH BOULEVARD INTERCHANGE PHASE II ENVIRONMENTAL ASSESSMENT, CITY OF LAKEWOOD, JEFFERSON COUNTY, COLORADO Prepared for: TEC Inc. 1746 Cole Boulevard, Suite 265 Golden, CO 80401 Prepared by: Paul C. Murphey, Ph.D. and David Daitch M.S. Rocky Mountain Paleontology 4614 Lonespur Court Oceanside, CA 92056 303-514-1095; 760-758-4019 www.rockymountainpaleontology.com Prepared under State of Colorado Paleontological Permit 2007-33 January, 2007 TABLE OF CONTENTS 1.0 SUMMARY............................................................................................................................. 3 2.0 INTRODUCTION ................................................................................................................... 4 2.1 DEFINITION AND SIGNIFICANCE OF PALEONTOLOGICAL RESOURCES........... 4 3.0 METHODS .............................................................................................................................. 6 4.0. LAWS, ORDINANCES, REGULATIONS AND STANDARDS......................................... 7 4.1. Federal................................................................................................................................. 7 4.2. State..................................................................................................................................... 8 4.3. County................................................................................................................................. 8 4.4. City.....................................................................................................................................
    [Show full text]
  • Was Dinosaurian Physiology Inherited by Birds? Reconciling Slow Growth in Archaeopteryx
    Was Dinosaurian Physiology Inherited by Birds? Reconciling Slow Growth in Archaeopteryx Gregory M. Erickson1,6*, Oliver W. M. Rauhut2, Zhonghe Zhou3, Alan H. Turner4,6, Brian D. Inouye1, Dongyu Hu5, Mark A. Norell6 1 Department of Biological Science, Florida State University, Tallahassee, Florida, United States of America, 2 Bayerische Staatssammlung fu¨r Pala¨ontologie und Geologie and Department of Earth and Environmental Sciences, LMU Munich, Mu¨nchen, Germany, 3 Key Laboratory of Evolutionary Systematics of Vertebrates, Institute of Vertebrate Paleontology & Paleoanthropology, Chinese Academy of Science, Beijing, China, 4 Department of Anatomical Sciences, Stony Brook University, Stony Brook, New York, United States of America, 5 Paleontological Institute, Shenyang Normal University, Shenyang, China, 6 Division of Paleontology, American Museum of Natural History, New York, New York, United States of America Abstract Background: Archaeopteryx is the oldest and most primitive known bird (Avialae). It is believed that the growth and energetic physiology of basalmost birds such as Archaeopteryx were inherited in their entirety from non-avialan dinosaurs. This hypothesis predicts that the long bones in these birds formed using rapidly growing, well-vascularized woven tissue typical of non-avialan dinosaurs. Methodology/Principal Findings: We report that Archaeopteryx long bones are composed of nearly avascular parallel- fibered bone. This is among the slowest growing osseous tissues and is common in ectothermic reptiles. These findings dispute the hypothesis that non-avialan dinosaur growth and physiology were inherited in totality by the first birds. Examining these findings in a phylogenetic context required intensive sampling of outgroup dinosaurs and basalmost birds. Our results demonstrate the presence of a scale-dependent maniraptoran histological continuum that Archaeopteryx and other basalmost birds follow.
    [Show full text]
  • The Sauropodomorph Biostratigraphy of the Elliot Formation of Southern Africa: Tracking the Evolution of Sauropodomorpha Across the Triassic–Jurassic Boundary
    Editors' choice The sauropodomorph biostratigraphy of the Elliot Formation of southern Africa: Tracking the evolution of Sauropodomorpha across the Triassic–Jurassic boundary BLAIR W. MCPHEE, EMESE M. BORDY, LARA SCISCIO, and JONAH N. CHOINIERE McPhee, B.W., Bordy, E.M., Sciscio, L., and Choiniere, J.N. 2017. The sauropodomorph biostratigraphy of the Elliot Formation of southern Africa: Tracking the evolution of Sauropodomorpha across the Triassic–Jurassic boundary. Acta Palaeontologica Polonica 62 (3): 441–465. The latest Triassic is notable for coinciding with the dramatic decline of many previously dominant groups, followed by the rapid radiation of Dinosauria in the Early Jurassic. Among the most common terrestrial vertebrates from this time, sauropodomorph dinosaurs provide an important insight into the changing dynamics of the biota across the Triassic–Jurassic boundary. The Elliot Formation of South Africa and Lesotho preserves the richest assemblage of sauropodomorphs known from this age, and is a key index assemblage for biostratigraphic correlations with other simi- larly-aged global terrestrial deposits. Past assessments of Elliot Formation biostratigraphy were hampered by an overly simplistic biozonation scheme which divided it into a lower “Euskelosaurus” Range Zone and an upper Massospondylus Range Zone. Here we revise the zonation of the Elliot Formation by: (i) synthesizing the last three decades’ worth of fossil discoveries, taxonomic revision, and lithostratigraphic investigation; and (ii) systematically reappraising the strati- graphic provenance of important fossil locations. We then use our revised stratigraphic information in conjunction with phylogenetic character data to assess morphological disparity between Late Triassic and Early Jurassic sauropodomorph taxa. Our results demonstrate that the Early Jurassic upper Elliot Formation is considerably more taxonomically and morphologically diverse than previously thought.
    [Show full text]
  • The Geologic History of Fossil Butte National Monument and Fossil Basin
    The Geologic History of Fossil Butte National Monument and Fossil Basin FOSSIL BUTTE The Geologic History of Fossil Butte National Monument and Fossil Basin NPS Occasional Paper No. 3 THE GEOLOGIC HISTORY OF FOSSIL BUTTE NATIONAL MONUMENT AND FOSSIL BASIN Paul O. McGrew Michael Casilliano Department of Geology, University of Wyoming National Park Service Occasional Paper Number Three 1975 TABLE OF CONTENTS index.htm Last Updated: 01-Mar-2005 http://www.nps.gov/history/history/online_books/fobu/index.htm[6/17/2013 12:42:49 PM] The Geologic History of Fossil Butte National Monument and Fossil Basin (Tables of Contents) FOSSIL BUTTE The Geologic History of Fossil Butte National Monument and Fossil Basin NPS Occasional Paper No. 3 CONTENTS COVER INTRODUCTION LOCATION HISTORICAL BACKGROUND OF RESEARCH STRATIGRAPHY Thaynes Limestone Evanston Formation Wasatch Formation Basal Conglomerate Member Lower Member Main Body Sandstone Tongue Mudstone Tongue Bullpen Member Tunp Member Age of the Wasatch Formation Depositional Environment Green River Formation Fossil Butte Member Angelo Member Age of the Green River Formation Depositional Environment Fowkes Formation Sillem Member Bulldog Hollow Member Gooseberry Member Age of the Fowkes Formation Depositional Environment QUATERNARY THE GEOLOGIC STRUCTURE OF FOSSIL BASIN PALEONTOLOGY Flora Invertebrates http://www.nps.gov/history/history/online_books/fobu/contents.htm[6/17/2013 12:42:58 PM] The Geologic History of Fossil Butte National Monument and Fossil Basin (Tables of Contents) Vertebrates Fish Amphibians Reptiles Birds Mammals PALEOECOLOGY AND TAPHONOMY ACKNOWLEDGMENTS GLOSSARY REFERENCES BIBLIOGRAPHY Library of Congress Cataloging in Publication Data McGrew, Paul Orman, 1909- The geological history of Fossil Butte National Monument and Fossil Basin.
    [Show full text]
  • A New Caenagnathid Dinosaur from the Upper Cretaceous Wangshi
    www.nature.com/scientificreports OPEN A new caenagnathid dinosaur from the Upper Cretaceous Wangshi Group of Shandong, China, with Received: 12 October 2017 Accepted: 7 March 2018 comments on size variation among Published: xx xx xxxx oviraptorosaurs Yilun Yu1, Kebai Wang2, Shuqing Chen2, Corwin Sullivan3,4, Shuo Wang 5,6, Peiye Wang2 & Xing Xu7 The bone-beds of the Upper Cretaceous Wangshi Group in Zhucheng, Shandong, China are rich in fossil remains of the gigantic hadrosaurid Shantungosaurus. Here we report a new oviraptorosaur, Anomalipes zhaoi gen. et sp. nov., based on a recently collected specimen comprising a partial left hindlimb from the Kugou Locality in Zhucheng. This specimen’s systematic position was assessed by three numerical cladistic analyses based on recently published theropod phylogenetic datasets, with the inclusion of several new characters. Anomalipes zhaoi difers from other known caenagnathids in having a unique combination of features: femoral head anteroposteriorly narrow and with signifcant posterior orientation; accessory trochanter low and confuent with lesser trochanter; lateral ridge present on femoral lateral surface; weak fourth trochanter present; metatarsal III with triangular proximal articular surface, prominent anterior fange near proximal end, highly asymmetrical hemicondyles, and longitudinal groove on distal articular surface; and ungual of pedal digit II with lateral collateral groove deeper and more dorsally located than medial groove. The holotype of Anomalipes zhaoi is smaller than is typical for Caenagnathidae but larger than is typical for the other major oviraptorosaurian subclade, Oviraptoridae. Size comparisons among oviraptorisaurians show that the Caenagnathidae vary much more widely in size than the Oviraptoridae. Oviraptorosauria is a clade of maniraptoran theropod dinosaurs characterized by a short, high skull, long neck and short tail.
    [Show full text]
  • M a R Y L a N D
    ' o J x l C a JJ¿ ' u¿»... /io hC i M A R T ü R J LES Y L w : A • i v & v ' :À N sr\lài«Q3/ D FRANK J. SCHWARTZ *IES 23102 Vlaams Instituut voor dl Zu Planden Marina Instituts MARYLAND TURTLE FRANK J. SCHWARTZ, Curator C h e s a p e a k e B i o l o g i c a l L a b o r a t o r y S o l o m o n s , M a r y l a n d University of Maryland Natural Resources Institute Educational Series No. 79 J u n e 1967 FOREWORD The 1961 publication of MARYLAND TURTLES resulted in an increased awareness of these interesting members of Maryland’s vertebrate fauna. New in­ formation stemming from this effort has been incorporated into this revision. The researches of Dr. J. Crenshaw, Jr. on the genus Pseudemys, especially Pseudemys floridana, Florida Cooter, has resolved much of the confusion regarding this species’ true distribution and systematica in the state. Its occurrence must now be relegated to an "introduced” or "escape” category. Additional information is also on hand to confirm the Bog Turtle’s tenacious survival in swampy-bog habitats adjacent to the Susquehanna River. Recent information has helped delineate the occurrence of the Atlantic Ridley turtle in the upper Chesapeake Bay. A new section has been added which discusses fossil turtles. It is hoped this edition will maintain interest in and further expand our knowledge of the turtles of the area.
    [Show full text]
  • Uppermost Cretaceous and Tertiary Stratigraphy of Fossil Basin, Southwestern Wyoming
    Uppermost Cretaceous and Tertiary Stratigraphy of Fossil Basin, Southwestern Wyoming By STEVEN S. ORIEL and JOSHUA I. TRACEY, JR. GEOLOGICAL SURVEY PROFESSIONAL PAPER 635 New subdivisions of the J,ooo-Joot-thick continental Evanston, Wasatch, Green River, and Fowkes Formations facilitate understanding of sediment genesis and Jl7yoming thrust-belt tectonic events UNITED STATES GOVERNMENT PRINTING OFFICE, WASHINGTON 1970 UNITED STATES DEPARTMENT OF THE INTERIOR WALTER J. HICKEL, Secretary GEOLOGICAL SURVEY William T. Pecora, Director Library of Congress catalog-card No. 70-604646 For sale by the Superintendent of Documents, U.S. Government Printing Office Washington, D.C. 20402 - Price 65 cents (paper cover) CONTENTS Page Wasatch Formation-Continued Abstract __________________________________________ _ 1 Fossils and age-Continued· Page Introduction ______________________________________ _ 2 Tunp Member______________________________ 28 Purpose ______________________________________ _ 2 Origin--------~-------------------------------- 28 Earlier work_ .. __ - __ - ___________________ - _-- _- __ 2 Tectonic implications ____________ -_-------------- 29 Acknowledgments __ . ___________________________ _ 2 Green River Formation ___ .. _______ ------------------ 30 General relations ___ -- _________________________ _ 5 Name and usage __________________ -------------- 30 Evanston Formation _______________________________ _ 5 Definition __________________ -_-------------- 30 N arne and usage _______________________________ _ 5 Lithologic heterogeneity.
    [Show full text]
  • Implications for Predatory Dinosaur Macroecology and Ontogeny in Later Late Cretaceous Asiamerica
    Canadian Journal of Earth Sciences Theropod Guild Structure and the Tyrannosaurid Niche Assimilation Hypothesis: Implications for Predatory Dinosaur Macroecology and Ontogeny in later Late Cretaceous Asiamerica Journal: Canadian Journal of Earth Sciences Manuscript ID cjes-2020-0174.R1 Manuscript Type: Article Date Submitted by the 04-Jan-2021 Author: Complete List of Authors: Holtz, Thomas; University of Maryland at College Park, Department of Geology; NationalDraft Museum of Natural History, Department of Geology Keyword: Dinosaur, Ontogeny, Theropod, Paleocology, Mesozoic, Tyrannosauridae Is the invited manuscript for consideration in a Special Tribute to Dale Russell Issue? : © The Author(s) or their Institution(s) Page 1 of 91 Canadian Journal of Earth Sciences 1 Theropod Guild Structure and the Tyrannosaurid Niche Assimilation Hypothesis: 2 Implications for Predatory Dinosaur Macroecology and Ontogeny in later Late Cretaceous 3 Asiamerica 4 5 6 Thomas R. Holtz, Jr. 7 8 Department of Geology, University of Maryland, College Park, MD 20742 USA 9 Department of Paleobiology, National Museum of Natural History, Washington, DC 20013 USA 10 Email address: [email protected] 11 ORCID: 0000-0002-2906-4900 Draft 12 13 Thomas R. Holtz, Jr. 14 Department of Geology 15 8000 Regents Drive 16 University of Maryland 17 College Park, MD 20742 18 USA 19 Phone: 1-301-405-4084 20 Fax: 1-301-314-9661 21 Email address: [email protected] 22 23 1 © The Author(s) or their Institution(s) Canadian Journal of Earth Sciences Page 2 of 91 24 ABSTRACT 25 Well-sampled dinosaur communities from the Jurassic through the early Late Cretaceous show 26 greater taxonomic diversity among larger (>50kg) theropod taxa than communities of the 27 Campano-Maastrichtian, particularly to those of eastern/central Asia and Laramidia.
    [Show full text]