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uoslriAc InEd

(asaJBlng) Brauaf palelar pue unllDqaqd ul saSuBqJlBJnlBlcuatuou puu sa!)adsi{aN

(8661)882-192:(Z)Z I,rsi'(rrtl Nr)rrld Vol. 12. No.2 (1998)

Thcscsuggested relationships, which werelargely bascd on floral characters,are now strongly supportedby seedmorphology which is distinctand uniformwithin eachof the abovegroups.

The traditionalclassilicaLion obviously rcquires attention. The options that are available in order to provider rnoren,rturrl r lrrsrilitrtirrn rrr':

L to unite wcler Phebu[iunrall the closelyrelated genera that are indicated above, or

2. to recognize that P he balium cLtnsists of thcspccies in sect. , with or withoutthe inclusion of M icrocl be, and to transferthosc spcciesin sect.GoniocLqdos to Rhadinothamnus,ard, thosc in scct.Eriostemoides to Nemcuolepis,while sect. would be given generic status.

It hasbeen decided to takcthc sccondcourse, in particularbecause each of thc resultantgenera is honrogeneousand clearly distinct morphologicalJy.Microcybe, which is closclytelatedto Phebalium but diffcrs in havingsessiJe flowers and 2 calpellaryovaries, is retainedas a distinctgenus.

"Flora Thispaper is aprccursor to anaccount ofthe gencra that will bepublished in Volume26 ofthe of Australia",theretbre descriptions are only providedfor newlyrecognized or newly circumscribed taxa.

Hilar strandsofthe seed

The ovariesand dcvelopingcarpels in dried herbariummaterial of a numberof generain the Rutaceaetribc Boronicacwcrc studiedin orderto establishthc homology ofcertain charactersevident on the sged.

In somegenera of the Boronieaethe seedpossesses a cream-colourcd ligament-like tissue on its adaxialsurfacc betwcen thc micropylc and thc chalazalopening. This tissuehas thc appearancewhen dly of a cartilaginousstrand; it surroundsthc hilum andextends, as a singlethread, to theraphe. This 'hilar cartilaginousmaterial, which is helecalled the strand',is shedon, or shortlyafter, the dispcrsal of the seed.

Hilar strandsof identicalform andorigin are found in seedof the following taxa: l. Chorilaena(Figurc l) 2. Phebalium sect. Canioclados 3. Rhadino t hannus eup he ntia e 4. Asteralasiap.p.

From ii study ol stagcsin the developmentof ovulesthrough to seedin the abovetaxa, and comparingthcse stllgos with similarstages in seedsof othermembers oI thc Boronieae,it bccame apparentthat the hiiar slrirndarises frorn thc narrow portion of the outcr testa(formed from the outer integument)that sullouncls thc hilum. In earlystages oi:seed development the futurc strand is intimately tlsed to thc restol thc oulerlcsta bLrt when nearly maturc it becomesseparated and eventually falls awayfrom the seed. Y

'x{1cl ':^r.lJu.rrrlJ lccrreqdsrureqeql 3 . r.qlo ,{qpauoddns sr r.rourS rsrqt urrm toclclrqsuorl-'1e: esolr n c)utssnuuDqloulpDqA vpweDuaollloLlS ur ,{lluopuodrpur rsolr spLm.rlsir,'lnl.Llt tuql {lilrlun srtrl

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694 ':3ueqr lc:ntrtruruioupuc'rrrJ" $ru rr,4,4J,r,/ 'ru tr\\ C tncd N&]rri.rVol. 12,No. 2 (1998) thc valvatcp,l1als, and thc non-glandularapiculum to the anthers.A relationshipwith A.r/erol4sl4is moreremotc sincc in thatgenus the calyx is cxtrcmclysmall, the are imbricate, and the anlhcrs havc a slandularaoiculunr.

I(cy to gcnera of the Phebalium group

I Anthcrsbasifixed, apcx with a prorninenlsphcrical gland; bractcolcs basalto pcdiccland insignilicant 2 Flowerspediccliatc; scpals unitedi carpels 5 ...... Phebalium 2: Flowersscssile; 1r-ee or unitcd;carpcls 2...... Microcybe l: Anthersversatilc, without an apicrl gland;bractcolcs modial or supra-medial on pcdicel 3 Sepalsunitcd; anthelsobtuseJy apiculaLc 4 Plantslepidote; llowers solitary or cyrnose. Rhadinothamnus 4: Plantswith stellatchairs; inllorescence a 6 tloweredumbel ...... Chorilaena 3.' Sepalsfrcc; anthersretuse rt apex 5 Plantslcpidote; sepals imbricatc; pctals imbricatc or unitcd...... 5: Plantglabrous or with simpleor stellatehairs; sepals valvate; pctals valvateor unitcd...... Leionema

Leionema

Leionema(F. Moell.) PaulG. Wilson.g (n. et comb.nov.

Eriostemo sect.LeionennF.Muell.,PL Vjctoria I : 125(1862).- Phebaliumsect. Leionemq (F. Muell.) Bcnth.,Fl. Austral.I : 337 ( 1863).Type. Leionemo biloDan (Lindl.) Paul G. Wilson.

Eriostemon secL.CfutriLuettoltsis F. Muell., op. cit. 13l. Type. Eriostenon phylicoides F. Muell. f= Leionemadiosmer,rtlr (A. Juss.)Paul G. WilsonI.

S/rrabs,glabrous or with simpleor stellatshairs. Branc,4es srnooth. leaves, alternate, simple, sessile or shortlypetiolate, glandular-punctate, smooth. Flowersterminal or axillary,cynose or solitary, pentarnerous.Pzdicels slender, rnedial)y bibractcolate. Sepals frce. PetqLsfrce or united,valvate, elliptic,usually glabrous; apcx inflcxed. Slamen.rl0; filamentsterete, glabrous: anthers versatilc, Ioculideeply scparatcd at base,apcx dccply retusc, terminal gland or apiculumabsent. Dl.sc usually presentand forming a shortgynophore. Carpels 5, glabrousor stellatchairy,apcx slcrile. Ovules2 per carpel. Slyle solitary.slcnder, glabrous, affixed to adaxiaimedial surface of carpels; minutelylobed. Scedsub- r'en iform, c. 3 mm long,adaxial margin t straight:outer testa thin, smooth; sclerotcstasmooth; hilum lincar to nanowly elliptic;raphe fleshy basalor sub basalwith a thin coriaccousto clustaceousglossy covcring; chalazal opening basal or sub basai,obscured by raphe; placentalendocarp thick, pcrsistcnt.(Figure 2)

Chromosomenumbcr. n=16 (Srnith-Whitc1954).

A gcnusof 22 specics, 2 I in thc casternStates of Australiaand one in New Zealand. () :osi :029t 'IAW 'lluolJrA'1ac:3 '1l;rn1q :olo11)ralpnW i' drungdtung.arl,{; (8981)t:l tttfil.r.1 gsnlnlnrras'g

'(OH. u,ir) 'uruErrrsDJ'purlst ' Lt6l uunD U srapurlC..rd,{J (SEi.ll)6 I t9 : l ursr.J.IJ .1)ooH LLtnltl)uuJld

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luoslrl\|D ^ou .1u.1o.r InDd( llenr{ C) sualqruEBrueuora-J

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:cqdpri :D,sa,rarno ro :.r,{do,:,'uu,,uin'q q '"1t"i:"trJili"i'.'l']"l'#",t,,;il]jdi:li:"'l]:ltrili"r; uoqras lllp?r purpnlrBuot - J:arrJrns lDrxrpn g i^\.r^ IDr.lrrt V Oll) ].nas u t t\.ttdt).td1ut\t)tDut)n1 aarfiU

s.turq3 Iprnnrtr!.rlroLrfrnr sirrids ^\.u rrrl,,4rry./ uoslrr\ D Intd Nryrid Vol. 12. No. 2 (1998) f)istribution. Ocoursin Victoria in Thc Grampians,Central Highlands, and west Gippsland;in Tasmaniaon the mainlandand on islandsin BassStrait.

Leionemacarruthersii (F. Mucll.) PaulG. Wilson,comb. nov.

Eriostemon carruther.rilF. Muell., Victori.in Nat.7: 46 (189O).- Phebqliumcqrruthersii (F. Muell.) Maiden & E. Betche,Census New SouthWales Pl. 116(1916). I1,perMoruya, New SouthWales, W. Bauerlen 564 (lcctoi MEL 4638; isolecro;MEL 4639)lde Wilson (1970).

Distributioil. Occursin thc BatemansBay to Begt districtof New SouthWales.

Leionema coxii (F. Muell.) Paul G. W jlson, comb.nov.

EriostemoncoxiiF. Muell., Aastralas. Chem. DruggistT .64 (Dec.1884). Phebaliumcoxii(F.Muell.) Maiden& E. Betche,Ccnsus New SouthWales Pl. 1l6 (1916).Type. Braidwood distdct, New South Wales,W BaeuerLen199 (holo: MEL 4649;i-so: NSW 70184).

Distribution. Occurschiefly in thc BudawangRange in southeastern New SouthWales.

Leionemadentatum (Sm.)Paul G. Wilson,camh. nov.

"Brought Phebaliumdentatunr Sn. /a Rccs,Cyclop.27:(1814). ?ype: by Gen.Grose from somepart of New Holland,and coIrlmuniclted to usby A.B. LambertEsq" (/rolo:LINN).

P. salicifoliun A. Juss.,Anr. Sci. Nal. ( Paris)4: 412 (1825). Type. Port Jackson,New South Wales, anon.\n.v. ).

Eriostemonumbellutus Tvcz., BulL.Soc. Ilnp. Nqturalistes Moscou22/2: l5 (1849). P. umbeLlatum (Ttrcz.)'lrrcz., op. cit.25/2: 160(l852). Type: 125nilcslc.200 kml from Sydney,New SouthWales, W. Stephenson(ftolo. KW photo seen).

Distribution. Ocoursin New SouthWalcs chielly nearthe coastfrom Illawarra north to Port Stephens, and also in thc GibraltarRangc.

Leionemadiosmeum (A. Juss.)Paul G .Wrlson, comb. nov.

Phebaliumdiosmeura A. Juss.,Ann. Scl. Nttt. ( Paris)4:472 (1825).- P.p hylic olde's Siebcr cx Spreng. nom.illeg.,Syst.Y eg.4 pt2: 1640(1827 ), supcrlluousname based on above.-Eriostemon phylicoides F. Muell. nonr.llleg., Fragm. l: 107(1859). 7]Pe. Port Jackson,New SouthWales, comm. J. Gay (lso: K).

Chorilaenq qngustifolla F. Muell., Trans. Philos. Soc. Victoria 1: l0 (1854) fas angustfolio]. Type: Argyle County [i.c. Goulburndistlict], New SouthWalcs, anon. 826 (hoLo:MEL 4680; iso: K, MEL 4812).

Distribution. Occursncar the south-easlcoast of New SouthWales. 'pcl.,\\ouqlnu'esour,{c'iturtu.ol puu eldrulsqtl,tr snoln:oqnd ,(lesreds'3uo1 urc Z J acuacsatol[u1 '3uo1 'csnlqo'elqLre 'epr,r 'cr1dr11c tuur g ol otoqcdp otur es€ql€ pe,\\orrcu urc Z'3uo1 tuc E o1 'pe11op-prcnllad'peurea 'snoeJEuaqJ 'sosrq {1e1euurd Burrupl:snorqulS rr,1ra7 ll]ol ut04 tuorn.?p sqF ol onp,ftp ueq^\ Jcln8uuleq^\cruos'snorqe13'fssolB'qtoorLrs seltlruD.rg qFrq ru 7 o1r7z-rr7g

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'do?,l 'uoslrlv|C .q quor lnpd ( llcnt{ C) ual{Jaq clsqnssngula Bruouorr.I

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'pozruSooolcrn sorladsqnso,u.1 sa1o1g

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'1gg1y 'selciN 'plerlrelueJ 1914 :o1oq) tEI uDng J qtnos ,roN rroN :2.1t (t9g l ) : 'lDj]snv Ott I IC'qluog('l|].el:.lNf) snuplawnllDqaqd - G981) 1g1:1 ur8rjg'l1cnyglroiJDla uolaa|sol)g

',1oa quor'uoslrld D Inud( llcnl l l) snqslasuauora.I

iuoslt se8ulqr lnrnt?l.uauou puu sarJads.nr.u xrrt,tgr?/./ D Inxd Nr)/ri.r Vol. 12,No. 2 (1998) fasciculatehairs; ped icels l-2 rnmlong; bractcolcs caducous. Flowers glabrous. Sepals 5, vcryshortly unitedat basc,flcshy, deltate, c. 0.8 rnrnIong. Prlah valvate,thick, firm, narrowlyoblong, c. 4.5 mm long, 1.3mm wide,slrongly kcelcd, white; apex thickcncd and slightly inflexed. S/amars glabrous; filamcntsslender, terete, shortly cxcceding pctals;anthcrs cordatc, c. l.2mm long,bluntly mucronulate. Gynophorcc. 0.5m lr high, dr:cply I 0-groovcd.Ovarl barrcl-shaped,c. I .3nm high,glabrous or with a lew minutehairs, tcrrrinal 1/3solid; style f ixedto nearbase ofcarpels, tcrctc, slightly shorter than pctals. l',"!it not segn.

SpecinterLsexumi.nerl. Known only Ilonr thc typc collcction.

Distribution. North eastQueensland.

Habitat. \f"/ittdsweptshrubland on t(Ir oi.mounlain.

'fhe EtynoLogy. epithetis de.ived liom thc Lalin cllipli.?.r andrcfers to theelliptical shapeofthe leavcs.

Noter. This speciesis only known lfom thelype collection. The flowersarc similarto thoscof other speciesol Leionent4qxcept for the antherswhich arebluntly muoronulate(not rctuse)and for the gynophorewhich is dccplygroovcd. Thcsc anomalous charactcrs suggest thatit is incorrectlyplaced in this genusllthough neitherT. Hartiey(CANB pers.comm.), nor I cansuggcst a morc appropriate onc. I considerthat it would bc inadvisablcli)r it to be describedas a new monotypicgenus whilc tiuiting rnatcrialis lacking.

Leionemaequestre (D.A. Cooke)Paul G. Wilson,co,,lrb. rov.

Phebaliuntequestre D.A. Cookc,J. Adclaide Bot. Gard.l0:241(1987). Z)pe. Kangaroo Island, South Australia,B.M. Overton435 (lso: PERTH).

Distributb . Endcmicto KangiuooIsland, South Australia.

Lcionemagracile (C.'f.White) PaulG. Wilson,comb. nov.

Phebqliunrgracile C.T. Whitc, l).).. Roy.Soc. Queensland50:69 (1939). Type: Mt Greville, Quccnsland,C.T. Whire9947 (ftolo.' BRI 0l 1387).

Distribution.Occurs in cxtr.cmcsouth-cast Queensland where it is apparentlyrestricted to thesummits of Mt Moon andMt Greville.

Leionemahillebrandii (J.l{.Willis) PaulG. Wilson,coab. nov.

Phebaliun.hilLebruari il J.H. Willis. Victorin Nat.'13:195 ( 1957).- Basedon Eriostemouhillebrandii var.hrevilltliusF.Mrcll. non illeg.,'lrun"-. Philos. Soc.Victoria l.l0 (1854).ryp?. MtLofty Ranges, South Austntlia, F. M ueUer(leuo: MEL 4590).lideJ.H. Wlllis Loc.cit.

DisLributiott.Occurs in thc ML Lolly l{angesof SouthAustralir. '(t :uts) gg osualoC'puEleez^\eN'?E,lrO :ad,(l talta?pg'uBuueloH (99 u) 'eE,^\O:puBluez,l\oN g1gl 'osuapJ 'rW'pu€lsl uraquouaqlJo ls€oclsBeeqluo toUPip adtJ (6981) I8I:Iur8€rC'lenl'C(\oolH)snpnuuowalso1]g-(EV8l)899'l:9'ld'uorl'looqwnpnuwllllDqaqd

'Aou qu.ro)'uosltll\ c lned ( )tooH) unpnu Gluauolarl

'Bruulljsel lsE6-tll.touJo sull?tunoul3ql ur stnaa(J uollnq1JlSlQ

'1)l :ul.s)Izt rruzrtt\pI a '€IuPurseJ'sul€lunotr'{ 3tpu1 8 e8Zuung u ure1se16;ad'{-;(2981) 6zl:l t?r.roltr^1o3 14 "ltent{ 't''looqwnuDluotutunuDqaqd C( {ooH)sruD$owuowalsoY{- (tt9l)992:lQ looH)toq

'^ou [aoJ uosltM c lnudI looH ) lunusluoul stuJuolJ-l

'€IIEllsnV 'elnsuluad 'ErJolcr^ uJelse^\l?J ol lsee qlnos el^A uJoqlnosul srnJxo uounqlllsl1

'S1;utstta11ttt1,ttr'crlurr\nV (829t lEt\ J 'elEJ qtnos 'fBg ur.JJoC.Beupue ,{eg relunocug :adKJ (JJ,681)fZ leJlsnv S rlDrlxg H qpuuH "qtueg ( qlueg) snn(qdtqJoiq uowatsotrg (€98t) Ing :l lE.IlsnV IC un]l4dtLltatq utntlDqaqd

'rrqLualdas 'uesoqca:aq ed,{1o1Jol (829t -Ighl :oQal)tr anW J tZ 'vrl€jlsnv ',{Eg qlnos r€lunocua :adtt1 lng s,luecur^ 15 puu s,rerued5 Jo saBuEJlsuoc 'nAol ' ttowatsotrg er11u6:uot1o11tadKa (gg8l) 66 il DuoJrtA2oS'sollqd .s11""iJllenlN I.tull'{qdo)11u1

'^ou qtuo)'uoslt$\'C InBd( llenl I C) tun11'{qdo'rc1uruuauolal

'uuo]or^ u.IelseauI puE 'sal8l[ 'uo1|nql]$1O 'spJE.\\qlnoseuolsl^d uto4 qlnos ,^\aNulelsBe Jo e8ue11Sutpr,ttq etp ut sJtlxx()

'OASN ') 'CV -IglN 'BtJolcIA . os?: 18 L, : opq) D anry I'6981^lEnul?f 'Ie^ru relsll€cutr{eqluo surstunou.'ad(J (sqs t ) Olg :t lErlsnv lc' qlueg ('ll"rtrN'9,)wn\(LldotdwDl '.2 'l?ryN'g ut1!ft1qaqd (698I) gt oltoictA 2oS wlDlld t UD\A sn11{qdo wttl ltowalsoug

'^ou qwo)'uosltN\'C lnEd ( llonrI C) utnfidqdoJdusl Burauola'I

'seu,(c ,{JsllIx€eql uI ro,^^oueuo fluo Jo eruaserdoql ,{q ,(lludtcuud uJelltuo4 peqstn8ullslpequeJlllnq l.utln7IIJU{qdTJosluqtlE^eurosol.lDlrurrsslsatJJdsslqJ raloN

'soplA qlnos ,{{eN'sut€lunohtr enlg aql ut sdorclnof1co.r uo Suuncco sirJads Jrrr V uottnqutstQ

'(IAIN'IUg .rv :)I .o1o4) 'ZZg 'sapiA 'sulPlunol 'qtEeqlcElg sol"A\ gg woqsutuunJ y I qtnos .^\eN trenlg a.fiJ (€68l ) [t 'wou 'J snrloltctltqd WnoS^\eN IC qpuu1' \alf atool xa llent[I ( uurl) Y) saploaDlttlcrtltu,. uowa$ou{ (gZ8l) Z0g selE^drllnos ^\eN luehtr SoeC'plelCU?uunJ V saPloaDltllrDlwnuDqalld

'^ou '\toslr$\ 'C qwo) lncd ( uunJ V) saploaBuqJBlBluauola'I

s€3u"q. luntvl.uetllou puv sorrads ^\eu 1rlrtr4t4d uoslnA C Ined Nr)rbiuvol. 12,No. 2 (1998)

Distribution. North Island,Ncw Zealancl.

N.,te. This is the only menbcr of the genusthat is foundoutside of Australia.

Leionemaobtusitbliunr (Paul G. Wilson)Paul G. Wilson,conb. nor' phebaliumobtusifo 1l/.rru Paul c. wilson , Nuyt.sial.107 (1970).Type: Upper reaches of Alice creek, (lzolo: about 8 miles fc. l3 kml north of Helidon,Queensland, August 1963,F D Hockings BRI 042851).

Distributktn. Occulsin the Helidonand Ravensbourne areas of south-castQueensland'

Leionemaoldfieldii (F. Muell.)Paul C. Wilson,c.),?1, nov

Eric)stemonottlliektii F. Muell., Frirgm.l : 3 ( 1858). PhebaliumotdJiekltl (F. Muell.) F. Muell. ex Benth.,Fl. Austral. l: 340 (1863). T)'/rE.Mount La Perousc,2TFebruary 1857, A Oldfield & C. Stuart 18'15(hoLo:MEL 4822;iso: K).

Distribution. Endcmicttl mountainsnear the westcoast of Tasmania

Leionemaphylicifoliurn (F. Muell.)Paul G. Wilson,corab nov.

- phebaliumphyliciJbl,.rnr F. Muell., Trans.& Proc. victo an Inst.Advancem. sci. l: 32 ( | 855). EriostemonphyticiJblius (F. Mucll ) F Mucll., Fragm l: 105(1859) Zype MunyangMountains 1855,F MuelLer(Lecto: fsnowy Mountainsl,also on the SnowyRiver' New SouthWales, January MEL ,1888;iso lecto: K) .fidePtul G Wilson(1970).

Distribution.Occurs in thc mountainsol'castern Victoda and of theextlcne south-eastofNew South Wales.

Noles. Seg lrotesLttldcr I'. lLrchttaeaides.

Leionemaralstonii (F. Muell.)Paul C. Wilson,conb. nov.

Eriostemont.al\to ii F. Mucll.,Fr.agrr. 2: l0 L t. l4 ( 1860).- Phebaliumralstonli (F. Muell.)Benth., Fl. Austrrl.i:339 (1863). f1?e.TwofbldBay, New SouthWales, FMueller (hoLo:MEL 4915; lso. K, MEL 4946).

Distribution. Occursin 1hcBega to EdcndislricL of Ncw SouthWalcs'

Leionemarotundifolium (Endl ) PaulG. Wilson,comD. nov'

Eriostemonfotu tliJollu.rEndl. l Endl.er cl., Enurn. Pl. Hucgcl l5(1837). Phebaliumroturulifolium (Endi.)Benth., Irl. Austral.I : 341 ( liJ(r3).7)Pe Mount Dangar,Hunters River, New SouthWales, A. Cunninghun55 (lso: K, MEL'1954). 'DIitrlSllV OlilUlOpuC SOla)ilds ua^i's_ lO SnuCilV

'(266 'i96 l .SuoJlsulvT .Jr?tb^ I rtrq1\-qlr LLr S) t)1=u .!dqu!u aluosout(uq)

(t.)Jn3rg)'snonPltip'snouutqutiulcllrJQpLtJ ltttuci)uld :lu.utl ll,lll :cceJJcrxupe JaA\oJ tro c;nlrcdu IEzDIf,qJ:usct riino qtr/\\s-nonurtuoa sr lullt ic'{rl snoocr:l,tocutqt {q 'eJnuede 'snout;tlrur:.r pale,\oc luzcluqopuu ulnllqJo estq ue3A\loqpclrntrs lIq^ iutns 1p-'utseqelur 'lErJrJJadns 'qloolus'snoc.uuor 'utql :Jltdrlla,{l,roJJpu unlrq:qloous nlsolorelos:el rI u|lus P}sc1l31no 'ttuoJIu.r-qns^ lqFIp:1Sur8Je ru lEIrcpe'3uo1ruu E 7 Z ot prosdrlla,{lptrorq ParS potpl ,{1ac,ttrsrr'a345 'sledrec;o 'aloJol',{rrlrlos ecegnslurpetu lclxupu ol pexrIIE ,1,!,S lod.rurlocl I saiir,r5rxodu e1t:e1s 'g 'asnlor 'osDq troqs B qtr^\ sladra3 lucurruordrsr.J rElnpuelStou ,(ltqBrlsxidu tt poleJrdis,{ldc.p IIncol 'oluesre,\sreqlue :,ftIeq etEIlots,{llps?q ro s^norqrlS'.torct peueltellsluotuxlll :ce4 '0l sr2rrrJs 'sno:qe13 .ctnirrqull'oa]I ro eloprdel'(sapvlpqaqdNu patrun)otecuqtur slrlrd slrrlrs .l.tldece.r 'olu lerog ,(qsegeql Surpuetqns,{leletperuut puu lucrdc.ro lartpadlo olpptu iu.Lr sap)nDtg 'snorauIEluad'^JDlrlos ro asoul^c'IlEllxe.r.ra,i{o/J qloous 'oln1rund-ru1npuu13'etelonod,{proqs '1rg 'eldut 'eluu + ts )e\1esalDa1 esoJrurc^ro tltooursspltlJuD.!g lra.// llcur\ ro tqrt.rtTs.rttrprdrl

'I zaJnl saplollDqeLldstdalontual'1 -] ^fiH sapiaaD.tto) S '(Eg8l) raftJ Vg:L'JSIW'ptDD aax pB'[ s,t>looH'^-n]FI x. tulunj(l 1unp4ado[t1dru'{g

'[uosltlN D Incd( l|t|',1l oawnnltsstda]orrutaN -l 'l]rqt-l 'lpuT 'tx.s snaaDnbsuowatsotl4 rad,,{J (0i81) 991I Id ucC saptoLuatsol.tll utnlLDqaqd

'7J.rnJ ' saptolloqaqd'19:adrt1 (2981) 8EI :(Z)9Ztlusow satstlD.rnDNAUJ ios lltltl zatrJ- s-ldaf)tDwaN

sldaloluureg

'sele^\ qtnos ^\?N's{jed lpuorltNeAUEU elSunqurnrfiiN pLtt? rrlndD) 1l^lut s]lxa? uollnqulslO 'Gv6r '8t61 'so|rlA 'eiuu11 fg},{ .oI? ://29 IASN:oloq) alE)isuoCJ? ,{Dt{8Z Wno5.uop el6unclurnrrEd\ 'urulunol '6L61) ,&a8noleg:adfi LII tI Dtst( N'uostr7'yC lt\vd uttuoL[1P1.!1^unltDqaqd

'^ou 'D qwo)'uoslr$\ lntd (uoslr/d D lnud)wn.rourpula Euauola'I

'solrld qlnos ,lreN'euotsl^)J lrlou sofluur.ql ur slncJo rlollt1qlJist1

'GL0LI7\O 'ASN ' HJdAd ../.ti:Zohtn96 eV :oloLl)uasu40f SV'l I E6l rcqur.ldesZ 'salEi[ 'ppullo.IE3N. qtnos,$aN ad(J (0f61)9lI:1ors1,{z1g'uos1r115 ltlt:dutlUD]a(lw,{s utll1,qaqd

'dot 'uoslld|C qtlor lncd (uostr1\\D 1nu4)wnluladru,{s etuauolal

'puulsuaon0 lo isue-qlnosetuoJtyc oql ur pup selu^\ qtnos ,\\eNtsEa quou Jo stculsrpuo13uuro1 pur l1c,trog6tU ul sincf,O 1t01lfiq1Jis1Q

'uost'r\A LL' sa8uBqr lxinllt.uaurou pur s:rr.ids ^\:rl r/r4rtrit./ C InDd N!)',ri.r Vol. 12, No. 2 (1998)

Fig\trc 3. Netwk epis.v/!dric.r socd (x25). A lalcral view; ts - adrxial surlacc; C transverseseclion with enlargcd section through testn; c - chnlazal apedure: h hilurD; In rnicropylc; ot outef tesia; r raphc; s sclerotesta. Drawn trcrr, F.A. RadhLt), 1179 (NSW)

Nematolepiselliptica (PaulG. Wilson)Paul G. Wilson,corzD. rror,.

Phebaliumel[ipticun Paul G. Wilson, l .341 (1914).Type: BigBadja Mountain, New South Wales,lP. Baker9Ol (hoto:NSW; lso: CANB, K, MEL, PERTH 01616587).

Distribution. Occursin thc rtrngcscirst ol Coorna,New SouthWalcs.

Nematolepisltondosa (N.C.Walsh & Alb.) PaulG. Wilson,comb. noy.

Phebalium frondoslrir N.G. Walsh& Alb.,Muetleria 6: 405(1988). Ilp". EastcrnVictoria [precise localitywithheldl, D.E. Albrecht 2815 (holo: MEL).

Distribution. Known only fiorr thc uppcrslopcs of a mountainin eastcrnVictoria.

Nematolepisovatifolia (F. Muell.)Paul G. Wilson,conb. nor,.

Pllebalium owttifollnn F. Muell., Tru1s. PhiLos.Soc. Victoria I : 99 ( I 855). Eriostemonoyatifolius 'Iype: (F.Muell.) F. Muell.,Fragm. I : 103( 1859). ln thcalpine parts of theMunyang Mountains, New SouthWales, F. Mueller (Lecto.MEL 4828) lide Wilson ( J970).

Distribution. OccuIsin the SnowyMountains of New SouthWalqs.

Nematolepisphebalioides Tulcz., Ball. Soc.lmp. Nqturqlistes Moscou25(2):158 ( 1 852). Ty,pe. Swan Rivcr Colony,Wcstcm Australia, -/. Drwnrnortd 5Lh coll. n. 194(holo. KW,photo scen;iJo.. K, TCD).

Slmphlopetaloncorraeoitles J. Dfumm. ex Harv.,Hooker's J. Bot. Ke$)Gard. Misc.7: 54 (1855). 71,pe:Near Middlc Mt Barren, J. I)rummond194 (holo;'lCD iso; K). -utuutuslr.lu,tilsli qljou ut s.tna.(, uollt1quls1Q

'(uc.solotld) :uts1atuatta1 Ai l! ,pnoe I'J[8] r?rut?tusr].1,:ati,(,L (OL6I) .c '(sggl)[g.l.urscJ t6:l rlsl(rN'uoslrl lnEd( looH) arsnlaL dsg|rsu natunttbs-1u114 Dtp1ld lJ 'JlooH(I@H)1u,rsnpl ./ )E^1ratptDllt q (nfgl)tEZtlQ):looH) pg t'lo<)Hru11s-ntatu!ntlDqaU(J

'dol./ ./lr.)J'uoslr/{ D lnEd( looH) ssnlor dsqn"-ecurunbs sldalolBureN.J

3uc:cu:1n31n.,\6:cou Lucl4pur? Je^e{.r€lsrlle.r?trJeql Jo pcoqeqt :u.u sur?]unoureql u t u uotf,rA u;Jlsuaulo.r1 u,{\o u1K1u11 uotgqulstlT

'() .os1 :!t8t -lgw:oloq)Ja IJ98l 'iruolct^ 'uot8ur anW { qclut { lle,rd1tr41 puco3u|1 rriurprrg urc,\\l:rg. ad,{y,

'(.0L6i 'uoslud n6: I ,?slf,/N D lntdLunarDitar dsqnsunaurnnbs wnt1nc1ar14

'^ou 'C qwoJ'uosltN\ lnEd(uoslrld C 1nc6)earurro dsqnsBarranbs srdalolEuraN.q

'sIuEursEJ 'euolcrA'selnl& 'purls^uc.n0 pue qlnoS,raNlElslloJ ulots?c-qtnosur s.rnxao uoltnqulste

'(t 'poo16 .opq) wDLt8uturrJ y'i:^r|rldqlno5 Aeg 8ur:dg1o ,{1rurcr,teql ur .ad,{; (gzgl) Itf salulAqlnog,nat\J uro61 SocD'plorC gi/? uunJrunpp LunllDqaqd

earuenbsdsqns uoslrl\|D InDd( llrcll,r.l)pculenbs srdalolsuaN'u

'pcznrSolcr oru sencclsqnseerql s;r1o,y

'Drlu4snVu,relsce ur puoldsoprlA uott tll1ts!e

'(9209 'EZoS-lgt{.osrl,) ',.uauorc-uen eruErusEJ alrdecur 1u1rqrH,,ra./,{/ (96i1I)I :j7Ill'Lllr+uezurud 'tEN ' 'tDN ''3a11t l8ug ( rqu-l)wnawDnb"^ unlpqaqd (gzgl) tf I :Z t^r.tDJ tst11:tog waly tuou 'ssr| 'y .u .do .soo11 , u alpJDlltqwnuDqald ,{luo euriruol sr (t l g I ) f :/Z 1r,{3 ut .'all j luou tl.ttLLts - 'lll '1yqr.'1 wnalua'tn wnttDq)qd (9081) Itl r :l dS td IloH ^oN snautnnbsuouatsotrg

dou gIl.r,'uoslld|D lnDd( IIlqr-l) ecuuubs s1dc1olvua5;

'sclrl/i\qlnos ,rcN tsce-qtnosJEJ ul surotunoWlcou alrqi[ puu ?o/i\ 3o/A uoc,{1eqnr"'etu1d oqt uo sI]trO u01tnqlrli.l(I

'GtOLIgI HJUad ;o.r1:61.2€99t tEW ;oloq) tttz tqra.tqlVI e'sote^\ qlnos 'qslE/ 'qlv ^\eN'tunot{ 30163o/ irrl(J (8861)Z0r:gDualp W C N T &/r1ri4tlo1t4{tlL utntloqaqrl

'uoslrl[ ^o?/411or .l Incd(qsluA|D N ry q1y)u11,{qdoprr,tq: srdc1o1eura61

',{ug 'ltounqutstO al!loD.rsIol tsee3un,(elqunq uro:1 erprlsnV uratsal!\ lo tstor qtnoscllt slt.ix,\totstrtl?O

se8utq. Itrntctiucurou pur s.n.ds ^\in ' trntt)q)ttJ hostri\\ D Inud 280 Nryv.r Vol.12, Nu. 2 (19981

Notes. This subspeciesgfadcs into thc variantol the typicalsubspecies that is found in north-cast Tasmania.It is cvidcntlyclosely related to thesubsp. corio.ceafromVicloria. Superficially it is similar to N. oyatifoLiabut it diflers in havinga glabrous(not lepidote)ovary and in not havingsub-floral bractcolcs.

Nematolepisrvilsonii (N.G. Wrlsh & Alb.) PaulG. Wilson,comb. nov.

Phebaliun n llsonll N.G.Walslt& Alb.,Mue[LericL 6: 399 (1988). T)pe. Ncar Mt Grant,Victoria, N.G. Wq.Lsh1494 (holo. MEL 1540265;iso: PERTH 009055069).

Distributiotl. Known only from the type localityin the CentralHighlands of Victoria.

Phebalium

PhebaliurnYent.,Jrld.Malr. 2: 102(I 805). EriostemonsecLPhebalium(Yent.)F.Muell., Pl. Victoria l: 129(f862). Phebaliumscct.EuphebaliunBenth. nom. lnval. , Fl. Austral.l:337 (1863).-Crowea sect.Plrebqlium (Vcnt.) Baillon, Dict. Bot. I l: 217 (1886). Type:P. squamulosumVent.

Shrubst coveredwhen young with a lepidoteindumentum. Branches often glandular verrucose. Le4re,ralternate, sirnplc, scssilcor shortly pctiolatc, glandular-punctate,often glandular-verrucose. Flowers tcrminal to branches,soJitary or umbellate,pentamerous. Bracteoles basal to pedicels and insignificant.Calyr 5-1obcd,lcpidole ou|side. Pel.rl.r lree, imbricatc, cliiptic, white, yellow, orpink, lcpidotcoutside, apex not inflexed. Stamensl0,lilaments slenderterete; anthers basifixed, Ioculi totallyunitcd, rpex roundcclwith a sphericalterminal gland. Di-rcnot apparent.C4lpels 5, Iepidote, apicalportions solid. Otules2 percarpel.Sryle terete, affixed to adaxia]medial surface ofcarpels; stigmasmall wilh shortlyspreading lobes. Seel oblongrenifbrm; surface longitudinally striate or coffugatedue to thefine corrugalionsof thcsclcrotcstai outcr testa membranous, black; hilum linear; raphesmall, fleshy and shrivellcd, situated in lowcrhalf of adaxialface, covered by membranouslayer thatis continous with oLltertesta; chalazal aperture on loweradaxial face; aril linearl placental endocarp thin, caducous.(Figure.l)

Chromosomenumbcr n= 16,32 (SrnithWhite i954).

A gcnusof 25 spcciesendenic to Australia.

Nores. This ,as circumscribcclin this pnpcr,is closclyrelated to MicrocybeTurcz. which is distinguishedby its sessilellowcrs, srnall lrcc or unitcdsepals, and bicarpellary ovary. The seedsof the two generaare similar.

Phebaliumappressum Paul G. Wilson,rp. ror,.

Ramulisparse glanduloso tuberculati. Folia dcnsa, scssilcs, crecta, ad ramulumadpressa, cordato ovata,c. 2 mm longa,L5 nlm lata,crassa, arote revoluta, supra aliquantum applanata, infra rotundata, laeves,virides, glabrr vcl sparseargenteo lepidota. Flores terminales, solitarii vel binati;pediccllus brcvis,crassus, c. J mm longus. Calyx c. 1.5mm altus,profunde deltato lobatus, cxtra ferrugineo lcpidolus. '(28E69 ' -lEI^t..rri ,t\SN I I8t i09l0S!96 (lV:oloq) ! '6E sulaH y J681 r?quetdes ZZ'EIlErtsnV uJ3tsclA'3uudS ouotrrn ueonf:r:ou dtlrtr'llcsicl !rrol.rA 'oL61) 'uoslrld patg :adt1 ZL: I ,?r'tLlN 5 gu4 w41[4d{t1t ]./ dsqnsLunsoln)taqnl wn.\Dqaqd

'^ou tDtsla 1/./or./'LroslllN!) lntd urnlloJln.a.rqurnllBqaqd

'slccrpad lcrql puEtroqs ,{Ja,\ a^Eq qclq,{\ sJ?^\oll ,{:r:1r1os {11unsn cllt ur puuedllqs Jucl uI xelcJtuocwn11,(t1dotJlw uuiltDqaq(l a1l Jo srcqrueorue,tale reqlo oq1ruo..r.l s-t.11tp setlecls sIqJ srloN

'uorlrsod 1uo1oqt ot s:elo.rlotlltclo otJrcrds aq1 {:JoPut,gg

'euO,{luor.rd

:r?rolCucrlursnV uretsed\rol epoJ uorle^rasuoJI/{'1VJ uoilrclo.tdj o paeuu r aq ,{eu i-. t..ds oqlltql stseSSnsqcrqlr asrclSururur r o11rcfqns ccru uD uro4 sruorlJelloc {rutrlos lq1 stllztsuo]tDLlaslta)

'Sururur.te1r lros1?Jol qtr,lr pellr1el uoaq peq leqt f,crr uu ur 3ur.uo,r11urulcl puus ^\ollo1 JrJr.lrll

()il6l p.rpag)lJUtsrC lEcruutog crp:u31oo3 eql ur sr qolq,{\{trltJol cd,{1oq1 uro,rl {1tto nt ou:] uotlnqu$!O

'ueas lou llr.rr' aprslnoctoprdcl-"^nourSn:re1'pccyll-alullap {Jdoop '8uol 'J 'atoprdel ur[r g'l x{p) 3uo1uur 91 J seloalJErqelteunJ ^l^\orrru p::a,les,{c1pol]uilqns 'IJrql 'leutut.tct -snour8nxej^lesuap'8uol iuru I J puu 1.roqslectped :pcJlrd ro ,{.tt1t1os s-ra,roJjt '.tn[o^a,l 'eloprdel-,{re^Irs,{lesJEds ro snorqvlS'uccrB 'qloours '^\oleq pcpunor '.^oqD ltlJ luql\ouos '{cr 'epl|^ '3uol ^lesolc ql tuuIg l ttlruZ J'tlp^o-oteproJ'qJuuiqot pcsscjdduPuu tcola'osticP'.llssas sa^Da? alElncJaqnt-.rulnpu1?l8,{1as:uds'e1oprdel,{re,t1r,'^slaltltunLg t13tt1uti o} qr./i/l popunou

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'uostrrt I8Z sa8uEqr lunllltrururou pue s.rr.ds l\ru UrrD./,r/(/ D inrd 282 N!)Iv,r Vol. 12. No 2 (lqq8)

Noles.This spccies is only Iounclin anarea around Queen Victoria Spring in theGreat Victoda Desert of Wcstcm Australia.It is helc isolatodliom otherspecies of PhebaLiumand showsno evidenceof intergradation.

Phebalium elegansPnul G. Wilson, .rp.acrv.

Ramuli lepidoti,gliurduloso tuberculati. Folia divaricata,atro-viridia, cuneata, retusa, crassa, c. 5mm longa,2 3 mm lata,glanduloso tubercuiata, marginis recurva, glanduloso undulata, supra glabra,infra argentcu vcl lcrrugincolepidota. lnflorescentia 2 5-noris; pcdicclli graciles, 5- 10mm longi. Calyx c. 1.5rnm altus. Pctalalalc clliplica,4-5 rnm longa,alba.

Typusr9 km WSW r)f Point Plcasant,Flaser llange, WesternAuslralia. 20 Scptcmber198,0, K. Newbey 7536 (hoLo:PERTH 00909726).

Spreadingshrub lo 90 crnbigh. Branchletsspreading, lepidote, glandular tuberculatc. Leaues sprcading,shortly petiolate, dark grccn;lamina cuneats, rctusc, thick, c. 5 mm long,2 3 mm widc, glandular-tuberculate(at leastwhen dry). Inarginsrccurved and glandular-undulate,upper suface glabrous,lower surfaccsilvei y- or ferruginous lepuJote.InJlorescence a terminalumbel oI2-5 flowers: pedicclsslender, 5- I 0 rnrnlong, lcpidotc. Calyxc. 1.5mm high, silvery-to ferruginous-lepidote outside,dividcd two-thirdsjnto deltatelobes. .sbroadly elliptic,4 5 mrn long, white,silvery to ferruginouslepidote outsidc. Fruitirtg cocci 5roadly oblong,c.3 mrr high,rounded at apexwith a small sprcadingapiculum on outerangle.

Distibution. SouthernWostcrn Ausfalia fronl Mt Day ( i20 km wcstofNorseman) east to theFraser Rangc,Coolgardie Botanioal District (Bcard 1980).

SeLectedspeciLnens exanLincd. WESTERN AUSTRALIA: 96 km E of Norseman,D.E. Albrecht 4032 (PERTH);9lt krr E of Norscman,/1.1 CronfieLd10065 (.PERTH);9 km Eof Norseman,C.A.Gardner 14222(PERTI{); Mt Drty,KR. NalDcy 5273(PERTH).

Habitat. In well drainedsrndy or gr.aniticloarn on rocky slopes.

Floweringperiod. July to September.

ConsenaLiotlstdtu.r. This spccicsis not rccorclcdlrom a rcscrvc,however, it is fbund over a wide area of pasturalland and vacantcrown lancllnd is not in ncedof protection.

Etymology.Thc specificepithet refers to the elegantappearance of the shrubas hasbccn notedby collectors.

Affinities.Tlr is species corrcspo nds most closcly to thepolymorphic P, ebaliumtuberculosum but it diffcrs in havinglong sienderpedicels, a cuncateleaf (not linear-terete),and a smallcalyx.

Nole,r. None of ths elevenspecies in what may be consideredthe Phebaliummicrophyllum - P. tubercuLo"^utttgroup can bc satislrctolilydiscriminated since each exhibits a differentvariant at cach differentlocality and cach appcars 1o hybridize rvith those of tllegroup it cortl(jsinto contact. A similar situationoccLrrs in P. clcgdir.rrvhich lt ils eastgrnextreme is silverylepidote and at itswcstgrn extreme fcnuginouslepidotc. Tho r,vostcrnvafianl may gradc1() thc southand cast rnto P. oboyatum. a

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'uoslr^\ €82 s.8urq) lunnf.uirrrou pur ser..ds ,rl)u ,ri4,r{rrl,1 D iDr.l 284 Nrlr.ri.r Vol. 12, No. 2 (1998)

a, Phebalium glandulosum subsp.nitidum Paul G. Wilson, subsp.nov.

PheboliogLandu[oso subsp. glartduLoso sinilis scdloliis grandioribusp)erumque 2 3crnlongis, 3-5 rnrn 1atis,apica truncatisprrum rctusis,supra convexis sgous costam leviter depressisnec canaliculatisdiff ert.

7j,pas.Warrumbungle Mountains, Ncw SouthWalcs, 25 May 1948,E. F. ConstabLe(holo:NSW 6462).

Branchlctssparsely glandular verrucose. Leaves shortly (2 3 mm) petiolatc;lamina narrowly oblongor narowly oblongelliptic, mostly 2-3 cm Jong,3-5 mm wide;margin crenate and glandular vefiucesc;base cuneate; apex truncateand slightiy retuse;upper surfaceconvcx with shallow depressionovcr midrib, glabrous,glossy, smooth or sparselyglandular-verrucose; lower surface smoothapart from thcsparscly verrucose prominent midrib, fawn lepidots. Pedicels slender, 5 7 mrr long. Calyx hernisphclical,glandular-vcrrucose, c. 2 mm highincluding thc broadly triangular lobes c. 0.7 mm long.

Specimenserantined. NEWSOUTHWALES:Head ofTooraweenah Creek, Wanumbungle Mountains, L.A.S. Johnson & E.F. Constqbte(NSW 20490); Mt Naman, 34 km SW of Coonabarabran, H. Streimann1 6 I (PERTH).

Distribution. Endemicto theWarrumbungle Range in north-easternNew SouthWales.

ITabitat. EvirJeniy conllncd to rocky basaltslopes.

Etymobgy. Thc cpithetis derivedfrom theLatin wordallidrls which meansshining and rclcrs to the uppersurface of thc leavcs.

N.rlen.This subspecieshas much largcr lcavcs than typical subsp. glandalosun but it is sirnilarto the large leavedv.rfiant ol lhat subspccicswhich gtowsin the GunderbookaRange south of Bourkein ccntralNcw SouthWalcs. Thc two subspccicsIn ay bc readilydistinguished by thc appcaranceof the uppersurihce of the leaves;in subsp.glarlrlllo.i&/lr there is a sharpdcpressed line over the midrib whereasin subsp.lltlrlurr thcfeis a gentledepression. In addition,the leavesof subsp.nltldrlm arc glossyabove rvhcn nalLn'c whcrcas in theGundelbooka Rangc variant the mature leaves are dull and oltcn retaina sparselepidote cover.

Phebaliumlaevigatunr Paul G. Wi)son,.rp.rov.

Larnina lolio angusteoblongl, I 2- I 5 rnmlonga, I .5-2.0tnm lata,obtusu, intcgru, supra co nvex a, glabrescens,nitida, Iacvis vcl lcvitercanaliculata, infl'a lepidota st manifcstecostata. Inflorescentia c.7 flota;pedicclliglac ilcs, c. ;l mrn longi. Caly x c. L5 mm altus,ad dimidiun in lobisdeltatis divisus. Pctalalatc clliptica.,15 rnrr iongir,llava vel alba,extra ferrugineo-lepidota.

'l'ypus; 48 knr ESli ,rl'Mclrcdir, WoslefnAustralia, N.N. Donner 46O0 (lolo. PERTH 896632).

Erccl slcnclcrrrnl/, to I rI] hi-qh.llrunchlets glanduhr tuberculate. Leaves asccnding; 2 mm long; lamina narrorvlyoblong, l2 l5 nlrn long, 1.5 2 mnl wide, obtuse,margin entirc; uppcr sorJedslJuusrp u sc uorlruioicJ s^lrn;.rrll,l\lt ]\ttll lse8Bnsslre,{ -sntuls fprql tsudeq1 re,lo stlpnls pletC slr qsllq€lsa,{lltoll ol tuolil.llnsulajo,{\ Lli.s suotloclloc,troJ .ql poqucseplsJrJueql& ellurlsnv ulalscl\ ]o leuJoclsrc-ql nos leJ cq1 uI punolsl uoxllls Iq.L IaloN

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Rhadinothamnus

RhadinothamnusPaulG.Wilson, Nalrsl a l: )97(197 l). Type:R. euphemiae (F. Muell.) Paul G. Wilson.

Pheballumsect. Gonioclodos Paul G Wilson,Niryr,rla l:96(1970). Type:Rhadinothamnus crnceps (DC.)Paul G. Wilson.

Silvery-lepidote slrubs. Br.lnchessmooth. aedre.raltsrnate, simple, shortly petiolate. Flowers cymosc or solitary and axillary. nedrally bibracteolate.Caly; patelliform or hemispherical, undulatclylobcd. PetaLsfree, elliptic, valvatc,slightly inflexedat tip, lcpidoteoutside, white. StaminalfiLanentsllatlcncd in lowcr part, tereteabove, glabrous; antheIs versatile, base cordate, apex obtuse,with a non-glandularapiculun, white. DlJc short. Carpels5, glabrousor lepidote,with or wilhout a short stclilc apcx. Oyrles 2 pcr carpel. S/yle solitary, terete,attached to adaxial medial surfacesof carpels;stigmi not or scarcalylobed. Seednanowly reniformor bluntlyellipsoid; outer testamembranous, dark brown, smooth;sclerotcsta smooth; hilum superficial,narrowly elliptic, borderedby cartilaginousstrands (hilar strands); raphe similar to hilar strandsin texture,sub-basal, coveredby mernbranouslayer that js continuouswith outertesta; aril linear,fleshy, situatcd between hilar strands,readily detached.

A genusof threespccies cndemic to WesternAustralia.

Not€s. The genus Rhadi otllumr?ir,iwas establishcdto acaommodatethe speciss Nemqtolepis euphemiae(syn. Phebttlium eup heniae) thatappeared to be anomalousin both ofthe generainto which it hadbeen placed. When the genuswas first dcscribed, comment was made on its closerelationship to tho specicsin Phebaliun scct. Gonioclados.but the speciesin this sectionwere not concurrently transf'erredto the new gcnus.This actionis now taken.

Someolthc charactcl'sthatscrvc to discrjmln te RhadinothtmnrAarefound in theseed,inparticular the rnannerin which cartilaginoidstrands surround the hilum. Thcsecharacters are described and discussedabove. Refer to Figure I , the seedof Chorilaena,which is thesame as th attorRha.dinothamnus.

Rhadinothamnusanceps (DC.) Paul C .Wilson,comb. nov.

PhebaliumancepsDC., Prodr. l:719 (1824). Eriostemonancep.s (DC.) Spreng.,Syst. Veg.2: 322 "Nouvellc ( 1825). Type. Hollande,cote orientale (Port du Roi Georgcs)"lKing GeorgeSound, Western Australial,/ide Jussieu,Mern. Soc. Hjst. Nat. Paris 2: 134(1825) (holo; G DC).

Distribution. (JccuISin the south-wcstol WcstcrnAustralia.

Rhadinothamnuseuphcmiae (F. Muell.) Paul G. Wilson, NuJtsiq 1. 198 (1971)., Nematolepis euphemiaeF.Muell.,Fragm. 3:149 t.25 (ApL.1863). Phebaliumeuphemiae(F. Muell.)C.A. Cardner, Enum.Pl. Austral.Occ. T0 (1931). TlTrc.Ncar Capc Arid, WesternAustrali a, G. MaxweLl(iso: K).

Pheb.Lliumbaxteri Benth., FI. Austrai. I : 345 (30 May 1863). NematoleplsDaxleri (Benth.) Engler in Englcr & Prantl, Nat. Pllanzenlam.III 4: 145( I 896). Tl,pe.South coast, Western Australia, W .Ba{te,' (holo: K).

Distribution.Occurs near the south coast of WesternAustfalia from thc Eyre Range east to Mt Ragged. 'uoslr116cuuurauuV {c1 po.ndcrtl c.to,tt suortrlrlslllli ctlJ

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References

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