Cloudbridge of Nature Reserve 2 km este de escuela San Gerardo de Rivas Perez Zeledon, www.cloudbridge.org

To book at tour, contact: [email protected]

For cabin rentals, contact: [email protected] Checklist &

Data inquiries or Monthly requests, contact: Abundances [email protected] October 2018

Contributors Alex Acott, Laurie Allnatt, Sarah Barnes, Kasey Bedford, Roopak Bhatt, Anna Bowland, Logan Bradley, Léo Chevillon, Sam Davies, Ryan Dibala, eBird, Baley Good, Neil Hancart, Olivia Hess, Spencer Kane, Nathan Marcy, Emilio Masotti-Black, Emma Noyes, Eugene Paradis, Brianna Powrie, Anna Rabone, James Ratcliff, Jeffrey Roth, Matt Smokoska, Frank Spooner, Richard Stanley, Chris Staunton, Amy Sutley, Joseph Taylor, Mélanie Thierry, Debbie Valliere, Willem Van Doorninck, Amelia Westhoff, Elisa Yang, & other interns and volunteers from Global Vision International. Photo: Green-crowned Brilliant (female).

Thijs Claes Quetzal. Timo Jäger Photo: Resplendent Photo: Ornate Hawk-Eagle (juvenile). Frank Spooner Checklist Key

Bird presence designations (except for Occasional or Accidental sightings) were calculated using the 2015-2018 data from Cloudbridge’s long-term bird monitoring survey. Designations are based on the percentage of total survey days when that bird was seen. Occasional and Accidental designations include birds seen outside of official surveys, and citizen science data (eBird 2018) provided the bird identification was from a trusted source or could be verified.

V Very Common: Sighted >75% of survey days

C Common: Sighted 50 - 75% of survey days

F Fairly Common: Sighted 25 - 50% of survey days

U Uncommon: Sighted 5 - 25% of survey days Birding in the Clouds Rare: Sighted >0 - 5% of survey days, every or R almost every year Cloudbridge is a 283 hectare (700 acre) cloud forest reserve on the Pacific slope of the Talamanca O Occasional: Sighted occasionally, but not every year mountains in Costa Rica. A reforestation project, the Accidental: Only 1 or 2 individuals ever reported, or A reserve is home to well established regenerating sightings occur over 4 years apart forest, as well as areas of primary montane (e) Endemic to Costa Rica and forest. At an altitude of 1500-2600 m (4900-8500 eBird. 2018. eBird: An online database of bird distribution and abundance ft), almost 300 mid- and high elevation bird species [web application]. eBird, Ithaca, New York. Available : http:// have be found within our borders. www.ebird.org. (Accessed: September 30, 2018).

Our hiking trails are open 365 days a year. Entry is by donation ($6 USD or 3000 colones is typical), to help keep the reserve accessible to all. All donations go back to the reserve to help maintain the trails, and fund research and education projects, like our long-term bird monitoring study.

Bird tours with local, knowledgeable guides can be arranged upon request. See our website for more information. Photo: Bay-headed Tanager. Tanager. Photo: Bay-headed Spooner Frank Tinamous J F M A M J J A S O N D Highland Tinamou Nothocercus bonapartei O O O O O O O Great Tinamou Tinamus major A Chachalacas, Guans, & Curassows Black Guan (e) Chamaepetes unicolor U F F U U U F U U F F U Gray-headed Chachalaca Ortalis cinereiceps O O O O New World Quail Buffy-crowned Wood-Partridge Dendrortyx leucophrys O O O O Spotted Wood-Quail Odontophorus guttatus F F F F F C C U F F C C Black-breasted Wood-Quail (e) Odontophorus leucolaemus A A Frigatebirds Magnificent Frigatebird Fregata magnificens A A Herons, Egrets, & Bitterns Cattle Egret Bubulcus ibis O O O O O Fasciated Tiger-Heron Tigrisoma fasciatum A Sandpipers Spotted Sandpiper Actitis macularius A Rails Gray-cowled Wood-Rail Aramides cajaneus R R R R R Vultures & Condors Turkey Vulture Cathartes aura U U U U U U U R R U U Black Vulture Coragyps atratus U U U U U U U U R R R C Osprey Osprey Pandion haliaetus O O Hawks, Eagles & Kites Short-tailed Hawk Buteo brachyurus R R R U Red-tailed Hawk Buteo jamaicensis U R R R R Broad-winged Hawk Buteo platypterus R R R R Swainson's Hawk Buteo swainsoni O O Hawks, Eagles & Kites (continued) J F M A M J J A S O N D Great Black Hawk Buteogallus urubitinga A Swallow-tailed Kite Elanoides forficatus R R R R R R Double-toothed Kite Harpagus bidentatus A A A A Gray-headed Kite Leptodon cayanensis A Barred Hawk Morphnarchus princeps O O O O Roadside Hawk Rupornis magnirostris O O O Ornate Hawk-Eagle Spizaetus ornatus U R R U U U U U Falcons & Caracaras Merlin Falco columbarius A Bat Falcon Falco rufigularis R U Laughing Falcon Herpetotheres cachinnans A A A A A Barred Forest-falcon Micrastur ruficollis R R R R Yellow-headed Caracara Milvago chimachima U R R R

Owls (night observations) Unspotted Saw-whet Owl Aegolius ridgwayi U U U U Mottled Owl Ciccaba virgata F V C C V F C F C F U F Costa Rican Pygmy-Owl (e) Glaucidium costaricanum F U Tropical Screech-Owl Megascops choliba A Bare-shanked Screech-Owl (e) Megascops clarkii U U U F F U U U U U Striped Owl Pseudoscops clamator A Nightjars Common Pauraque Nyctidromus albicollis R R R R Swifts Costa Rican Swift (e) Chaetura fumosa O O Vaux's Swift Chaetura vauxi R R R R Spot-fronted Swift Cypseloides cherriei A Lesser Swallow-tailed Swift Panyptila cayennensis O O Chestnut-collared Swift Streptoprocne rutila O O O White-collared Swift Streptoprocne zonaris U R U R R R U R R U J F M A M J J A S O N D Snowy-bellied (e) Amazilia edward U U R R U U R R U U U F Steely-vented Hummingbird Amazilia saucerrottei U Rufous-tailed Hummingbird Amazilia tzacatl U R U R U U U R R U F

Magenta-throated Woodstar (e) Calliphlox bryantae U U U R U U U Violet Sabrewing Camplyopterus hemileucurus U R U R R U U U U F F F Garden Emerald (e) Chlorostilbon assimilis O O

Lesser Violetear Colibri cyanotus C C F R U U F F U U F C Brown Violetear Colibri delphinae R U R U

Green-fronted Lancebill Doryfera ludovicae R U White-tailed Emerald (e) Elvira chionura R U R R R R U U R Talamanca Hummingbird (e) Eugenes spectabilis R R U U U

Stripe-tailed Hummingbird Eupherusa eximia F F U U U F F F F F F F White-necked Jacobin Florisuga mellivora O O O

Bronzy Hermit Glaucis aeneus U R Green-crowned Brilliant Heliodoxa jacula U R U U F F U U U U U U Long-billed Starthroat Heliomaster longirostris O O O O

Purple-crowned Fairy Heliothryx barroti U U R U U U U U U Violet-headed Hummingbird Klais guimeti R U R

Purple-throated Mountain-Gem (e) Lampornis calolaemus R R R White-throated Mountain-gem (e) Lampornis castaneoventris C F F F C C C F F F C C Fiery-throated Hummingbird (e) Panterpe insignis O O O O O

Scaly-breasted Hummingbird Phaeochroa cuvierii A Green Hermit Phaethornis guy U U U U R U U U U F F F Long-billed Hermit Phaethornis longirostris R R R R

Stripe-throated Hermit Phaethornis striigularis O O O O Volcano Hummingbird (e) flammula R R R R R R

Scintillant Hummingbird (e) Selasphorus scintilla F U F C F F F U U U F F Pigeons & Doves J F M A M J J A S O N D Plain-breasted Ground-Dove Columbina minuta A A

Ruddy Ground-Dove Columbina talpacoti R R R R R Gray-headed Dove Leptotila plumbeiceps A White-tipped Dove Leptotila verreauxi R R R R Band-tailed Pigeon Patagioenas fasciata U R U U U R R Scaled Pigeon Patagioenas speciosa A A

Ruddy Pigeon Patagioenas subvinacea R R R R R Chiriqui Quail-Dove (e) Zentrygon chiriquensis A Buff-fronted Quail-Dove (e) Zentrygon costaricensis R R R R True Parrots Barred Parakeet Bolborhynchus lineola U U R R R White-crowned Parrot Pionus senilis R R R R R U

Crimson-fronted Parakeet (e) Psittacara finschi R R R U Brown-hooded Parrot Pyrilia haematotis U R R U U R Sulphur-winged Parakeet (e) Pyrrhura hoffmanni F C F C C C V F C F F F Red-fronted Parrotlet (e) Touit costaricensis R R U Cuckoos Black-billed Cuckoo Coccyzus erythropthalmus O O

Smooth-billed Ani Crotophaga ani O O Squirrel Cuckoo Piaya cayana F U U U U R U U F F F F Trogons & Quetzals Resplendent Quetzal Pharomachrus mocinno R U U U U U U U R R U Gartered Trogon Trogon caligatus O O O Collared Trogon Trogon collaris F F U U U F C U F F C F

Slaty-tailed Trogon Trogon massena O O O Motmots Lesson's Motmot Momotus lessonii U U U U U U U U U U U U Barbets J F M A M J J A S O N D Red-headed Barbet Eubucco bourcierii F F F F U F F U F F F F Toucans Northern Emerald-Toucanet Aulacorhynchus prasinus C C C C C C C F C C F C Fiery-billed Aracari (e) Pteroglossus frantzii O O O O O Woodpeckers Golden-olive Woodpecker Colaptes rubiginosus R R R R R U R Lineated Woodpecker Dryocopus lineatus O Acorn Woodpecker Melanerpes formicivorus R R R U U R R R

Red-crowned Woodpecker Melanerpes rubricapillus U R R U U U Smoky-brown Woodpecker Picoides fumigatus R R U Hairy Woodpecker Picoides villosus U U U U U U U U F U U Rufous-winged Woodpecker (e) Piculus simplex O O Olivaceous Piculet Picumnus olivaceus R R U

Yellow-bellied Sapsucker Sphyrapicus varius A Ovenbirds Scaly-throated Foliage-gleaner Anabacerthia variegaticeps R R R Chiriqui Foliage-gleaner (e) Automolus exsertus R R R U R R Buff-throated Foliage-gleaner Automolus ochrolaemus R R U U U U R R Brown-billed Scythebill Campylorhamphus pusillus R U F R R R U

Red-faced Spinetail Cranioleuca erythrops F U F U U U C U U F C U Tawny-winged Woodcreeper Dendrocincla anabatina A Black-banded Woodcreeper Dendrocolaptes picumnus A A Wedge-billed Woodcreeper Glyphorynchus spirurus R R R R Spot-crowned Woodcreeper Lepidocolaptes affinis U U F F F C C F F F U U

Streak-headed Woodcreeper Lepidocolaptes souleyetii R U R R R R U Ruddy Treerunner (e) Margarornis rubiginosus R R R U U U R U R Spotted Barbtail Premnoplex brunnescens U U U U U F F U U U F F Buffy Tuftedcheek Pseudocolaptes lawrencii R U U R Ovenbirds (continued) J F M A M J J A S O N D Tawny-throated Leaftosser Sclerurus mexicanus R R R U

Olivaceous Woodcreeper Sittasomus griseicapillus R R R R U R Lineated Foliage-gleaner Syndactyla subalaris U R U U U F U U U U F Streak-breasted Treehunter (e) Thripadectes rufobrunneus U R U R R U R R R U Plain Xenops Xenops minutus R R R Streaked Xenops Xenops rutilans U R U U U R R U U

Spotted Woodcreeper Xiphorhynchus erythropygius U Antbirds Slaty Antwren Myrmotherula schisticolor U R U U U R R U R Tapaculos Silvery-fronted Tapaculo (e) Scytalopus argentifrons U R R R R R R Tyrant Flycatchers Bright-rumped Attila Attila spadiceus R R R U Tropical Pewee Contopus cinereus U R U U R Olive-sided Flycatcher Contopus cooperi R R F U U

Dark Pewee (e) Contopus lugubris U U R Ochraceous Pewee (e) Contopus ochraceus R U R Western Wood-Pewee Contopus sordidulus U U R U Eastern Wood-Pewee Contopus virens R R R

Lesser Elaenia Elaenia chiriquensis R U U R Yellow-bellied Elaenia Elaenia flavogaster R R R R Mountain Elaenia Elaenia frantzii R R U U R U U U U Black-capped Flycatcher (e) Empidonax atriceps U R R R U U F Yellowish Flycatcher Empidonax flavescens F F F F F F F F F F F F

Yellow-bellied Flycatcher Empidonax flaviventris U U U U U R U Least Flycatcher Empidonax minimus R R R U U Piratic Flycatcher Legatus leucophaius R R Slaty-capped Flycatcher Leptopogon superciliaris U R U U U U U R R R U Tyrant Flycatchers (continued) J F M A M J J A S O N D Scale-crested Pygmy-Tyrant Lophotriccus pileatus R R U F R R R R Boat-billed Flycatcher Megarhynchus pitangua R R U U U U U U U R U Ochre-bellied Flycatcher Mionectes oleagineus R R R

Olive-striped Flycatcher Mionectes olivaceus U U U R U U U U U U R U Tufted Flycatcher Mitrephanes phaeocercus U U U U U U U U U F F F Great Crested Flycatcher Myiarchus crinitus O

Dusky-capped Flycatcher Myiarchus tuberculifer U U U U R U F U U U U U Golden-bellied Flycatcher (e) Myiodynastes hemichrysus U F U U U U U U U U U U

Sulphur-bellied Flycatcher Myiodynastes luteiventris O O Streaked Flycatcher Myiodynastes maculatus A Gray-capped Flycatcher Myiozetetes granadensis U R R R

Social Flycatcher Myiozetetes similis U R U U U R R R Rough-legged Tyrannulet Phyllomyias burmeisteri O O

Great Kiskadee Pitangus sulphuratus U R U U U F U U U F White-throated Spadebill Platyrinchus mystaceus R R Eye-ringed Flatbill Rhynchocyclus brevirostris R R U U R

Black Phoebe Sayornis nigricans U R R R U U U R R Torrent Tyrannulet Serpophaga cinerea R R R R U R U U U

Common Tody-flycatcher Todirostrum cinereum U R R R R U U U Yellow-olive Flycatcher Tolmomyias sulphurescens O O O O O Tropical Kingbird Tyrannus melancholicus U U U U R U R U U U

Paltry Tyrannulet Zimmerius vilissimus F U F F U R F U F F F F Tityras & Allies Rose-throated Becard Pachyramphus aglaiae A A Black-and-white Becard Pachyramphus albogriseus R R R U

Barred Becard Pachyramphus versicolor U R U R U U U U U U R Masked Tityra Tityra semifasciata U R U R R U U R U R Manakins J F M A M J J A S O N D White-ruffed Manakin Corapipo altera A Cotingas Three-wattled Bellbird (e) Procnias tricarunculatus A A A Crows, Jays, Magpies & Allies Silvery-throated Jay (e) Cyanolyca argentigula O O O O Brown Jay Psilorhinus morio U U U U R U U U U Swallows Barn Swallow Hirundo rustica A Gray-breasted Martin Progne chalybea A Blue-and-white Swallow Pygochelidon cyanoleuca U R R R R U R

Southern Rough-winged Swallow Stelgidopteryx ruficollis U R R R R R U Northern Rough-winged Swallow Stelgidopteryx serripennis R R R R R Thrushes Orange-billed Nightingale- Catharus aurantiirostris R R U R R U Ruddy-capped Nightingale-Thrush Catharus frantzii U U F F F F F U U U U U Slaty-backed Nightingale-Thrush Catharus fuscater R U U F R R U U

Black-billed Nightingale-Thrush (e) Catharus gracilirostris R R R U U R R U Swainson's Thrush Catharus ustulatus U R F C U C F U Wood Thrush Hylocichla mustelina A Black-faced Solitaire (e) Myadestes melanops V V C C C C V F V V V V White-throated Thrush Turdus assimilis O O O

Clay-colored Thrush Turdus grayi U U U U U U R U U U U U (e) Turdus nigrescens U R R Mountain Thrush Turdus plebejus F F U U R U U U U F F C Dippers American Dipper Cinclus mexicanus U U R Mockingbirds Tropical Mockingbird Mimus gilvus R R R Silky-flycatchers J F M A M J J A S O N D Black-and-yellow Silky-flycatcher (e) Phainoptila melanoxantha O O O O Long-tailed Silky-flycatcher (e) Ptiliogonys caudatus R R F F U U U U R Wrens Isthmian Wren (e) Cantorchilus elutus R R R U U R R U R

Gray-breasted Wood-Wren Henicorhina leucophrys F F F F U C C F C C C V White-breasted Wood-Wren Henicorhina leucosticta R R R R

Scaly-breasted Wren Microcerculus marginatus U F F F C C U F F C U Rufous-breasted Wren Pheugopedius rutilus F U U U U U F U F U F R House Wren Troglodytes aedon U R U U U U U R U U U U

Ochraceous Wren (e) Troglodytes ochraceus R U R U U U F U U U U R Vireos Rufous-browed Peppershrike Cyclarhis gujanensis U R U U R R R U U U U

Lesser Greenlet Pachysylvia decurtata U R U R R R R R Yellow-winged Vireo (e) Vireo carmioli R R U U U R Yellow-throated Vireo Vireo flavifrons U U U U U U

Yellow-green Vireo Vireo flavoviridis O Brown-capped Vireo Vireo leucophrys F U C C F C C F C C C C Red-eyed Vireo Vireo olivaceus A

Philadelphia Vireo Vireo philadelphicus F F F F U U U C Green Shrike-Vireo Vireolanius pulchellus O O O New World Warblers Golden-crowned Warbler Basileuterus culicivorus U U F U F U F F U F F C Black-cheeked Warbler (e) Basileuterus melanogenys U U U U U U U R U F U U Costa Rican Warbler (e) Basileuterus melanotis U U F U U F U U U U U F

Rufous-capped Warbler Basileuterus rufifrons O O O O Wilson's Warbler Cardellina pusilla F F V C R F V V V

Kentucky Warbler Geothlypis formosa R R R Mourning Warbler Geothlypis philadelphia A A A New World Warblers (continued) J F M A M J J A S O N D MacGillivray's Warbler Geothlypis tolmiei A Worm-eating Warbler Helmitheros vermivorum U U U U Black-and-white Warbler Mniotilta varia F F F R C V V C Slate-throated Redstart Myioborus miniatus V V C V V V V V V V V V Collared Redstart (e) Myioborus torquatus U R U U U R U U U U U Buff-rumped Warbler Myiothlypis fulvicauda R U R U U U U Flame-throated Warbler (e) Oreothlypis gutturalis U U U U R U U U F F U U Tennessee Warbler Oreothlypis peregrina C F C F R U U F C Louisiana Waterthrush Parkesia motacilla R R R U Prothonotary Warbler Protonotaria citrea R R R Yellow-rumped Warbler Setophaga coronata R R R Yellow-throated Warbler Setophaga dominica O O O Blackburnian Warbler Setophaga fusca U U F U R R U F U U Magnolia Warbler Setophaga magnolia A A Chestnut-sided Warbler Setophaga pensylvanica R R R R U R Tropical Parula Setophaga pitiayumi U U U U U U F U F F F U Townsend's Warbler Setophaga townsendi U R U R R U F F U Black-throated Green Warbler Setophaga virens F U F U U F C C Golden-winged Warbler Vermivora chrysoptera U U F U U F C Wrenthrush (e) Zeledonia coronata U R Tanagers Peg-billed (e) Acanthidops bairdi R R R U Green Honeycreeper Chlorophanes spiza R R R Bananaquit Coereba flaveola U R R R R R U R R U Red-legged Honeycreeper Cyanerpes cyaneus O O O O O O Shining Honeycreeper Cyanerpes lucidus A A A Blue Dacnis Dacnis cayana O O O O O Scarlet-thighed Dacnis Dacnis venusta R U U F U U U U F F U R Slaty Flowerpiercer (e) Diglossa plumbea R R R U R U U R U Tanagers (continued) J F M A M J J A S O N D Gray-headed Tanager Eucometis penicillata A A Cherrie's Tanager (e) Ramphocelus costaricensis U U U U U U U R U U U U Buff-throated Saltator Saltator maximus U U U U U F U R U U U U Streaked Saltator Saltator striatipectus U Variable Seedeater Sporophila corvina R R R R R U Thick-billed Seed-Finch Sporophila funerea A A Yellow-bellied Seedeater Sporophila nigricollis A White-collared Seedeater Sporophila torqueola A A White-shouldered Tanager Tachyphonus luctuosus O O Spangle-cheeked Tanager (e) Tangara dowii R R R R R R U R U U U U Speckled Tanager Tangara guttata U U U F U U U U F U U F Bay-headed Tanager Tangara gyrola U R U U U U U U U U U U Silver-throated Tanager Tangara icterocephala C F C V C V V F C C C V Golden-hooded Tanager Tangara larvata U R R U U U U R R R R U Blue-gray Tanager Thraupis episcopus U R U U U U R R U U Palm Tanager Thraupis palmarum U U R U U R R R U Yellow-faced Grassquit Tiaris olivacea U U U U U U U R R U U U Blue-black Grassquit Volatinia jacarina O O O O New World Sparrows Chestnut-capped Brush-finch Arremon brunneinucha F U U F U U C U C F F F Costa Rican Brush-finch (e) Arremon costaricensis O O Sooty-faced Finch (e) Arremon crassirostris A Black-striped Sparrow Arremonops conirostris O O O White-naped Brushfinch Atlapetes albinucha U U R U U U F U U U U U Common Chlorospingus Chlorospingus flavopectus V V V V V V V V V V V V Sooty-capped Chlorospingus (e) Chlorospingus pileatus U R R R R R Large-footed Finch (e) Pezopetes capitalis A Yellow-thighed Finch (e) Pselliophorus tibialis F F F F U F F U C F F F Rufous-collared Sparrow Zonotrichia capensis U U U U U U U U U U U U Old World Sparrows J F M A M J J A S O N D House Sparrow Passer domesticus A Cardinals Blue Seedeater Amaurospiza concolor R U R R R

Blue-black Grosbeak Cyanocompsa cyanoides A Indigo Bunting Passerina cyanea A

Rose-breasted Grosbeak Pheucticus ludovicianus R R R U R Black-thighed Grosbeak (e) Pheucticus tibialis U U R U R

Flame-colored Tanager Piranga bidentata U F F F U F F U C F U U Hepatic Tanager Piranga flava R U White-winged Tanager Piranga leucoptera U R U U F U U U U U U U

Scarlet Tanager Piranga olivacea O O O Summer Tanager Pirangra rubra U R U U R U F F Blackbirds & Allies Yellow-billed Cacique Amblycercus holosericeus O O Melodious Blackbird Dives dives R R R R

Baltimore Oriole Icterus galbula U R U R Bronzed Cowbird Molothrus aeneus R U U R

Chestnut-headed Oropendola Psarocolius wagleri R R Golden-browed Chlorophonia (e) Chlorophonia callophrys U R U U U U U U U U U F Elegant Euphonia elegantissima R U F F F C F U U U U

Yellow-throated Euphonia Euphonia hirundinacea R R R R Spot-crowned Euphonia (e) Euphonia imitans R R R R R U U

Yellow-crowned Euphonia (e) Euphonia luteicapilla R R R R R Lesser Goldfinch Spinus psaltria R R R R R U

Yellow-bellied Siskin Spinus xanthogastrus R R U