Is Alnus Viridis 'A' Glacial Relict in the Black Forest?
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December 2012 Number 1
Calochortiana December 2012 Number 1 December 2012 Number 1 CONTENTS Proceedings of the Fifth South- western Rare and Endangered Plant Conference Calochortiana, a new publication of the Utah Native Plant Society . 3 The Fifth Southwestern Rare and En- dangered Plant Conference, Salt Lake City, Utah, March 2009 . 3 Abstracts of presentations and posters not submitted for the proceedings . 4 Southwestern cienegas: Rare habitats for endangered wetland plants. Robert Sivinski . 17 A new look at ranking plant rarity for conservation purposes, with an em- phasis on the flora of the American Southwest. John R. Spence . 25 The contribution of Cedar Breaks Na- tional Monument to the conservation of vascular plant diversity in Utah. Walter Fertig and Douglas N. Rey- nolds . 35 Studying the seed bank dynamics of rare plants. Susan Meyer . 46 East meets west: Rare desert Alliums in Arizona. John L. Anderson . 56 Calochortus nuttallii (Sego lily), Spatial patterns of endemic plant spe- state flower of Utah. By Kaye cies of the Colorado Plateau. Crystal Thorne. Krause . 63 Continued on page 2 Copyright 2012 Utah Native Plant Society. All Rights Reserved. Utah Native Plant Society Utah Native Plant Society, PO Box 520041, Salt Lake Copyright 2012 Utah Native Plant Society. All Rights City, Utah, 84152-0041. www.unps.org Reserved. Calochortiana is a publication of the Utah Native Plant Society, a 501(c)(3) not-for-profit organi- Editor: Walter Fertig ([email protected]), zation dedicated to conserving and promoting steward- Editorial Committee: Walter Fertig, Mindy Wheeler, ship of our native plants. Leila Shultz, and Susan Meyer CONTENTS, continued Biogeography of rare plants of the Ash Meadows National Wildlife Refuge, Nevada. -
( Su Manco (Astragal 5-Ye Ummary Os Milkve Lus
Mancos Milkvetch (Astragalus humillimus) 5-Year Review Summary and Evaluation Photo: Robert Sivinski U.S. Fish and Wildliffee Service New Mexico Ecological Services Office Albuquerque, New Mexico July 2011 5-YEAR REVIEW Mancos milkvetch/Astragalus humillimus 1.0 GENERAL INFORMATION 1.1 Reviewers Lead Regional Office: Southwest Regional Office, Region 2 Susan Jacobsen, Chief, Threatened and Endangered Species, 505-248-6641 Wendy Brown, Endangered Species Recovery Coordinator, 505-248-6664 Maggie Dwire, Recovery Biologist, 505-248-6663 Julie McIntyre, Recovery Biologist, 505-248-6507 Lead Field Office: New Mexico Ecological Services Field Office Eric Hein, Terrestrial Branch Chief, 505-761-4735 Thetis Gamberg, Fish and Wildlife Biologist, 505-599-6348 Laura Hudson, Vegetation Ecologist, 505-761-4762 1.2 Purpose of 5-Year Reviews: The U.S. Fish and Wildlife Service (Service or USFWS) is required by section 4(c)(2) of the Endangered Species Act (Act) to conduct a status review of each listed species at least once every 5 years. The purpose of a 5-year review is to evaluate whether or not the species’ status has changed since it was listed (or since the most recent 5-year review). Based on the 5-year review, we recommend whether the species should be removed from the list of endangered and threatened species, be changed in status from endangered to threatened, or be changed in status from threatened to endangered. Our original listing as endangered or threatened is based on the species’ status considering the five threat factors described in section 4(a)(1) of the Act. These same five factors are considered in any subsequent reclassification or delisting decisions. -
Reproductive Success and Soil Seed Bank Characteristics of <Em>Astragalus Ampullarioides</Em> and <Em>A. Holmg
Brigham Young University BYU ScholarsArchive Theses and Dissertations 2011-07-06 Reproductive Success and Soil Seed Bank Characteristics of Astragalus ampullarioides and A. holmgreniorum (Fabaceae): Two Rare Endemics of Southwestern Utah Allyson B. Searle Brigham Young University - Provo Follow this and additional works at: https://scholarsarchive.byu.edu/etd Part of the Animal Sciences Commons BYU ScholarsArchive Citation Searle, Allyson B., "Reproductive Success and Soil Seed Bank Characteristics of Astragalus ampullarioides and A. holmgreniorum (Fabaceae): Two Rare Endemics of Southwestern Utah" (2011). Theses and Dissertations. 3044. https://scholarsarchive.byu.edu/etd/3044 This Thesis is brought to you for free and open access by BYU ScholarsArchive. It has been accepted for inclusion in Theses and Dissertations by an authorized administrator of BYU ScholarsArchive. For more information, please contact [email protected], [email protected]. Reproductive Success and Soil Seed Bank Characteristics of Astragalus ampullarioides and A. holmgreniorum (Fabaceae), Two Rare Endemics of Southwestern Utah Allyson Searle A Thesis submitted to the faculty of Brigham Young University in partial fulfillment of the requirements for the degree of Master of Science Loreen Allphin, Chair Bruce Roundy Susan Meyer Renee Van Buren Department of Plant and Wildlife Sciences Brigham Young University August 2011 Copyright © 2011 Allyson Searle All Rights Reserved ABSTRACT Reproductive Success and Soil Seed Bank Characteristics of Astragalus ampullarioides and A. holmgreniorum (Fabaceae), Two Rare Endemics of Southwestern Utah Allyson Searle Department of Plant and Wildlife Sciences, BYU Master of Science Astragalus ampullarioides and A. holmgreniorum are two rare endemics of southwestern Utah. Over two consecutive field seasons (2009-2010) we examined pre- emergent reproductive success, based on F/F and S/O ratios, from populations of both Astragalus ampullarioides and A. -
Stehlik 2002
American Journal of Botany 89(12): 2007±2016. 2002. GLACIAL HISTORY OF THE ALPINE HERB RUMEX NIVALIS (POLYGONACEAE): A COMPARISON OF COMMON PHYLOGEOGRAPHIC METHODS WITH NESTED CLADE ANALYSIS1 IVANA STEHLIK2 Institute of Systematic Botany, University of Zurich, Zollikerstrasse 107, CH-8008 Zurich, Switzerland The glacial history of the alpine herb Rumex nivalis was investigated using ampli®ed fragment length polymorphisms (AFLPs) and restriction fragment length polymorphisms with polymerase chain reaction (PCR-RFLPs) of cpDNA. Both traditional statistical methods widely applied in phylogeographic research and nested clade analysis were used. The AFLPs indicated little geographic structure probably due to the wind-pollinated reproductive system of the dioecious R. nivalis. Because cpDNA haplotypes exhibited distinct distributional patterns, correlation between AFLPs and PCR-RFLPs was low. The results of common statistical methods and of nested clade analysis were largely congruent. Both supported in situ survival of one group of common haplotypes in the Central Alps. For another group of common haplotypes, classical phylogeographic analyses gave strong evidence for survival in peripheral refugia at the northern alpine border, whereas this conclusion was not as clearly supported in the nested clade analysis. Nested clade analysis provided several detailed insights on past and ongoing populational demographic processes. Thus, it is a valuable tool in the phylo- geographical analysis of haplotype data, but it should preferably be combined with other statistical analyses. In situations with low genetic variation in cpDNA, classical phylogeographic analytical tools on nuclear DNA will still be the methods of choice. Key words: AFLP; glacial survival; methodological comparison; migration; nested clade analysis; nunataks; PCR-RFLPs of cpDNA; Pleistocene glaciation. -
Astragalus Missouriensis Nutt. Var. Humistratus Isely (Missouri Milkvetch): a Technical Conservation Assessment
Astragalus missouriensis Nutt. var. humistratus Isely (Missouri milkvetch): A Technical Conservation Assessment Prepared for the USDA Forest Service, Rocky Mountain Region, Species Conservation Project July 13, 2006 Karin Decker Colorado Natural Heritage Program Colorado State University Fort Collins, CO Peer Review Administered by Society for Conservation Biology Decker, K. (2006, July 13). Astragalus missouriensis Nutt. var. humistratus Isely (Missouri milkvetch): a technical conservation assessment. [Online]. USDA Forest Service, Rocky Mountain Region. Available: http:// www.fs.fed.us/r2/projects/scp/assessments/astragalusmissouriensisvarhumistratus.pdf [date of access]. ACKNOWLEDGMENTS This work benefited greatly from the input of Colorado Natural Heritage Program botanists Dave Anderson and Peggy Lyon. Thanks also to Jill Handwerk for assistance in the preparation of this document. Nan Lederer at University of Colorado Museum Herbarium provided helpful information on Astragalus missouriensis var. humistratus specimens. AUTHOR’S BIOGRAPHY Karin Decker is an ecologist with the Colorado Natural Heritage Program (CNHP). She works with CNHP’s Ecology and Botany teams, providing ecological, statistical, GIS, and computing expertise for a variety of projects. She has worked with CNHP since 2000. Prior to this, she was an ecologist with the Colorado Natural Areas Program in Denver for four years. She is a Colorado native who has been working in the field of ecology since 1990. Before returning to school to become an ecologist she graduated from the University of Northern Colorado with a B.A. in Music (1982). She received an M.S. in Ecology from the University of Nebraska (1997), where her thesis research investigated sex ratios and sex allocation in a dioecious annual plant. -
A Guide to Frequent and Typical Plant Communities of the European Alps
- Alpine Ecology and Environments A guide to frequent and typical plant communities of the European Alps Guide to the virtual excursion in lesson B1 (Alpine plant biodiversity) Peter M. Kammer and Adrian Möhl (illustrations) – Alpine Ecology and Environments B1 – Alpine plant biodiversity Preface This guide provides an overview over the most frequent, widely distributed, and characteristic plant communities of the European Alps; each of them occurring under different growth conditions. It serves as the basic document for the virtual excursion offered in lesson B1 (Alpine plant biodiversity) of the ALPECOLe course. Naturally, the guide can also be helpful for a real excursion in the field! By following the road map, that begins on page 3, you can determine the plant community you are looking at. Communities you have to know for the final test are indicated with bold frames in the road maps. On the portrait sheets you will find a short description of each plant community. Here, the names of communities you should know are underlined. The portrait sheets are structured as follows: • After the English name of the community the corresponding phytosociological units are in- dicated, i.e. the association (Ass.) and/or the alliance (All.). The names of the units follow El- lenberg (1996) and Grabherr & Mucina (1993). • The paragraph “site characteristics” provides information on the altitudinal occurrence of the community, its topographical situation, the types of substrata, specific climate conditions, the duration of snow-cover, as well as on the nature of the soil. Where appropriate, specifications on the agricultural management form are given. • In the section “stand characteristics” the horizontal and vertical structure of the community is described. -
Annotated Checklist of the Vascular Plant Flora of Grand Canyon-Parashant National Monument Phase II Report
Annotated Checklist of the Vascular Plant Flora of Grand Canyon-Parashant National Monument Phase II Report By Dr. Terri Hildebrand Southern Utah University, Cedar City, UT and Dr. Walter Fertig Moenave Botanical Consulting, Kanab, UT Colorado Plateau Cooperative Ecosystems Studies Unit Agreement # H1200-09-0005 1 May 2012 Prepared for Grand Canyon-Parashant National Monument Southern Utah University National Park Service Mojave Network TABLE OF CONTENTS Page # Introduction . 4 Study Area . 6 History and Setting . 6 Geology and Associated Ecoregions . 6 Soils and Climate . 7 Vegetation . 10 Previous Botanical Studies . 11 Methods . 17 Results . 21 Discussion . 28 Conclusions . 32 Acknowledgments . 33 Literature Cited . 34 Figures Figure 1. Location of Grand Canyon-Parashant National Monument in northern Arizona . 5 Figure 2. Ecoregions and 2010-2011 collection sites in Grand Canyon-Parashant National Monument in northern Arizona . 8 Figure 3. Soil types and 2010-2011 collection sites in Grand Canyon-Parashant National Monument in northern Arizona . 9 Figure 4. Increase in the number of plant taxa confirmed as present in Grand Canyon- Parashant National Monument by decade, 1900-2011 . 13 Figure 5. Southern Utah University students enrolled in the 2010 Plant Anatomy and Diversity course that collected during the 30 August 2010 experiential learning event . 18 Figure 6. 2010-2011 collection sites and transportation routes in Grand Canyon-Parashant National Monument in northern Arizona . 22 2 TABLE OF CONTENTS Page # Tables Table 1. Chronology of plant-collecting efforts at Grand Canyon-Parashant National Monument . 14 Table 2. Data fields in the annotated checklist of the flora of Grand Canyon-Parashant National Monument (Appendices A, B, C, and D) . -
The Evolutionary Theory of Sex
TWO SEXES. Why? ♂ ♀ ♀ ? ♀ The Evolutionary Theory of Sex \ Two Sexes – Why? The Evolutionary Theory of Sex Wilmington 2011 Two Sexes – Why? The Evolutionary Theory of Sex — Wilmington, 2011. What do we need two sexes for? What is the evolutionary significance of this phenomenon? What is the nature of the differences between the sexes? What is sexual dimorphism and what does it mean? How is it connected to other life phenomena? The new evolutionary theory of sex differentiation developed by Dr. Biol. Sci. Vigen Geodakian has the answers. It provides a scientific justification for the existence of men and women, and the evolutionary roles that they play in family and society. Pages: х + 230. Ill. 49, tabl. 33, bibliography 637. ISBN xxxxx (PDF) Compiled and edited by S. V. Geodakyan. The materials presented in the book, are submitted selectively and if they contain any omissions, the responsibility for them bears the originator of this work. In case of any questions and uncertainties readers should refer to the original papers listed in the bibliography. Copyright © 2011 by Sergey Geodakyan. All rights reserved. Printed in the United States of America. No part of this publication may be reproduced or transmittedin any form or by any means without the written permission of the publisher. E-mail: [email protected] PREFACE The first publication of Vigen Geodakian devoted to the problem of sex differentiation, appeared in 1965 in cybernetic magazine. Then came an article in popular science magazine “Science and Life”. Due to multimillion distribution this article received much attention. Many scientists still remember just this one small aspect of the theory—how to influence the sex of offspring. -
The Marble Canyon Milk-Vetch
The Marble Canyon milk-vetch Astragalus cremnophylax var. hevronii A 10-year Monitoring Update Redwall Site, Coconino County, AZ Daniela Roth Navajo Natural Heritage Program P.O. Box 1480 Window Rock AZ 86515 INTRODUCTION The Marble Canyon milk-vetch (Astragalus cremnophylax var. hevronii) is one of three varieties of Astragalus cremnophylax, all three of which are extremely rare local endemics to the Grand Canyon area of northern Arizona. The Sentry milk-vetch, Astragalus cremnophylax var. cremnophylax, was Listed Endangered in 1990, primarily due to severe threats to known populations through trampling by park visitors of plants and habitat along the south rim of the Grand Canyon. The Cliff milk-vetch, A. cremnophylax var. myriorrhapis, is only known from the Buckskin Mountains, on the Arizona/Utah border, where it is managed both by the U.S. Forest Service and the BLM. A population of A. cremnophylax var. cremnophylax from the north rim of the Grand Canyon has shown to be genetically different from the south rim populations as well as the other varieties and may be described as a new species (Allphin et al. 2005). All three varieties are narrow endemics and are vulnerable to extinction due to their habitat specificity, limited habitat availability, low fecundity levels and the overall small number of plants in existence. The Redwall population of A. cremnophylax var. hevronii was first discovered by Bill Hevron, former botanist with the Navajo Natural Heritage Program, in 1991 and described by Rupert Barneby of the New York Botanical Garden in 1992. In 1997 the Grand Canyon National Park received a grant from the National Fish and Wildlife Foundation to set up monitoring plots for all three varieties of A. -
Docketed 08-Afc-13C
November 2, 2010 California Energy Commission Chris Otahal DOCKETED Wildlife Biologist 08-AFC-13C Bureau of Land Management TN # Barstow Field Office 66131 2601 Barstow Road JUL 06 2012 Barstow, CA 92311 Subject: Late Season 2010 Botanical Survey of the Calico Solar Project Site URS Project No. 27658189.70013 Dear Mr. Otahal: INTRODUCTION This letter report presents the results of the late season floristic surveys for the Calico Solar Project (Project), a proposed renewable solar energy facility located approximately 37 miles east of Barstow, California. The purpose of this study was to identify late season plant species that only respond to late summer/early fall monsoonal rains and to satisfy the California Energy Commission (CEC) Supplemental Staff Assessment BIO-12 Special-status Plant Impact Avoidance and Minimization, requirements B and C (CEC 2010). Botanical surveys were conducted for the Project site in 2007 and 2008. In response to above average rainfall events that have occurred during 2010, including a late season rainfall event on August 17, 2010 totaling 0.49 inch1, additional botanical surveys were conducted by URS Corporation (URS) for the Project site. These surveys incorporated survey protocols published by the Bureau of Land Management (BLM) (BLM 1996a, BLM 1996b, BLM 2001, and BLM 2009). BLM and CEC staff were given the opportunity to comment on the survey protocol prior to the commencement of botanical surveys on the site. The 2010 late season survey was conducted from September 20 through September 24, 2010. The surveys encompassed the 1,876-acre Phase 1 portion of the Project site; select areas in the main, western area of Phase 2; a 250-foot buffer area outside the site perimeter; and a proposed transmission line, which begins at the Pisgah substation, heads northeast following the aerial transmission line, follows the Burlington Northern Santa Fe (BNSF) railroad on the north side, and ends in survey cell 24 (ID#24, Figure 1). -
U.S. Fish and Wildlife Service Species Assessment and Listing Priority Assignment Form
U.S. FISH AND WILDLIFE SERVICE SPECIES ASSESSMENT AND LISTING PRIORITY ASSIGNMENT FORM Scientific Name: Astragalus microcymbus Common Name: skiff milkvetch Lead region: Region 6 (Mountain-Prairie Region) Information current as of: 04/08/2014 Status/Action ___ Funding provided for a proposed rule. Assessment not updated. ___ Species Assessment - determined species did not meet the definition of the endangered or threatened under the Act and, therefore, was not elevated to the Candidate status. ___ New Candidate _X_ Continuing Candidate ___ Candidate Removal ___ Taxon is more abundant or widespread than previously believed or not subject to the degree of threats sufficient to warrant issuance of a proposed listing or continuance of candidate status ___ Taxon not subject to the degree of threats sufficient to warrant issuance of a proposed listing or continuance of candidate status due, in part or totally, to conservation efforts that remove or reduce the threats to the species ___ Range is no longer a U.S. territory ___ Insufficient information exists on biological vulnerability and threats to support listing ___ Taxon mistakenly included in past notice of review ___ Taxon does not meet the definition of "species" ___ Taxon believed to be extinct ___ Conservation efforts have removed or reduced threats ___ More abundant than believed, diminished threats, or threats eliminated. Petition Information ___ Non-Petitioned _X_ Petitioned - Date petition received: 07/30/2007 90-Day Positive:08/18/2009 12 Month Positive:12/15/2010 Did the Petition -
Wojciechowski Quark
Wojciechowski, M.F. (2003). Reconstructing the phylogeny of legumes (Leguminosae): an early 21st century perspective In: B.B. Klitgaard and A. Bruneau (editors). Advances in Legume Systematics, part 10, Higher Level Systematics, pp. 5–35. Royal Botanic Gardens, Kew. RECONSTRUCTING THE PHYLOGENY OF LEGUMES (LEGUMINOSAE): AN EARLY 21ST CENTURY PERSPECTIVE MARTIN F. WOJCIECHOWSKI Department of Plant Biology, Arizona State University, Tempe, Arizona 85287 USA Abstract Elucidating the phylogenetic relationships of the legumes is essential for understanding the evolutionary history of events that underlie the origin and diversification of this family of ecologically and economically important flowering plants. In the ten years since the Third International Legume Conference (1992), the study of legume phylogeny using molecular data has advanced from a few tentative inferences based on relatively few, small datasets into an era of large, increasingly multiple gene analyses that provide greater resolution and confidence, as well as a few surprises. Reconstructing the phylogeny of the Leguminosae and its close relatives will further advance our knowledge of legume biology and facilitate comparative studies of plant structure and development, plant-animal interactions, plant-microbial symbiosis, and genome structure and dynamics. Phylogenetic relationships of Leguminosae — what has been accomplished since ILC-3? The Leguminosae (Fabaceae), with approximately 720 genera and more than 18,000 species worldwide (Lewis et al., in press) is the third largest family of flowering plants (Mabberley, 1997). Although greater in terms of the diversity of forms and number of habitats in which they reside, the family is second only perhaps to Poaceae (the grasses) in its agricultural and economic importance, and includes species used for foods, oils, fibre, fuel, timber, medicinals, numerous chemicals, cultivated horticultural varieties, and soil enrichment.