INSECTA MUNDI, Vol. 12, No. 3-4, September-December, 1998 189

Descriptions ofnew luperine genera and species from Mexico, with keys to related taxa (Coleoptera: Chrysomelidae: )

Shawn M. Clark Plant Industries Division West Virginia Department of Agriculture 1900 Kanawha Blvd., East Charleston, WV 25305-0191

Abstract. The section Scelidites ofthe subtribe is diagnosed and described. A key is provided to separate the genera within this section. Cyphotarsis Jacoby is reduced to a junior synonym of Metacoryna Jacoby. Microscelida, new genus, is erected to include Agelastica viridis Jacoby, subcostatus Jacoby, Luperus subglabratus Jacoby, Scelidopsis violacea Jacoby, and seven newly described species. A key and diagnoses are provided to enable recognition of each species within this new genus. Scelidacne, new genus, is erected to include a single newly described species.

Introduction Many New World species of Scelidites were As classified by Seeno & Wilcox (1982), the originally described in Luperodes Motschulsky, a section Scelidites is placed in the family Chry­ genus properly belonging to the Monoleptites (a somelidae, subfamily Galerucinae, tribe Luperini, quite unrelated world-wide group that can usually and subtribe Luperina. That classification is fol­ be recognized by the very long basitarsus of the lowed in this study. Even so, I favor the recent hind leg). Many other American species of Scelid­ classifications of Reid (1995) and other authors ites were formerly classified in Luperus Muller. who treated the traditional subfamilies Galeruci­ However, the type species ofthat genus belongs in nae and Alticinae as mere tribes in the expanded the Luperites, a group that is only distantly relat­ subfamily Galerucinae. If their classifications are ed, is restricted to the Eastern Hemisphere, and followed, the ranks of tribe, subtribe, and section may be recognized by the form of the male anten­ may eventually need to be adjusted downward. nae (antennomere II is very short, and III is In the Old World, genera of Scelidites are unusually long). similar to those found in America, but, with the The biology of the Scelidites is essentially single exception of Eugalera Brancsik (already unknown. To my knowledge, the larvae of only one recognized as a synonym of Crotch by New World species have been reported (Wilcox, Wilcox, 1973), I have not found them to be synon­ 1965), and they are similar to root feeders such as ymous. Genera ofthe principally Old World section Chevrolat. Some species of Scelidites Exosomites are also similar to Scelidites in many are apparently restricted in their acceptance of respects, and further study may show that the two host plant species, but others may survive well on taxa should be combined. several hosts. Unfortunately, the hosts of most In the Western Hemisphere, species of Scelid­ species are not known. ites are found from Canada to Panama, but they are Methods most numerous and diverse in the southwestern Although species of Scelidites are currently United States and in Mexico. No species are known recognized from North America, Europe, Asia, and from South America. Africa, only the New World species were carefully In the New World, the genera of Scelidites are studied during my recent investigations, with Old similar to Pteleon Jacoby (section Exosomites) World genera being examined only enough to and, to a lesser extent, to Chevrolat determine that they are not synonymous with (section Phyllobroticites). Beyond this, the Scelid­ those of the Western Hemisphere. ites are quite distinct from all other American Specimens were borrowed from the following Galerucinae, although they were often confused curators and collections (acronyms following the with other groups by earlier workers. entries are referred to throughout this paper); 190 Volume 12, No. 3-4, September-December, 1998, INSECTA MUNDI

Sharon L. Shute, British Museum (Natural Histo­ Pubescence of head largely confined to more ry) [BMNH]; Robert S. Anderson, Canadian Muse­ distal areas. Interocular sulcus present though um of Nature [CMNC]; Laurent LeSage, Canadian sometimes weakly developed. Frontal tubercles National Collection ofInsects [CNCI]; Edward G. usually separated from mesal frontal carina by Riley (private collection), College Station, Texas distinct sulci. Antennae subdivided into 11 ant en­ [EGRC]; John B. Kethley, Field Museum of Natu­ nomeres; antennomere II distinctly shorter than I ral History [FMNH]; Kathryn C. McGiffen, Illinois or IV, slightly shorter than III. Natural History Survey [INHS]; Alfred F. Newton, Pronotum wider than long; lateral beads well­ Jr. and Scott Shaw, Museum of Comparative Zool­ developed; long seta present at each angle; fringe ogy [MCZC]; Richard E. White, United States of short setae usually present along all margins, National Museum [NMNH]; Charles A. Triple­ although often obsolete on lateral margins; disc horn, Ohio State University [OSUC]; Paul K. Lago lacking pubescence. (private collection), University, Mississippi Elytra lacking conspicuous pubescence, usual­ [PKLC]; Shawn M. Clark (private collection), ly with scattered setae on apical half; elytral Charleston, West Virginia [SMCI]; Edward G. punctures confused; epipleuron well developed Riley, Texas A. & M. University [TAMU]; Univer­ near base, often extending to near apex. sity of Arizona [UAlC]; Saul I. Frommer, Univer­ Ventral portion of prothorax with fringe of sity of California, Riverside [UCRV]; Mark F. setae along anterior and posterior margins. Front O'Brien, University of Michigan [UMMZ]. coxae contiguous or rather narrowly separated by I have placed much emphasis on the form ofthe prosternum; front coxal cavities open behind. Mid­ aedeagi, a brief preliminary study indicating that dle coxae usually narrowly separated, sometimes the spermathecae lacked useful diagnostic charac­ contiguous. Metathorax, especially episternum, ters. Aedeagal preparations were made following usually pubescent. Hind coxae well separated. techniques similar to those described by Smith Tarsi subdivided into five tarsomeres; tarsomeres (1979) and Reid (1992). I-III densely setose beneath; III bilobed; IV very small. Tarsomere I of hind leg usually shorter than Section Scelidites Chapuis II-V combined. Tarsal claws usually appendiculate, rarely bifid. Scelidites Chapuis, 1875:184; Wilcox, 1965:94; Seeno and Abdomen with five visible segments, usually Wilcox, 1982:109. pubescent; terminal segment of male often im­ Androlyperini Leng, 1920:298. pressed, usually with subrectangular lobe at apex; terminal segment of female not impressed, round­ Diagnosis. In this group, the antennae extend ed or angulate at apex. Aedeagus lacking well­ at least to near the middle of the elytra but not to developed basal spurs. the apex. The elytra are, at most, sparsely and Comments. The tribe Galerucini differs from rather inconspicuously pubescent. The epipleura, Scelidites in that the abdomen of the male lacks a at least near the base, are easily visible and not rectangular lobe, and the aedeagus usually has unusually narrow. The basal tarsomere ofthe hind well-developed basal spurs. Also, the tarsal claws leg is usually shorter than the following tarsomer­ are usually bifid, and the elytra are often conspic­ es combined. The tarsal claws are usually appen­ uously pubescent. diculate. The abdomen of the male usually has a The tribe Metacyclini differs from Scelidites in distinct, apical, subrectangular lobe that is either that the abdomen of the male lacks a rectangular much shorter than wide or that is distinctly im­ lobe, and the aedeagus has well-developed basal pressed or bent toward the dorsum. And the spurs. aedeagus lacks well-developed basal spurs. These The tribe Sermylini is an Old World group, but characters will enable recognition of most New Agelastica alni (Linnaeus) and Sermylassa halen­ World species of Scelidites, distinguishing them sis (Linnaeus) have been reported from North from other galerucines (for questionable taxa, see America. This tribe differs from Scelidites in that additional discussion in the comments section the lobe at the apex of the male abdomen is rounded below). rather than rectangular, and the basal foramen of Description. Form elongate or elongate-oval; the aedeagus is nearly circular. head narrower than prothorax; prothorax narrow­ Within the tribe Luperini, the subtribe Diabro­ er than elytra. ticina differs from Scelidites in that the apex of the INSECTA MUNDI, Vol. 12, No. 3-4, September-December, 1998 191 male abdomen lacks a rectangular lobe. Also, the Pseudoluperus, a no doubt polyphyletic assem­ tarsal claws of many species are bifid. Within this blage that will almost certainly require further subtribe, the section Phyllecthrites is especially resolution. likely to be confused with Scelidites but differs in I consider Cyphotarsis Jacoby to be synony­ that the male has a notch near the apex of the mous with Metacoryna Jacoby, and it therefore middle tibia. does not appear in the generic key presented Within the subtribe Luperina, the New World below. The reasons for this synonymy will be representatives of the section Phyllobroticites presented in a forthcoming revision of Metacory­ differ from Scelidites in that the epipleuron is na. absent or very much reduced. The section Exo­ Many of the genera in Scelidites are best somites is represented in the New World by four recognized by male characters. Because of this, the species of Pteleon in which the antennae do not following key is based largely on males. However, extend much beyond the humeri of the elytra. The the experienced worker will discover that unasso­ section Monoleptites has a sclerotized covering at ciated females can also be identified, although the aedeagal orifice, and tarsomere I of the hind leg often with considerably more difficulty. is usually extremely long. Within the Scelidites, some species of Scelida Key to New World genera of Scelidites (males) Chapuis have unusually long basitarsi on the hind legs. However, in such species, the apex of the male 1. Basal bead of pronotum complete although of- abdomen is strongly impressed and the aedeagal ten small, extending without interruption orifice lacks a sclerotized covering. Either ofthese between posterior angles (this character ob­ characters distinguishes Scelida from non-Scelid­ scured by extremely coarse pronotal puncta­ tion in Synetocephalus autumnal is Fall, a ites with such tarsi. yellow or reddish species from California) .. Scelida metallica Jacoby and some species of ...... 2 Jacoby have bifid tarsal claws. However, Basal bead of pronotum absent, or restricted to the rectangular lobe at the apex of the male extreme lateral areas (pronotal punctation abdomen is conspicuous and distinguishes these never coarse, except in metallic blue Sceli- species from non-Scelidites with bifid claws. dacne from Mexico) 8 In several species of Scelidites, the rectangu­ lar lobe at the apex ofthe abdomen is very short and 2(1). Elytra with a distinct impression at posterolat- not easily visible. It is especially inconspicuous in eral angle; distal portion of epipleuron bent upward at point of impression; genal length some species of Metacoryna Jacoby Scelidac­ and as great as or often greater than length of ne, new genus, but these species may be recognized antennomere II; all tibiae with terminal spurs; by the presence of either abdominal appendages or elytra not metallic Androlyperus Crotch swollen antennae in the males. Elytra normal, lacking obvious modifications. In some species of Triarius, the lobe at the ...... 3 apex ofthe male abdomen is about as long as broad. Many species of non-Scelidites (Monoleptites) also 3(2). Intermediate or distal antennomeres distinctly have subquadrate lobes, but they differ from Tri­ swollen; genae about as long as antennomere arius in that the aedeagal orifice has a sclerotized II; tibial spurs absent; elytra entirely dark ...... Metacoryna Jacoby covering, of leg and the basal tarsomere the hind Antennae usually slender; if antennae are swol­ is usually longer than tarsomeres II-V combined. len, length of genae is about halfthat of anten- The aedeagi of some species of Pseudoluperus nomere II 4 Beller & Hatch have small basal spurs that might be confused with the larger spurs found in the 4(3). Rectangular lobe at apex of abdomen at least tribes Galerucini or Metacyclini. However, the half as long as broad, large, not strongly bent male abdomens of these species of Pseudoluperus towards dorsum; length of antennomere I have the impressed, rectangular lobe that is char­ subequal to maximum diameter of eye; tarsal acteristic of Scelidites. claws often bifid; elytra not metallic; tibiae with large, conspicuous, terminal spurs Tri­ Following several new combinations published arius Jacoby herein and in forthcoming papers, most of the New Rectangular lobe at apex of abdomen either World genera of Scelidites will form well-defined, shorter or distinctly bent towards dorsum; homogeneous groups. One notable exception is antennomere I usually shorter; if tarsal claws 192 Volume 12, No. 3-4, September-December, 1998, INSECTA MUNDI

are bifid, elytra are metallic; tibial spurs of- 8(1). Basitarsus of hind leg enlarged, much different ten smaller or absent 5 in form from basitarsi of front or middle legs ...... 9 5(4). Distance between antennal fossae equal to at Basitarsus of hind leg not conspicuously en- least twice diameter of fossa; mesal frontal larged 10 carina broad, nearly continuous with frontal tubercles, delimited behind by shallow, some­ 9(8). Abdomen with ventral appendages; hind tibia times indistinct impression; elytra usually with curved apical appendage; basitarsi of all pale with darker stripes or entirely pale, legs enlarged, those of front and middle legs rarely entirely dark with very faint metallic about as broad as apices oftibiae, that of hind luster, never with obvious metallic luster; leg very irregular in form; color nearly entire­ aedeagal orifice often with a sclerotized cov- ly metallic blue ...... Scelidacne, new genus ering Synetocephalus Fall Abdomen normal, without ventral appendages; Distance between antennal fossae equal to less hind tibia without apical appendage; basitar­ than twice diameter of fossa; mesal frontal si of front and middle legs much narrower carina narrower, usually separated from fron­ than apices oftibiae, similar in form to basi­ tal tubercles by distinct sulci; transverse in­ tarsi of female; color largely yellow, with terocular sulcus usually distinct, abruptly elytra bicolored Keitheatus Wilcox delimiting frontal tubercles behind; elytral color variable, often with distinct metallic 10(8). Front femora much larger than middle or hind luster; orifice of aedeagus lacking sclerotized femora; aedeagus strongly asymmetrical ..... covering 6 . Lygistus Wilcox Front femora not abnormally large; aedeagus 6(5). Elytra with a conspicuous transverse impres­ symmetrical 11 sion at basal third; body, including appendag­ es, strongly metallic; species occurring in 11(10). Orifice of aedeagus with a sclerotized covering; Costa Rica Inbioluperus Clark at least hind legs with terminal spurs; abdo­ Transverse impression of elytra absent or weak­ men lacking ventral appendages Microsceli­ ly developed; front tibiae or other body parts da, new genus lacking metallic luster; distribution in Mexi­ Orifice of aedeagus usually lacking sclerotized co and northward [except Pseudoluperus fla­ covering (other characters variable); if scle­ uomarginatus (Jacoby), a pale, non-metallic rotized covering present, terminal spurs ab­ species known from Guatemala and Panama] sent from all tibiae and abdomen with ventral ...... 7 appendages 12

7(6). Mesal frontal carina narrow, usually forming 12(11). Antennae short, reaching only a little beyond angulate ridge; frontal tubercles often ex­ the humerus; antennomeres II-IV about equal tending to and continuous with orbit, rarely in length and width .. delimited laterally by poorly defined impres­ ...... (Pteleon Jacoby, section Exosomites) sion; genae usually about as long as terminal Antennae longer, reaching to middle of elytra article of maxillary palp; vertex often with or farther 13 alutaceous microsculpture; head and elytra dark, often metallic; apex of aedeagus often 13(12). Pronotum usually entirely pale, but if partially truncate, emarginate, or asymmetrical, rare- black then abdomen with conspicuous ventral ly symmetrical and pointed . appendages; tibiae without terminal spurs; if ...... Sceloiyperus Crotch elytra partially pale, body longer than 5.0 mm If mesal frontal carina is narrow or acutely ...... Scelida Chapuis elevated, frontal tubercles are delimited lat­ Pronotum often partially or entirely dark; if erally by distinct impression; genae usually pronotum entirely pale, then elytra largely or not much more than half as long as terminal entirely pale also, tibiae with terminal spurs, article of maxillary palp; vertex not distinctly and body length less than 5.0 mm . alutaceous; head and elytra sometimes large­ ...... Pseudoluperus Beller & Hatch (part) ly pale; apical portion of aedeagus usually pointed and symmetrical, rarely strap-like Scelidacne Clark, and asymmetrical, not truncate or emargin­ new genus ate ...... Peeudoluperus Beller & Hatch (part) Diagnosis. Males of this genus are immedi­ ately recognizable by the enlarged, oddly shaped INSECTA MUNDI, Vol. 12, No. 3-4, September-December, 1998 193 basitarsus of the hind leg and by the curved apical appendage of the hind tibia (fig. 2b). Either males or females are separable from other Scelidites by the absence of a basal bead on the pronotum, in combination with the coarsely punctate pronotum and the nearly uniform metallic blue color. Description. Form elongate; head slightly narrower than prothorax; prothorax much nar­ rower than elytra. Pubescence of head largely confined to more distal areas. Frontal tubercles well defined, sepa­ rated from each other by deep sulcus. Mesal frontal carina well-developed, short. Antennae subdivided into 11 antennomeres; antennomere III much longer than II, distinctly longer than IV, shorter than 1. Pronotum wider than long; lateral beads well­ developed; a long seta present at each angle; a fringe of short setae present along anterior and posterior margins; disc lacking pubescence. Elytra with a few scattered, inconspicuous setae; elytral punctures confused; epipleuron broad basally, narrow apically. Ventral portion of prothorax with a fringe of setae along anterior and posterior margins. Front coxae very narrowly separated by prosternum, nearly contiguous; front coxal cavities open be­ Figure 1. Scelidacne andrewi, new species, male. hind. Middle coxae narrowly separated. Metatho­ rax, especially episternum, pubescent. Hind coxae widely separated. Terminal spurs present on all tibiae offemale, absent from all tibiae of male. Each Scelidacne andrewi Clark, tarsus subdivided into five tarsomeres; tarsomer­ new species es I-III densely setose beneath',,III broad bilobed:, (Figures 1, 2a-e) IV very small. Male with basal tarsomeres of front and middle legs broadened, with basal tarsomere Diagnosis. This, the single known species of hind leg greatly enlarged, flattened (fig. 2b). included in this genus, can be recognized by the Tarsal claws appendiculate. characters given in the generic Diagnosis. Abdomen with five visible segments, pubes­ Description. Form elongate; prothorax nar­ cent; sternites of male greatly modified, with well­ rower than elytra. Color metallic blue, with anten­ developed appendages (fig. 2a). Aedeagus lacking nae and tarsi brown. Male with abdominal append­ basal spurs. ages (fig. 2a) and enlarged basal tarsomeres (figs. Type species. Scelidacne andre wi, new spe­ 1,2b). cies. Head shining, not alutaceous. Vertex largely Comments. The rectangular lobe at the apex glabrous, but with several setae near each eye. of the male abdomen, so characteristic of most Distal and subocular areas densely pubescent. Scelidites, is very much reduced in the single Eyes slanting, with internal margins diverging known species of this genus. However, based on posteriorly. Interocular distance, measured along other similarities, this genus clearly fits into this group. posterior margin of frontal tubercles, equal to 0.6 times maximum width of head across eyes. Frontal The name, Scelidacne, is a euphonious varia­ tubercles sub-triangular, mesially contiguous for a tion of Scelida, not intended to have meaning in distance slightly greater width of antenno­ Latin or Greek languages. I designate its gender as than female. mere I, separated from each other by deep sulcus, 194 Volume 12, No. 3-4, September-December, 1998, INSECTA MUNDI

Elytra together 1.7 times as long broad, broad­ est near apical fourth. Several poorly developed longitudinal costae sometimes present, most no­ ticeable in some females. Punctures coarse, dense, mostly separated by a distance about equal to diameter of puncture, sometimes partially conflu­ ent and forming short, transverse rugae. Inter­ punctural areas opaque, distinctly less shining than interspaces of pronotum. Ventral areas of prothorax shining, not aluta­ ceous; anterior margin clothed with double fringe of long setae, with one fringe directed anteriorly /\ /) and another directed ventrally. Mesothorax and ,\ \ /( meta thorax largely pubescent, especially lateral­ I\ /I I I ly. Legs of female normal, with basitarsi much narrower than apices of tibiae. Male with basitarsi ) \ ,/,/ offront and middle legs greatly expanded, about as \~)I wide as tibial apices (fig. 1); hind tibia with curved I !\ I I appendage at apex (fig. 2b); hind basitarsus greatly II I enlarged, irregular in shape, with fine basal ridges i ' opposite tibial appendage (fig. 2b). Abdomen pu­ I i bescent, alutaceous; female lacking unusual mod­ ifications, with terminal segment rounded apical­ I ly; segment III of male with trilobed appendage, \ I: , ;' c D E the lateral lobes being densely pubescent, the '... ~_/ v median glabrous lobe being bifurcate apically (fig. Figure 2. Scelidacne andrewi, new species. a) abdomen 2a); male with terminal segment very short, with of male, ventral view; b) distal portion of hind leg of male; apical margin concave. c) aedeagus, dorsal view; d) aedeagus, ventral view; e) Aedeagus symmetrical in dorsal view, with aedeagus, lateral view. acute tip (fig. 2c). Two dorsally directed flanges present near mid-length, one on each side of orifice (fig. 2e). Ventral surface with a median grove abruptly delimited behind, separated from mesal extending from basal foramen to near apex, this frontal carina by short sulcus, delimited laterally grove terminating in a deep, round, subapical by broad impression adjacent to eye. Antennal concavity (fig. 2d). Basal spurs absent. fossae separated from each other and from eyes by Material examined. Holotype male and allo­ distance less than diameter of fossa. Mesal frontal type female: Mexico, Chiapas, 5 mi. W. San Cris­ carina short, convex, not acute. Genal length slightly tobal L. C., 6 May 1969, H. F. Howden (CMNC). greater than width of distal article of maxillary Paratypes: same data as holotype (I male, 1 fe­ palp. Antennae extending to near middle of elytra; male, CMNC; 1 male, 1 female, SMCI); same data antennomere III 0.6 times as long as I, 2.7 times as as holotype except 10 May 1969 (1 female, CMNC); long as II, 1.4 times as long as IV; IV-X subequal in same data as holotype except 11 May 1969 (1 length; XI 1.4 times as long as X, emarginate near female, CMNC); same data as holotype except 3 apical third, pointed at apex. May 1969 and beating oak (2 females, CMNC). Pronotum 1.5 times as wide as long, 0.7 times Comments. The male abdominal appendages as wide as elytra across humeri, widest near (fig. 2a) call to mind certain species of the genus anterior third; sides sinuate in dorsal view. Lateral Scelida. However, the modifications of the male beads present; anterior and basal beads absent. legs (fig. 2b), as well as the presence of tibial spurs Disc with anterior third shallowly, transversely in the females, are unlike the species in that genus. impressed; a deep fovea present on each side, The name of this new species honors my son, slightly behind middle. Surface densely, coarsely Andrew. punctate; area between coarse punctures shining, not alutaceous, but with a few fine punctures. INSECTA MUNDI, Vol. 12, No. 3-4, September-December, 1998 195

Microscelida Clark, Basal half of pro notal disc with a conspicuous new genus fovea at each side (fig 3); diameter of fovea subequal to one-fourth width of prothorax; Diagnosis. The absence of a basal bead on the length 3.8-4.1 mm; Mexico (Michoacan to Mex­ ico to Guerrero) ...... foveicollis, new species pronotum, in combination with the sclerotized orificial covering of the aedeagus, usually distin­ 3(2). Interspaces ofelytra polished, with minute punc- guishes this from other New World genera of tures either absent or widely spaced 4 Scelidites. An orificial covering is also present in Most areas of elytra with numerous minute, an undescribed species of Scelida, but the tibiae of closely spaced punctures in area between any that species lack terminal spurs. In this new genus, two larger punctures; length 4.5 mm; Mexico spurs are present on at least the hind tibiae. The (Guerrero) viridis (Jacoby) elytra of Microscelida are largely or entirely dark. Description. Form elongate or elongate oval; 4(3). Aedeagus gradually narrowed to near apex, usually with a short, median slit or emargin­ prothorax narrower than elytra. Pronotum wider ation at apex (fig. fie); elytra of female dis­ than long; margins setose; disc lacking pubes­ tinctly costate; elytra of male sometimes cence; lateral beads well developed; anterior and vaguely costate; length 4.4-4.9 mm; Mexico basal beads absent. Elytra dark, with a few erect (Michoacan to Veracruz to Chiapas) . setae in distal half; epipleura well-developed, ex­ ...... subcostata (Jacoby) tending nearly to apex. Ventral areas of prothorax Apex of aedeagus usually not emarginate; if largely glabrous; anterior margin setose; front apex of aedeagus is emarginate, apical por­ coxae contiguous or nearly so; front coxal cavities tion of aedeagus is more rounded (fig. 4b); open behind. Metathorax pubescent, at least later­ elytra not costate 5 ally. Terminal spurs present on hind tibiae, often 5(4). Apex of aedeagus pointed 6 present on other tibiae. Tarsal claws appendicu­ Apex of aedeagus rounded or with median emar­ late. Abdomen pubescent, lacking appendages; gination (fig. 4b); length 3.6-4.5 mm; Mexico terminal segment of male with an impressed, more (Guerrero) subglabrata (Jacoby) or less rectangular, apical lobe; terminal segment of female rounded at apex. Aedeagus symmetrical, 6(5). Orifice of aedeagus largely covered by a median lacking basal spurs; orifice covered by a median, sclerotized process and by two lateral sclero­ sclerotized process which is continuous basally tized processes, one at each side (figs. fid-e); with tubular portion of aedeagus; additional scle­ lateral processes meeting or nearly meeting medially, distal to median process 8 rotized processes sometimes extending from lat­ Covering of aedeagal orifice consisting of a me­ eral areas and covering orifice. dian sclerotized process; lateral processes Type species. Microscelida viridipennis, new absent (figs. 4d, 5a) 7 species. Comments. Most of the species in this genus 7(6). Ventral surface of aedeagus with longitudinal are similar in appearance. Often, accurate identi­ groove; length 3.4-4.1 mm; Mexico (Oaxaca to fication of species is possible only through exami­ Chiapas) wilcoxi, new species nation of the aedeagus. Ventral surface of aedeagus without longitudi­ In many characters, this genus resembles Scel­ nal groove; length 3.1-4.1 mm; Mexico (Oax- aca) moweri, new species ida, but the species are smaller. Accordingly, I have chosen the name Microscelida. 8(6). Distal fourth of aedeagus attenuate, gradually narrowed to acute apex (fig. 5e); length 3.1-4.5 Key to species of Microscelida mm; Mexico (Jalisco to Zacatecas to Veracruz to Oaxaca) viridipennis, new species 1. Femora, at least offront legs, largely or entirely Distal eighth of aedeagus more abruptly nar­ pale, testa ceous; distal portion offemora dark rowed to acute apex (fig. 5d); length 3.9 mm; in some species 2 Mexico (Guerrero) ...... wellsi, new species Femora dark brown or black 9 9(1). Vertex and elytra distinctly alutaceous; tibiae 2(1). Pronotal disc evenly convex or slightly, mesial­ conspicuously paler than femora; length 4.8 ly flattened; distinct pronotal impressions mm; Mexico (Morelos) . small or absent 3 ...... alutacea, new species 196 Volume 12, No. 3-4, September-December, 1998, INSECTA MUNDI

Vertex and elytra polished, not alutaceous; tib­ nal fossae subequal to length of antennomere 1. iae not obviously paler than principal color of Mesal frontal carina convex. Genal length subequal femora 10 to three times diameter of an ommatidium. Anten­ nae brown; antennomere II much shorter than I, 10(9). Aedeagus with a short, median slit or emargin­ slightly shorter than III, about half as long as IV. ation at apex (fig. 5c); apical fourth of aedea­ gus with a narrow, median, ventral groove; Pronotum 1.3 times as wide as long, 0.7 times elytra of female distinctly costate; elytra of as wide as elytra across humeri, widest slightly male sometimes vaguely costate; length 4.4­ anterior to middle; sides arcuate in dorsal view. 4.9 mm; Mexico (Michoacan to Veracruz to Lateral beads present; anterior and basal beads Chiapas) subcostata (Jacoby) absent. Punctures distinct, especially near scutel­ Apex of aedeagus pointed, not emarginate, lack­ lum. Interspaces shining, not alutaceous. Color ing median, ventral groove; elytra not costate testaceous ...... 11 Elytra together 1.7 times as long as broad; sides nearly parallel in anterior two-thirds. Punc­ 11(10). Distal fifth of aedeagus very narrow, attenuate (fig. 4c); length 3.8-4.3 mm; Mexico (Guerrero) tures distinct, separated by a distance subequal to ...... whitingi, new species twice diameter of a puncture. Interspaces aluta­ Distal portion of aedeagus broadly curved to ceous. Color dark brown with metallic blue luster. apex, not attenuate (figs. 4d-e) 12 Ventral areas of prothorax testaceous, largely glabrous; front coxae contiguous; front coxal cav­ 12(11). Orifice of aedeagus largely covered by a median ities open behind. Mesothorax dark brown. Met­ sclerotized process and by two lateral sclero­ athorax dark brown, covered with pale pubes­ tized processes, one at each side (fig. 4e); cence. Coxae, trochanters, and femora dark brown. length 3.5-3.8 mm; Mexico (Guerrero) . Tibiaepale brown; male with terminal spurs present ...... uiolacea (Jacoby) Orificial covering of aedeagus consisting of a on hind tibiae only. Tarsi pale brown; basitarsi of median sclerotized process (fig. 4d); lateral male not obviously dilated; tarsomere I of hind leg coverings absent; length 3.1-4.1 mm; Mexico about as long as II-III combined; tarsal claws (Oaxaca) moweri, new species appendiculate. Abdomen brown, pubescent; termi­ nal segment of male semicircularly impressed at apex. Microscelida alutacea Clark, Aedeagus symmetrical. Orifice with a sclero­ new species tized covering (fig. 5b). (Figure 5b) Material examined. Mexico, Morelos, Xochi­ calco, 15 Sept. 1942, W. F. Foshag (holotype male, Diagnosis. The alutaceous microsculpture of NMNH). the vertex and elytra, in combination with the tibiae that are distinctly paler than the femora, Microscelida foueicollis Clark, distinguish this from other species of Microsceli­ new species da. (Figures 3, 4a) Description. Form elongate; prothorax nar­ rower than elytra. Vertex, elytra, mesosternum, Diagnosis. The two large, conspicuous im­ metasternum, abdomen, and femora black or dark pressions, one at each side of the pronotum, brown; elytra with blue luster. Antennae, protho­ distinguish this from other species of Microscelida rax, tibiae, tarsi, and distal areas of head paler (fig. 3). brown or testaceous. Length of male 4.8 mm. Description. Form elongate oval; prothorax Vertex punctate and pubescent near frontal narrower than elytra. Color black or dark brown, tubercles, distinctly alutaceous, black. Transverse with prothorax, legs, and at least part of head interocular sulcus deep; each half arcuate. In­ testaceous; elytra with blue or purple luster. Length terocular distance equal to 0.6 times maximum of male 4.1 mm; length of female 3.8-4.0 mm. width of head across eyes. Frontal tubercles and Head shining, not alutaceous; color entirely more distal areas of head brown or testaceous. testaceous, or testaceous with vertex and orbits Frontal tubercles separated from each other and dark brown, nearly black. Vertex minutely punc­ from mesal frontal carina by deep sulci; surface tate, with a few lateral setae, without mesal setae. shining, not alutaceous. Distance between anten- Transverse interocular sulcus deep. Interocular INSECTA MUNDI, Vol. 12, No. 3-4, September-December, 1998 197

greater and sometimes less than diameter of a puncture; punctures nearer base or apex finer. Interspaces shining, not alutaceous. Color black or dark brown, with strong blue or purple luster; epipleuron sometimes pale. Ventral areas of prothorax testaceous, shin­ ing, largely glabrous, not alutaceous; front coxae contiguous or nearly so; front coxal cavities open behind. Metathorax black or dark brown, pubes­ cent. Legs testaceous, sometimes with darker tarsi; terminal spurs present on all tibiae of fe­ male, present on only hind tibiae of male; basitarsi of male not dilated, similar to those of female; basitarsus of hind leg subequal in length to tars om­ eres II-III combined; tarsal claws appendiculate. Abdomen black or dark brown, pubescent; termi­ nal segment of male apically, semicircularly im­ pressed. Aedeagus symmetrical in dorsal view, acute at Figure 3. Microscelida foueicollis, new species. apex. Orifice covered by three sclerotized process­ es, one median and two lateral; lateral processed meeting or nearly meeting medially distal to me­ distance equal to 0.6 times maximum width of head dian process (fig. 4a). across eyes. Frontal tubercles mesially contiguous Material examined. Holotype male and allo­ for a distance less than width of antennomere II, type female: Mexico, Mexico, 1 mi. N. Valle de separated from each other and from mesal frontal Bravo, 6500', VIII-7-82, C. & L. O'Brien & G. carina by deep sulci, separated from orbit by Wibmer (NMNH). Paratypes: Mexico, Guerrero, distinct impression. Antennal fossae separated 13 km. N. ofTaxco, 5300' CRt. 3, km 147), oak pine from each other by a distance greater than width scrub, VI-21-1948, W. Nutting, F. Werner (1 male, of antennomere 1. Mesal frontal carina convex. 1 female, SMCI); Mexico, Guerrero, 38 km. W Genal length subequal to width of distal article of Iguala, 21 July 1987, R. Turnbow (1 male, RHTC); maxillary palp. Antennae slender, extending to Mexico, Mexico, 1 mi. N. Valle de Bravo, 6500', near middle of elytra; color black or dark brown, VIII-7-82, C. & L. O'Brien & G. Wibmer (1 female, with basal portion largely or entirely paler; length EGRC); Mexico, Michoacan, 49 mi. se. Aquila, July of antennomere II about half that of! or IV, about 13, 1984, Carroll, Schaffner, Friedlander (1 male, two-thirds that of III; V slightly shorter than IV, 1 female, SMCI; 3 males, 4 females, TAMU); slightly longer than VI; VI-X subequal in length; XI Mexico, Michoacan, 14.3 km. S. Uruapan, 1370­ longer than X. 1465 m, 29.vii.88, R. S. Anderson, oak-Acacia wood­ Pronotum 1.3 times as wide as long, 0.7 times land, 88-10 (1 male, 1 female, TAMU). as wide as elytra across humeri, widest near Comments. The elytra of some specimens anterior third; sides rounded or weakly sinuate in have several longitudinal costae, this being most dorsal view. Lateral beads well-developed; anteri­ pronounced in females. However, this character is or and basal beads absent. Disc with two distinct variable, the costae being entirely absent from foveae, one at each side slightly behind middle, other specimens. each separated from lateral margin, basal margin, and other fovea by a distance slightly less than Microscelida mouieri Clark, diameter of fovea (fig. 3). Surface minutely, incon­ new species spicuously punctate, shining, not alutaceous. Col­ (Figure 4d) or testaceous. Elytra together 1.8 times as long as broad, Diagnosis. The vertex and elytra of this spe­ sometimes costate; sides nearly parallel in anteri­ cies are polished, without alutaceous microsculp­ or three-fourths. Punctures near basal third con­ ture. Also, the distal portion of the aedeagus is not spicuous, coarse, separated by a distance not much unusually slender or attenuate, and the aedeagal 198 Volume 12, No. 3-4, September-December, 1998, INSECTA MUNDI orifice is covered by a single, median, sclerotized process (fig. 4d). Ventrally, the aedeagus lacks a median groove. These characters distinguish this from other species of Microscelida. Description. Form elongate oval; prothorax narrower than elytra. Color dark brown or black; A B elytra with faint blue luster. Length of male 3.1-4.0 mm; length of female 3.6-4.1 mm. Head dark brown or black, shining, not aluta­ ceous. Vertex shallowly, mesially impressed adja­ cent to frontal tubercles, with a few setae near each eye, otherwise nearly glabrous. Transverse in­ terocular sulcus deep; each half arcuate. Interoc­ ular distance equal to 0.6 times maximum width of c head across eyes, nearly as great as length of antennomere 1. Frontal tubercles separated from y each other by deep sulcus, delimited laterally by broad impression, often nearly confluent with mesal frontal carina. Mesal frontal carina nearly flat. Genal length subequal to width of distal article of maxillary palp. Antennae slender, black or dark D E brown distally, paler near base; antennomere II much shorter than I, slightly shorter than III; III nearly as long as IV. Figure 4. Microscelida spp., dorsal and lateral views of Pronotum 1.4 times as wide as long, 0.8 times aedeagi. a) M. foveicollis, new species; b) M. subglabrata as wide as elytra across humeri, widest near (Jacoby); c) M. whitingi, new species; d) M. moweri, new anterior third; sides arcuate in dorsal view. Lateral species; e) M. violacea (Jacoby). beads well developed; anterior and basal beads absent. Disc evenly convex. Punctures sometimes sclerotized process (fig. 4d). Ventral surface with­ conspicuous near margins, otherwise minute. Col­ out median groove. or testaceous, dark brown, or black. Material examined. Holotype male and allo­ Elytra together 1.6 times as long as broad; type female: Mexico, Oaxaca, 13 mi. SE. Tlalcolula, sides nearly parallel in anterior two-thirds. Most 6100', VI-2-1974, O'Brien & Marshall (NMNH). elytral punctures fine or minute, widely spaced; Paratypes: Mexico, Oaxaca, 7.7 mi. s. Ejutla, July punctures near basal third larger. Interspaces 20,1974, Clark, Murray, Ashe, Schaffner (2 males, shining, not alutaceous. Color black or dark brown, 2 females, TAMU); Mexico, Oaxaca, 2.7 mi. nw. El with weak blue luster. Camaron, July 14, 1971, Clark, Murray, Hart, Ventral surface black or dark brown; protho­ Schaffner (1 male, 2 females, TAMU); Mexico, rax sometimes testaceous. Prothorax largely gla­ Oaxaca, 3.4 mi. se. Matatlan, July 12, 1981, Bogar, brous, shining, not alutaceous; front coxae contig­ Schaffner, Friedlander (1 male, TAMU); Mexico, uous or nearly so; front coxal cavities open behind. Oaxaca, 3 mi. se. Matatlan (Microondas road), elev. Metathorax pubescent. Legs variable in color, 6650 ft., July 17,1987, Kovarik, Schaffner (1 male, usually dark brown, sometimes pale yellowish SMCI); Mexico, Oaxaca, 2 mi. n. Miahuatlan, July brown; terminal spurs present on all tibiae of 14, 1973, Mastro & Schaffner (1 male, 1 female, female, present on hind tibiae of male; basitarsi of SMCI; 8 males, 9 females, TAMU); Mexico, Oax­ male similar to those of female, not dilated; tar­ aca, 1800m, 11 km E Mitla, 3-VIII-1986, H.&A. somere I of hind leg about as long as II-III com­ Howden (1 male, CMNC); Mexico, Oaxaca, Oaxaca, bined; tarsal claws appendiculate. Abdomen brown, 6-24 June 1968, G. Pollard (8 males, 7 females, pubescent; terminal segment of male impressed at CMNC); Mexico, Oaxaca, Oaxaca, 16 June 1968, G. apex. Pollard (15 males, 6 females, CMNC; 1 male, 1 Aedeagus symmetrical in dorsal view; apex female, SMCI); Mexico, Oaxaca, 3.2 mi. s. Ocotlan, acutely angulate. Orifice covered by a median, July 20, 1974, Clark, Murray, Ashe, Schaffner (2 males, 1 female, TAMU); Mexico, Oaxaca, 1 mi. se. INSECTA MUNDI, Vol. 12, No. 3-4, September-December, 1998 199

emargination at the apex of the aedeagus (fig. 5c). Due to the costate elytra, females are likely to be confused only with M. foueicollis from which they differ in the absence of pronounced pronotal fove­ A ae. B Description. Form oval or elongate oval. Pro­ thorax and distal areas of head testaceous, orange, or pale brown; vertex and legs variable in color; elytra metallic blue; metasternum and abdomen dark brown or black. Length of male 4.5-4.9 mm; length of female 4.4-4.8 mm. Head shining, not alutaceous. Vertex and fron­ c tal tubercles variable in color, testaceous, orange, dark brown, or black; distal areas of head testa­ ceous or orange. Vertex nearly glabrous. Trans­ verse interocular sulcus deep, with each half strong­ ly curved. Interocular distance equal to 0.6 times maximum width of head across eyes. Frontal tu­ bercles mesially contiguous for a distance sub equal to width of antennomere I, separated from each other by deep sulcus, separated from mesal frontal D E carina by shallow sulci, delimited laterally by broad impression. Antennal fossae separated by a dis­ tance equal to about 1.5 times diameter of fossa. Mesal frontal carina poorly developed, weakly Figure 5. Microscelida spp., dorsal and lateral views of convex. Genal length subequal to twice diameter of aedeagi. a) M. wilcoxi, new species; b) M. alutacea, new an ommatidium. Antennae slender, extending to species; c)M. subcostata (Jacoby); d) M. wellsi, newspecies; near middle of elytra; color variable, testaceous to e) M. uiridipennis, new species. very dark brown, usually paler near base; antenno­ mere II about half as long as I or III; IV about as long Rio Hondo, July 22, 1974, Clark, Murray, Ashe, as III or V. Schaffner (2 males, 4 females, TAMU); Mexico, Pronotum 1.3 times as wide as long, 0.7 times Oaxaca, 23 km. NW. Totolapan, 28 June 1981, John as wide as elytra across humeri, widest slightly D. Pinto (1 male, 1 female, UCRV); Mexico, Oaxaca, anterior to middle; sides sinuate in dorsal view. 16.1 mi. nw. Totolapan, July 21,1974, Clark, Mur­ Lateral beads present; anterior and basal beads ray, Ashe, Schaffner (1 male, TAMU). absent. Surface minutely, very inconspicuously Comments. As indicated in the preceding punctate, shining, not alutaceous. Color orange or description, this species is variable in color. Some testaceous. specimens, including the holotype, have a dark Elytra together 1.7-1.9 times as long as broad; prothorax. Others have a pale prothorax. The legs sides nearly parallel in basal two-thirds. Punctures may be dark or pale. This species is named in honor deep, conspicuous, moderately coarse. Interspac­ of my good friend, Robert C. Mower, who has es shining, with a few minute punctures, not accompanied me on many collecting trips. alutaceous. Longitudinal costae present in female, absent or very weakly developed in male. Color Microscelida subcostata (Jacoby), dark brown or black, with metallic blue or purple new combination luster. (Figure 5c) Ventral areas of prothorax largely or entirely orange or testaceous, largely glabrous, shining; Luperus subcostatus Jacoby, 1888:597. front coxae contiguous or nearly so; front coxal Pseudoluperus subcostatus: Wilcox, 1973:457. cavities widely open behind. Mesothorax variable in color, testaceous, orange, or brown. Metaster­ Diagnosis. Males ofthis species may be distin­ num dark brown or black, glabrous or sparsely guished from other Microscelida by the small pubescent mesially, densely pubescent laterally; 200 Volume 12, No. 3-4, September-December, 1998, INSECTA MUNDI metaepisternum black or dark brown, densely ined belong to a single species. Additional investi­ pubescent. Legs variable in color, entirely testa­ gation may prove otherwise. ceous, entirely dark brown, or bicolored; terminal Jacoby had two specimens at the time he described spurs present on all tibiae of female, present on this species. As noted above, I have seen both of hind tibiae of male; basitarsi of male and female these. Each bears a "type" label. However, the slender; tarsal claws appendiculate. Abdomen major part of Jacoby's description is based on the brown, pubescent; terminal segment of male im­ male specimen with a dark vertex; the female pressed apically, with a rectangular lobe at apex of specimen with a pale vertex is listed as a variety. impression. Therefore, I conclude that the male specimen Aedeagus symmetrical in dorsal view, attenu­ (BMNH) is the holotype and the female specimen ate in distal fourth; apex with a short, medial slit (MCZC) is a paratype. or emargination. Ventral surface with a narrow, median groove. Basal spurs absent. Orifice cov­ Microscelida subglabrata (Jacoby), ered by three sclerotized processes, one median new combination and two lateral (fig. fie). (Figure 4b) Type locality. "Mexico, Iguala in Guerrero." Material examined. Mexico: Chiapas: Chia­ Luperus subglabratus Jacoby, 1888:598. pa de Corzo, 23 June 1965, Burke, Meyer, Schaffner Pseudoluperus subglabratus: Wilcox, 1973:457. (1 female, TAMU); EI Chorreadero E. ofTuxla Gtz., 13 June 1989, P. K. Lago, S. Testa (1 male, PKLC); Diagnosis. This differs from other species of Jct. Hwys 190-195, 6 June 1969, H. F. Howden (1 Microscelida in that the apex of the aedeagus is male, 2 females, CMNC; 1 male, 1 female, SMCI); rounded and not angulate (fig. 4b). EI Sumidero, km. mk. 9-12, 24 June 1990, R. Description. Form elongate oval; prothorax Turnbow (2 males, 7 females, RHTC); Sumidero much narrower than elytra. Vertex, elytra, met­ Nat. Pk., 19 June 1989, S. Testa, P. K. Lago (1 male, athorax, abdomen, and distal portion of antennae EGRC; 1 male, PKLC); Sumidero Nat. Pk., 22 June and legs brown; other areas paler, testaceous; 1989, S. Testa, P. K. Lago (1 male, 1 female, PKLC); elytra with blue luster. Length of male 3.6 mm; 17 mi. S.E. Teopisca, Rt. 24, 3-4 June 1969, H. length of female 3.7-4.5 mm. Howden (2 females, CMNC); 17 Km W. Tutl. Gtz., Head shining, not alutaceous, with vertex large­ 3300', 1-8 July 1986,J. E. Wappes (1 male, lfemale, ly dark brown, with other areas testaceous. Vertex EGRC). Guerrero: Iguala, Hoge (male holotype, 1 nearly glabrous. Transverse interocular sulcus female paratype, and 1 other female [see com­ deep; each half arcuate. Interocular distance equal ments below]; BMNH, MCZC); 11.2 mi. N. 19u1a, to 0.6 times width of head across eyes. Frontal 4300', 5 July 1987, Kovarik, Schaffner (1 female, tubercles separated from each other by deep sul­ TAMU); 2 mi. S. of Mexcala, 18 July 1957, R. B. & cus, separated from mesal frontal carina by shal­ J. M. Selander leg. (2 females, FMNH); Zopilote low sulci, separated from orbit by broad depres­ Cyn., 20-23 June 1989, Jim Cope (1 female, SMCI). sion. Distance between antennal fossae distinctly Michoacan: 24 km. wsw. C. Tancitaro, 1500 m., 1 greater than width of antennomere 1. Mesal frontal July 1947, T. H. Hubbell (1 female, SMCI; 1 male, carina poorly developed, broad. Genal length sub­ 1 female, UMMZ); Uruapan, 1610 m., 13 July 1947, equal to twice diameter of an ommatidium. Anten­ T. H. Hubbell (l male, UMMZ). Veracruz: Santeco­ nae slender, extending to near middle of elytra, mapan, 10 June 1969, H. Howden(1 female, CMNC). brown with basal portion sometimes paler; anten­ Comments. This species is somewhat vari­ nomere III 1.5 times as long as II; IV slightly longer able. Some specimens are more elongate than than III, about as long as I or V. others. Furthermore, the antennae, mesothorax, Pronotum 1.3 times as wide as long, 0.7 times metathorax, and legs vary from testaceous to dark as wide as elytra across humeri, widest near brown, or they may be bicolored. In the specimens anterior third; sides arcuate in dorsal view. Lateral from Chiapas, the aedeagus is more abruptly beads distinct; anterior and basal beads absent. narrowed near the apical fourth with the distal Disc evenly convex, lacking obvious impressions. portion more nearly parallel sided, and the short Surface nearly impunctate, shining, not aluta­ median slit at the apex is sometimes indistinct. ceous. Color testaceous. Even so, I believe that all of the specimens exam- Elytra together 1.8 times as long as broad, broadest near posterior fourth. Punctures fine, INSECTA MUNDI, Vol. 12, No. 3-4, September-December, 1998 201 sometimes inconspicuous. Interspaces shining, not for a distance subequal to width of antennomere II, alutaceous. Color brown with faint blue luster. separated from each other and from mesal frontal Ventral areas of prothorax testaceous, shin­ carina by deep sulci, separated from orbit by broad ing, largely glabrous, not alutaceous; front coxae impression. Antennal fossae separated by a dis­ nearly contiguous; front coxal cavities open be­ tance slightly greater than diameter of fossa. hind. Mesothorax testaceous. Metathorax brown, Mesal frontal carina convex. Genal length subequal pubescent. Legs testaceous with tarsi and some­ to width of distal article of maxillary palp. Anten­ times tibiae darker, brown; terminal spurs present nae slender, dark brown, paler near base; antenno­ on all tibiae of male and female; basitarsi of male mere III about 1.5 times as long as II, conspicuously and female slender; tarsal claws appendiculate. shorter than IV; V slightly shorter than IV, about Abdomen brown, pubescent; terminal segment of as long as 1. male with an impressed apical lobe. Pronotum 1.5 times as wide as long, 0.7 times Aedeagus symmetrical in dorsal view, rounded as wide as elytra across humeri, widest near at apex. Orifice covered by three sclerotized pro­ anterior third; sides sinuate in dorsal view. Lateral cesses, one median and two lateral (fig. 4b). beads well developed; anterior and basal beads Type locality. "Mexico, Acapulco in Guerre­ absent. Surface minutely, inconspicuously punc­ ro." tate, shining, not alutaceous. Color very dark Comments. The preceding treatment is based brown, almost black. on one male syntype (MCZC) and on two female Elytra together 1.8 times as long as broad; syntypes (BMNH). One of the females bears the sides nearly parallel in anterior three-fourths. following six labels: "Type" [a disk with a red Punctures coarse, separated on average by a dis­ margin], "Acapulco, Guerrero. Hoge," "Godman­ tance subequal to diameter of a puncture. Inter­ Salvin ColI., Biol. Centr.-Amer.," "Luperus subgla­ spaces shining, sparsely, minutely punctate, not bratus Jac." [a blue label], "Syntype" [a disk with alutaceous. Color dark brown with distinct purple a blue margin], and "examined by S. M. Clark #5." luster. This specimen is hereby designated as lectotype. Ventral surface dark brown. Prothorax largely The biology of this species is unknown. glabrous, shining; front coxae contiguous; front coxal cavities open behind. Metathorax largely Microscelida violacea (Jacoby), pubescent. Legs dark brown; basitarsi of male not new combination conspicuously dilated; tars omere I of hind leg (Figure 4e) about as long as II-III combined; tarsal claws appendiculate. Abdomen pubescent; terminal seg­ Scelidopsis (?) violacea Jacoby, 1892:339. ment of male with a semicircular impression in distal half, with a rectangular lobe at apex of Diagnosis. In this species, the legs and pro­ impression. thorax are dark, and the orifice of the aedeagus is Aedeagus symmetrical in dorsal view, acutely covered by a median sclerotized process and by two pointed at apex. Basal spurs absent. Orifice cov­ lateral sclerotized processes, one at each side (fig. ered by three sclerotized processes, one median 4e). These characters will enable recognition ofthis and two lateral; lateral processes meeting medially species. beyond median process (fig. 4e). Description. Form elongate; body 2.3 times Type locality. Xucumanatlan, 7000 ft., Guer­ longer than broad; prothorax much narrower than rero, Mexico. elytra. Color dark brown or black; elytra with Material examined. Mexico: Guerrero: Xu­ purple luster. Length of male 3.5 mm; length of cumanatlan, 7000 ft., July, H. H. Smith (female female 3.8 mm. lectotype, BMNH; 1 male paralectotype, BMNH; 1 Head shining, not alutaceous. Vertex black male paralectotype, MCZC); Chilpancingo, 4600 with faint green luster; frontal tubercles dark ft., July (1 male, SMCI). brown; clypeus pale brown. Vertex evenly convex, Comments. As noted above, I have examined with a few punctures and setae near each eye, three syntypes of this species. Two of these are otherwise nearly glabrous. Transverse interocular labeled "type" (female, BMNH; male, MCZC); one sulcus deep; each half strongly curved. Interocular is not (male, BMNH). I have chosen to designate distance equal to 0.6 times maximum width of head the female from the British Museum (Natural across eyes. Frontal tubercles mesially contiguous History) as lectotype. 202 Volume 12, No. 3-4, September-December, 1998, INSECTA MUNDI

Microscelida viridipennis Clark, puncture. Interspaces with minute scattered punc­ new species tures, shining, not alutaceous. Color dark brown or (Figure 5e) black, with distinct metallic green or blue luster. Ventral areas of prothorax testaceous, shin­ Diagnosis. In addition to the larger, conspic­ ing, largely glabrous, not alutaceous; front coxae uous punctures, the elytra of this species also have contiguous or nearly so; front coxal cavities open minute punctures. However, unlike M. viridis, the behind. Mesothorax testaceous; mesosternum and minute punctures are widely spaced such that the mesoepisternum largely glabrous; mesoepimeron elytra do not appear at all granulate or alutaceous. pubescent. Metathorax pubescent, usually dark Furthermore, the distal fourth of the aedeagus is brown or black, sometimes yellow. Legs testa­ gradually, attenuately narrowed to the apex, and ceous; terminal spurs present on all tibiae of male the apex is not emarginate (fig. fie). These charac­ and of female; male with tarsomere I of front leg ters distinguish M. viridipennis from other spe­ distinctly enlarged; tarsomere I of hind leg about cies of Microscelida. as long as II-III combined; tarsal claws appendic­ Description. Form elongate; prothorax nar­ ulate. Abdomen pubescent, varying from yellow to rower than elytra. Vertex and elytra dark with dark brown or black, sometimes with weak metallic metallic blue or coppery green luster; metaster­ luster; terminal segment of male impressed apical­ num and abdomen usually dark brown or black, ly, with a short, truncate lobe at apex of impres­ sometimes testaceous; pronotum often marked sion. with black; other areas testaceous. Length of male Aedeagus symmetrical in dorsal view; distal 3.1-4.5 mm; length of female 4.1-4.5 mm. fourth gradually narrowed to acute apex (fig. 5e). Head shining, not alutaceous. Vertex black Distinct basal spurs absent. Orifice covered by with weak green luster, with a few punctures and three sclerotized processes, one median and two setae in lateral areas. Transverse interocular sul­ lateral; lateral processes meeting medially distal cus distinct; each half arcuate. Interocular dis­ to median process. tance equal to 0.6 times maximum width of head Material examined. Holotype male and allo­ across eyes. Frontal tubercles and more distal type female: Mexico, Guerrero, Chilpancingo, 4000', areas of head testaceous. Frontal tubercles sepa­ 19-VII-1962, H. E. Milliron (CNCI). Paratypes: rated from each other by deep sulcus, separated Mexico, Guerrero, 2 mi. N. Cacahuamilpa, VII-19­ from mesal frontal carina by shallow sulci, separat­ 84, Carroll, Schaffner, Friedlander (1 male, TAMU); ed from orbit by broad impression. Distance be­ Mexico, Guerrero, 2 mi. N. Cacahuamilpa, July 19, tween antennal fossae subequal to one-third in­ 1984, J. B. Wooley (5 males, 7 females, TAMU); terocular distance. Mesal frontal carina poorly Mexico, Guerrero, Chilpancingo, 4000', 19-VII­ developed, convex. Genal length subequal to width 1962, H. E. Milliron (23 males, 3 females, CNCI; 2 of distal article of maxillary palp. Antennae testa­ males, SMCI); Mexico, Guerrero, 8 km W Chilpanc­ ceous or pale brown, slender, extending to near ingo, 23 July 1987, R. Turnbow (l male, EGRC); middle of elytra; antennomere II nearly half as long Mexico, Guerrero, 2.4 mi. n. Mazatlan, July 9, 1974, as I, distinctly shorter than III; IV slightly longer Clark, Murray, Ashe, Schaffner (1 male, TAMU); than III, about as long as V. Mexico, Guerrero, 38 km. W Iguala, 21 July 1987, Pronotum 1.4 times as wide as long, three­ R. Turnbow (1 male, EGRC); Mexico, Guerrero, 6.2 fourths as wide as elytra across humeri, widest mi SW Xochipala, 13-VII-1985, J. Wooley, G. Zol­ near anterior third; sides sinuate in dorsal view. nerowich, 85/056 (3 males, TAMU); Mexico, Guer­ Lateral beads well developed; anterior and basal rero, 6.2 mi. sw. Xochipala, elev. 5670 ft., July 6, beads absent. Disc evenly convex or slightly flat­ 1987, Kovarik, Schaffner (3 males, SMCI); Mexico, tened mesially. Surface minutely, inconspicuously Jalisco, 16km.n.Autlan,July31-Aug. 2,1978, Plitt punctate, shining, not alutaceous. Color testa­ & Schaffner (2 males, TAMU); Mexico, Jalisco, ceous, sometimes with a central dark area, rarely Tuxpan, 9/6 1903, J. F. McClendon (1 male, 1 almost entirely dark with only lateral margins female, MCZC); Mexico, Oaxaca, 10 mi. n. Milte­ pale. pec, July 15-16, 1971, Clark, Murray, Hart, Elytra together 1.9 times as long as broad; Schaffner (1 male, TAMU); Mexico, Oaxaca, 10 mi. sides nearly parallel in basal three-fourths. Larger n. Miltepec, July 26, 1974, Clark, Murray, Ashe, punctures distinct, separated on average by a Schaffner (1 male, TAMU); Mexico, Oaxaca, 4 mi. distance not much greater than diameter of a ne. Miltepec, July 21, 1984, Carroll, Schaffner, INSECTA MUNDI, Vol. 12, No. 3·4, September-December, 1998 203

Friedlander (2 males, 1 female, TAMU); Mexico, mere I paler, testaceous; antennomere I twice as Puebla, 4.3 mi. sw. Acatepec, July 16, 1971, Clark, long as II; III 1.5 times as long as II, distinctly Murray, Hart, Schaffner (3 males, TAMU); Mexi­ shorter than IV; IV-X subequal in length; XI longer co, Puebla, 4.4 mi SW Acatepec, July 26, 1974, than X. Clark, Murray, Ashe, Schaffner (1 male, 1 female, Pronotum 1.4 times as wide as long, three­ SMCI; 5 males, 10 females, TAMU); Mexico, Pue­ fourths as wide as elytra across humeri, widest bla, 4 mi. w. Acatepec, July 26, 1973, Mastro & near anterior fourth; sides weakly sinuate in dorsal Schaffner (1 male, TAMU); Mexico, Puebla, 30 mi. view. Margins setose. Lateral beads well devel­ se. Tehuacan, July 7,1973, Mastro & Schaffner (1 oped; anterior and basal beads absent. Surface male, 2 females, TAMU); Mexico, Puebla, 5 miles minutely, inconspicuously punctate, shining, not SWTehuacan, July 8, 1977, J. C. Schaffner (1 male, alutaceous. Color testaceous. TAMU); Mexico, Puebla, 1 mi. sw. Zapotitlan, July Elytra together 1.7 times as long as broad; 11, 1973, Mastro & Schaffner (1 male, TAMU); sides nearly parallel in anterior two-thirds. Color Mexico, Veracruz, Tierra Blanca, 500', 15-VIII­ dark brown with metallic green luster; an elongate, 1962, H. E. Milliron (10 males, CNCl); Mexico, subsutural, testaceous spot present in basal half. Zacatecas, 39.7 km. S. Juchipala, 1524 m, 88-26, Punctures fine near base and in testaceous area, 6.viii.88, R. S. Anderson, Acacia thorn scrub (4 elsewhere rather coarse and separated by a dis­ males). tance subequal to diameter of a puncture. Inter­ spaces, except near base, covered with numerous Microscelida viridis (Jacoby), closely spaced, minute punctures. new combination Ventral areas of prothorax testaceous, shin­ ing, largely glabrous, not alutaceous; front coxae Agelastica oiridis Jacoby, 1892:334. contiguous; front coxal cavities open behind. Me­ Pseudoluperus viridis: Wilcox, 1973:457. sothorax testaceous; sternum and episternum gla­ brous; epimeron pubescent. Metathorax dark Diagnosis. The pale femora and the microscu­ brown, covered with pale pubescence. Coxae, tro­ lpture on the distal half of the elytra will distin­ chanters, and femora testaceous. Tibiae testa­ guish this from other species of Microscelida. The ceous basally, elsewhere dark brown; terminal elytral interspaces appear alutaceous, but close spurs present on all tibiae of female. Tarsi brown; examination reveals that they are actually covered tarsomere I of hind leg about as long as II-III by numerous, closely spaced, minute punctures. combined; tarsal claws appendiculate. Abdomen Description. Form elongate; prothorax nar­ brown, pubescent. rower than elytra. Color largely brown; vertex and Type locality. Chilpancingo, Mexico. elytra with faint green luster; head (except ver­ Comments. I have examined a female syntype tex), prothorax, femora, and a subsutural spot on (BMNH), and this specimen is hereby designated each elytron testaceous. Length of female 4.5 mm. as lectotype. It bears the following six labels: Vertex largely black with faint blue-green lus­ "Type" [a disk with a red margin], "Chilpancingo, ter; other areas of head, including area adjacent to Guerrero. Hoge", "Godman-Salvin CoIl., BioI. Cen­ frontal tubercles, testaceous; surface shining, tr.-Amer.", "Agelastica viridis Jac." [a blue label], vaguely alutaceous, with a few long setae. Trans­ "SYNTYPE" [a disk with a blue margin], "examined verse interocular sulcus distinct; each half curved. by S. M. Clark #109." Interocular distance equal to nearly two-thirds The lectotype is the only specimen I have seen width of head across eyes. Frontal tubercles me­ that is assignable to this species. It has pale sially contiguous for a distance less than width of markings on the elytra. However, these may not be antennomere I, nearly continuous with mesal fron­ present on all specimens. Such markings were not tal carina, separated from each other by distinct mentioned in Jacoby's original description. sulcus, separated from orbit by broad, shallow I have seen a second syntype from Acapulco, impression. Distance between antennal fossae about Guerrero, Mexico (female, BMNH). However, it is 1.5 times as great as diameter of fossa. Mesal not conspecific with the lectotype. It is similar to frontal carina very poorly developed, convex. Ge­ Microscelida viridipennis but lacks the dark ver­ nal length subequal to width of distal article of tex common in that species. Accordingly, M. uiri­ maxillary palp. Antennae slender, extending to dis may not occur in Acapulco, from where it was near middle of elytra; color brown with antenno- recorded by Jacoby (1892). 204 Volume 12, No. 3-4, September-December, 1998, INSECTA MUNDI

Microscelida wellsi Clark, area near apex; middle femur testaceous with new species distal half dark brown on one side; hind femur (Figure 5d) testaceous in basal two-thirds, dark brown in distal third. Tibiae testaceous or pale brown; at least hind Diagnosis. This differs from most other spe­ tibia of male with a terminal spur. Tarsi brown; cies of Microscelida in that the distal third of the basitarsi not dilated; basitarsus of hind leg nearly hind femur is dark brown and the basal two-thirds as long as tarsomeres II-III combined; tarsal claws are testaceous. However, the most reliable way to appendiculate. Abdomen dark brown, pubescent; identify this species is through examination of the terminal segment of male with an apical, im­ aedeagus (fig. 5d). pressed, rectangular lobe. Description. Form elongate; prothorax nar­ Aedeagus symmetrical in dorsal view, with rower than elytra. Color dark brown with head, apex acute. Basal spurs absent. Orifice covered by prothorax, mesothorax, and legs at least partially three sclerotized processes, one median and two testaceous; elytra with blue luster. lateral; lateral processes medially contiguous dis­ Head shining, not alutaceous; color testaceous tal to median process (fig. 5d). except on vertex which is largely darker brown. Material examined. Mexico, Guerrero, Vertex largely glabrous, with a small, mesal im­ Chilpancingo, 4000', 19-VII-1962, H. E. Milliron pression adjacent to frontal tubercles. Transverse (male holotype, CNCl). interocular sulcus rather shallow but well-defined; Comments. Because of the position of the each half arcuate. Interocular distance equal to 0.6 legs, I am unable to see terminal spurs on the front times maximum width of head across eyes. Frontal or middle tibiae, but they may be present. If these tubercles mesially contiguous for a distance sub­ characters are found to be critical in future taxo­ equal to width of antennomere I, separated from nomic investigations, relaxation of the specimen each other, from mesal frontal carina, and from will be necessary. This species is named in honor of orbits by distinct sulci. Distance between antennal Samuel A. Wells, a coleopterist who has assisted in fossa equal to about one-third interocular dis­ many of my entomological investigations. tance. Mesal frontal carina convex, not at all acute. Genal length subequal to width of distal article of Microscelida whitingi Clark, maxillary palp. Antennae largely brown, with basal new species portion paler, testaceous; antennomere II dis­ (Figure 4c) tinctly shorter than III, about half as long as I or IV. Diagnosis. The head and legs of this species Pronotum 1.3 times as wide as long, 0.7 times are dark. Also, the distal fifth of the aedeagus is as wide as elytra across humeri, widest near very narrow and attenuate (fig. 4c). These charac­ anterior third; sides arcuate in dorsal view. Lateral ters distinguish this from other species of Mi­ beads present; anterior and basal beads absent. croscelida . Disc shallowly, inconspicuously impressed anteri­ Description. Form elongate oval; prothorax or to scutellum. Surface nearly impunctate, shin­ much narrower than elytra. Color black or dark ing, not alutaceous. Color testaceous. brown, except prothorax which may be testaceous; Elytra together 1.9 times as long broad. Punc­ elytra with violet or blue luster. Length of male 3.8 tures near anterior third distinct, separated by a mm; length of female 4.0-4.3 mm. distance about twice as great as diameter of a Head black or dark brown; vertex sometimes puncture; basal and distal punctures finer. Inter­ with green or violet luster; surface shining, not spaces with a few scattered, minute punctures, alutaceous. Vertex evenly convex, largely gla­ shining, not alutaceous. Color dark brown with brous. Transverse interocular sulcus deep; each distinct metallic blue luster. half strongly arcuate. Interocular distance equal to Ventral areas of prothorax testaceous, largely 0.6 times maximum width of head across eyes. glabrous, shining, not alutaceous; front coxae sub­ Frontal tubercles separated from each other and contiguous; front coxal cavities open behind. Me­ from mesal frontal carina by deep sulci, separated sothorax testaceous. Metathorax dark brown, large­ from orbit by broad impression. Distance between ly pubescent. Front and middle coxae testaceous; antennal fossae subequal to one-third interocular hind coxae slightly darker. Trochanters testa­ distance. Mesal frontal carina rather broad, con­ ceous. Front femur testaceous with a small dark vex. Genal length two to three times greater than INSECTA MUNDI, Vol. 12, No. 3-4, September-December, 1998 205 diameter of an ommatidium. Antennae slender, Microscelida wilcoxi Clark, extending to near middle of elytra, dark brown with new species basal few antennomeres largely paler; antenno­ (Figure 5a) mere II about half as long as I or III; IV slightly longer than III. Diagnosis. The legs, at least the front femora Pronotum 1.4 times as wide as long, 0.7 times of most specimens, are largely or entirely testa­ as wide as elytra across humeri, widest near ceous, and the interspaces of the elytra are pol­ anterior third; sides sinuate in dorsal view. Lateral ished, without numerous closely spaced, minute beads well developed; anterior and basal beads punctures. Also, the orifice of the aedeagus is absent. Surface largely impunctate, shining, not covered by a single, median, sclerotized process alutaceous. Color uniform; testaceous, black, or (fig. 5a), and the ventral surface of the aedeagus is dark brown. longitudinally grooved. These characters will to­ Elytra together 1.6 times as long as broad; gether distinguish this from other species of Mi­ sides nearly parallel in anterior three-fourths. croscelida. Punctation distinct, with most punctures separat­ Description. Form elongate oval; prothorax ed by about three times diameter of a puncture. narrower than elytra. Color largely brown; elytra Interspaces shining, not alutaceous. Color black or with weak blue luster; head, prothorax, mesotho­ dark brown, with distinct blue or violet luster. rax, and legs largely testaceous. Length of male Ventral areas dark brown or black, except 3.2-4.1 mm; length of female 3.7-4.1 mm. prothorax which may be testaceous. Prothorax Head shining, not alutaceous; color testaceous largely glabrous, shining, not alutaceous; front with posterior portion of vertex darker, brown. coxae contiguous; front coxal cavities open behind. Vertex evenly convex, with a few setae near each Mesosternum and mesoepisternum largely lack­ eye. Transverse interocular sulcus deep; each half ing pubescence; mesoepimeron sparsely pubes­ arcuate. Interocular distance equal to 0.6 times cent. Metathorax pubescent. Legs dark brown or maximum width of head across eyes. Frontal tu­ black; front tibia of male slightly broader than bercles mesially contiguous for a distance subequal normal; terminal spurs present on all tibiae of to width of antennomere II, separated from each female, present on onlyhind tibiae of male; basitar­ other and from mesal frontal carina by distinct si of male not obviously dilated, similar to those of sulci, separated from orbit by broad impression. female; hind basitarsus about as long as tarsomer­ Distance between antennal fossae about one-third es II-III combined; tarsal claws appendiculate. as great as interocular distance. Mesal frontal Abdomen brown or black, pubescent; terminal carina broad, weakly convex. Genal length sub­ segment of male with a short, impressed, subrect­ equal to width of distal article of maxillary palp. angular lobe at apex. Antennae brown, paler ventrally and at base, Aedeagus symmetrical in dorsal view; distal slender, extending to near middle of elytra; ant en­ fifth very narrow, attenuate to acute apex (fig. 4c). nomere II about half as long as I or IV; III longer Large basal spurs absent; short, acute basal tuber­ than II, shorter than IV; IV-X subequal in length; cles present. Orifice with a median sclerotized XI longer than X. covering. Pronotum 1.4 times as wide as long, three­ Material examined. Mexico, Guerrero, Tax­ fourths as wide as elytra across humeri, widest co, July, N. L. H. Krauss (male holotype, female near anterior third; sides weakly sinuate in dorsal allotype, 1 female paratype, NMNH; 1 female view. Surface minutely punctate, shining, not aluta­ paratype, SMCI). ceous. Comments. As indicated in the preceding Elytra together 1.6 times as long as broad; description, this species is variable in color. I have sides nearly parallel in anterior three-fourths. seen three specimens in which the prothorax is Surface finely punctate; interspaces shining, not entirely dark. However, the prothorax of one other alutaceous. Color brown with weak blue luster. specimen, a female, is entirely pale. Ventral areas of prothorax testaceous, largely This species is named in honor of Michael F. glabrous, shining, not alutaceous; front coxae con­ Whiting, a superb entomologist and a good friend. tiguous; front coxal cavities open behind. Mesotho­ rax testaceous. Metathorax distinctly darker than prothorax, brown, pubescent. Legs testaceous with tarsi and distal portion of tibiae sometimes slightly 206 Volume 12, No. 3-4, September-December, 1998, INSECTA MUNDI paler; terminal spurs present on all tibiae of I thank Niki Jefferies who inked figures 4 and female; basitarsus of hind leg about as long as 5 and did the complete preparation of figure 3. I tarsomeres II-III combined; tarsal claws appendic­ likewise thank Laura Torres-Miller who prepared ulate. Abdomen pubescent, brown, similar in color figures 1 and 2. to metathorax; terminal segment of male semicir­ Also, I extend special thanks to Charles A. cularly impressed at apex. Triplehorn (Ohio State University) who advised Aedeagus symmetrical in dorsal view, with during portions of this study that were completed acute apex. Basal spurs absent. Orifice with a while I was under his supervision as a graduate single, median, sclerotized covering (fig. fia). Ven­ student. Additionally, John A. Wilcox is gratefully tral surface with median, longitudinal groove. acknowledged for generously sharing his expertise Material examined. Holotype male and allo­ with the Galerucinae. type female: Mexico, Chiapas, 2 mi. W.Chiapa de Terry N. Seeno and R. Wills Flowers each Corzo, 23 June 1955, R. B. & J. M. Selander reviewed a late version of the manuscript. Their (OSUC). Paratypes: Mexico, Chiapas, 2 mi. W. suggestions led to significant improvements. Chiapa de Corzo, 23 June 1955, R. B. & J. M. Selander (4 males, 4 females, INHS; 3 males, 1 References female, SMCI); Mexico, Chiapas, 2 km S Chicoa­ sen, 18 June 1989, E. Lago, E. Zuccaro (1 male, Chapuis, F.1875. Famille LXIX. Phytophages. In Lacor­ PKLC); Mexico, Chiapas, EI Sumidero, 20 June daire, T. and F. Chapuis, Histoire naturelle des 1955, R. B. & J. M. Selander (2 males, 2 females, insectes. Genera des coleopteres ou expose method­ INHS; 2 males, 1 female, OSUC; 2 males, 1 female, ique et critique de tous les genres proposes jusq'ici SMCI); Mexico, Chiapas, 16 mi. E. Teopisca, 14 dans cet ordre d'insectes. Volume 11. La Librairie May 1969, H. Howden (5 males, 5 females, CMNC); encylopedique de Roret, Paris, 420 pp. Mexico, Oaxaca, Oaxaca, 6-24June 1968, G. Pollard Jacoby, M. 1888. Biologia Centrali-Americana, Insecta, (4 females, CMNC); Mexico, Oaxaca, Oaxaca, 16 Coleoptera, Galerucidae, vol. 6, pt. 1, pp. 585-625. June 1968, G. Pollard (l male, 1 female, CMNC; 1 Jacoby, M.1892. Biologia Centrali-Americana, Insecta, male, 1 female, SMCI); Mexico, Oaxaca, Oaxaca, Coleoptera, Supplement to Phytophaga, vol. 6, pt. 1, July 1969, G. Pollard (l male, CMNC). Suppl., pp. 313-348. Comments. Terminal spurs are present and Leng, C. W. 1920. Catalogue of the Coleoptera of Amer­ rather easily visible on the tibiae of the females. ica, North of Mexico, 470 pp. Mount Vernon, New However, although spurs are present on all tibiae York. of at least some males, I am unable to find them on Reid, C. A. M. 1992. Making full use of internal abdom­ the front and middle tibiae of others. Therefore, I inal characters. Chrysomela, Report 27: 2. conclude that this character is variable, or perhaps Reid, C. A. M. 1995. A cladistic analysis of subfamilial that the spurs are easily broken off. relationships in the Chrysomelidae sensu lata (Chry­ Although the preceding treatment reflects the someloidea), pp. 559-631. In: J. Pakaluk and S. A. color pattern of the holotype, this species is vari­ Slipinski (eds.), Biology, Phylogeny, and Classifica­ able. The dark area on the vertex may be reduced tion of Coleoptera, Papers Celebrating the 80th or entirely absent, the pronotum may have dark Birthday of Roy A. Crowson. Muzeum i Instytut lateral margins, the anterior half of the epipleuron Zoologii PAN, Warszawa. may be testaceous, and the legs vary from entirely Seeno, T. N., and J. A. Wilcox. 1982. Leafbeetle genera pale to largely dark. (Coleoptera: Chrysomelidae). Entomography 1: 1­ This species is named in honor of John A. 221. Wilcox who has generously shared his understand­ Smith, E. H. 1979. Techniques for the dissection and ing of the Galerucinae. mounting of the male (aedeagus) and female (sper­ matheca) genitalia of the Chrysomelidae (Co­ Acknowledgments leoptera). Coleopterists Bulletin 33(1): 93-103. Wilcox, J. A. 1965. A synopsis of the North American I express thanks to all individuals and institu­ Galerucinae (Coleoptera: Chrysomelidae). New York tions that have loaned specimens (see methods State Museum and Science Service, Bulletin No. section). I am especially grateful to Sharon L. 400, pp. i-iv, 1-226. Wilcox, J. A. 1973. Coleopter­ Shute of the British Museum (Natural History). orum Catalogus Supplementa, Chrysomelidae: Gal­ Had she not enabled the loan of type specimens of erucinae (editio segundo), pars 78, fasc. 3, pp. 433­ species described in the Biologia Centrali-Ameri­ 664. cana, the completion of this project would not have Wilcox, J. A. 1973. Coleopterorum Catalogus supple­ been possible. menta, Chrysomelidae: Galerucinae (editio segun­ do), pars 78, fasc. 3, pp. 433-664.