Coleoptera: Carabidae: Psydrinae)

Two new species of the genus Mecyclothorax Sharp from New Guinea (Coleoptera: Carabidae: Psydrinae)

Martin Baehr

Two new species of the psydrine genus Mecyclothorax Sharp from New Guinea are described: M. lackneri sp. n., from central Papua (former Irian Jaya), western New Guinea, and M. kubor sp. n., from central Papua New Guinea, in the eastern half of the island. The first species was collected at median altitude and is most closely related to M. jiwikae Baehr of the same region, but is distinguished by wider pronotum and even shorter elytra, and by the structure of the aedeagus. The second species was sampled at high altitude of Kubor Range and is closely related to M. sedlaceki Darlington of the nearby Bismarck Range, but is distinguished by its larger body size and the presence of five deeply impressed setiferous punctures on the elytra. A revised key to the New Guinean species of Mecyclothorax is presented. Additional records of M. sedlaceki Darlington from Mt. Wilhelm are presented. Martin Baehr, Zoologische Staatssammlung, Münchhausenstr. 21, D-81247 München, Germany. [email protected]

Introduction only known from single or few specimens. It is yet The psydrine genus Mecyclothorax Sharp, 1903 is unknown whether their recorded scattered occur- widely distributed in the area between Australia rences simply demonstrates our limited knowledge, (Moore 1984, 1992, Moore et al. 1987, Baehr 2000, or whether they mirror their truly restricted ranges. 2003), New Guinea (see below), New Caledonia Thus far, most species were recorded from central (Jeannel 1944, Deuve 1987, unpublished records), and eastern Papua (former Irian Jaya). In spite of New Zealand (Larochelle & Larivière 2001), Java much longer and better exploration, from the east- (Louwerens 1949), Borneo (Baehr & Lorenz 1999), ern half of the island (Papua New Guinea) only few and far out in the Pacific on Tahiti (Perrault 1978, species have been recorded. The extremely scattered 1992) and Hawaii (Britton 1948, Liebherr 2005, records may reflect the yet absolutely inadequate 2007, 2008). In both latter archipelagos, the genus sampling of these small beetles that probably are best has developed its highest species diversity. Most collected by sieving or Berlese extraction of litter or probably the genus is of Australian origin; it is very soil. Apparently, such sampling methods so far have widely distributed in the southern part of Australia been very rarely employed. and along the whole east coast. In view of their phy- As a conclusion, in New Guinea the genus Mecyclot- logenetic status the most plesiotypic species of the horax is yet unknown from Owen Stanley Range, genus apparently occur in southern Australia. Huon Peninsula, the western part of Papua New In New Guinea the genus was repeatedly subject of Guinea, and also from the whole western part of descriptions of new species. Revisions (Darlington, Irian Jaya (now Papua) including Snow Mountains 1962, 1971; Baehr, 1992, 1995, 1998, 2002) show and Vogelkop Peninsula. Very probably species that the mecyclothoracine fauna of New Guinea is will be discovered in most of these areas in future, by no means adequately recorded yet. because the genus is as well distributed to the east of Until now, altogether twelve species were described New Guinea as to the west. from the whole island of New Guinea, most of them

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Material and methods – Lateral margin of elytra not perceptibly sinu- The new species were detected within a sample of ate in front of the rather obtuse basal angles carabid beetles collected in western New Guinea (Baehr 1995, figs 3, 4). Papua ...... 5 and kindly sent on loan for identification from the 4. Elytra with 5 discal punctures and setae Zoölogisch Museum of the University of Amsterdam (Fig. 3); size larger, length 4.4 mm; aedea- (ZMAN) and in the unidentified carabid material gus unknown. Kubor Range at 3600 m of the National Museum of Natural History Natu- ...... kubor sp. n. ralis, Leiden (RMNH). The holotypes are located – Elytra with 2–3 discal punctures and setae; in these collections, one paratype of M. lackneri is size smaller, length < 4.0 mm; aedeagus with stored in the working collection of the author at the narrow, elongate and slightly sinuate apex Zoologische Staatssammlung, München (CBM). and with two elongate sclerites in internal Additional records of M. sedlaceki Darlington were sac (Fig. 2). Bismarck Range (Mt. Wilhelm), obtained from the collections of RMNH and of the above 3200 m . . . sedlaceki Darlington, 1971 Hungarian National Museum of Natural History, 5 Elytra with 4 discal punctures and setae; base Budapest (HNMB). of pronotum coarsely punctate; elytral striae For dissection of the male and female genitalia well impressed, crenulate, intervals convex, specimens were soaked in a wet jar for a night, then stria 7 fairly well developed (Baehr 1995, the genitalia were cleaned for a short while in hot fig. 3). Juliana Top (Gunung Mandala), east- 4% KOH. For the descriptions normal taxonomic ern central Papua, at 3500 m ...... methods have been used...... julianae Baehr, 1995 Measurements were taken using a stereo microscope – Elytra with 2 discal punctures and setae; with an ocular micrometer. Length has been meas- base of pronotum impunctate; elytral striae ured from the apex of labrum to the apex of elytra. weakly impressed, outer striae consisting of Length of pronotum was measured along midline, rows of punctures only, intervals depressed, width of base of pronotum at the extreme tips of the stria 7 scarcely indicated (Baehr 1995, fig. 4). basal angles. Measurements and ratios were obtained Eipomek-Langda area, eastern central Papua, in the same manner as in Baehr (1992, 1995, 1998, at 3500 m ...... eipomeki Baehr, 1995 2002). 6. Basal angles of pronotum almost rectangular, lateral margin near base distinctly sinuate; aedeagus see Baehr (1995: figs 5, 7; 2002: Recognition fig. 1) ...... 7 A new key to the New Guinean Mecyclothorax is given – Basal angles of pronotum obtuse, lateral mar- below. For easier identification, figures from former gin near base not or barely sinuate; aedeagus papers (Baehr, 1992, 1995, 1998, 2002) are quoted see fig. 1 and Baehr (1992: fig. 2; 1995: where necessary. Although range is not a prima facie figs 11, 12; 1998: fig. 2; 2002: fig. 2) . . . . . 9 distinguishing character, the ranges of most species 7. Size larger, length > 5 mm; surface at least seem to be so restricted that they can be used as sup- with traces of microreticulation; aedeagus port for the differentiation of the species. Therefore, see Baehr (1995: figs 5, 7). Eastern central geographic and altitudinal ranges are included in the Papua, above 3200m ...... 8 key as exactly as possible. – Size smaller, length < 4 mm; surface without traces of microreticulation; aedeagus see Revised key to the New Guinean Baehr (2002: fig. 1). Bulldog Rd., Morobe Mecyclothorax Prov., eastern central Papua New Guinea, at 2550m ...... cuccodoroi Baehr, 2002 1. Elytral striae 3 and 5 with setiferous punc- 8. Larger species, length > 5.7 mm; microre- tures. Wilhelmina Top (Gunung Trikora), ticulation on head and pronotum barely central Papua (former Irian Jaya), at 4200 m . indicated, on elytra superficial, but present ...... toxopei Darlington, 1962 on apical part of elytra; basal angles of – Only elytral stria 3 with setiferous punc- pronotum obtuse at apex, lateral margins tures ...... 2 distinct, explanate and slightly upturned; 2. Posterior lateral seta of pronotum present . . . 3 at least two or three median elytral striae – Posterior lateral seta of pronotum absent . . . 5 deeply impressed, intervals distinctly convex 3. Lateral margin of elytra distinctly sinuate (Baehr 1995: fig. 8); aedeagus see Baehr (1995: in front of the subdentiform basal angles fig. 5). Gunung Elit, eastern central Papua, (Fig. 3). Papua New Guinea ...... 4 at 3200–3300 m ...... eliti Baehr, 1995

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– Smaller species, length c. 5.3 mm; micro- central Papua, at 1900–2300 m ...... reticulation distinct, even very conspicuous ...... bilaianus Baehr, 1998 on elytra, but absent on apical part of elytra; – Apex of aedeagus elongate, not falciform, basal angles of pronotum rectangular, lateral only feebly curved up (Baehr 1992: fig. 2; margins very narrow, barely explanate and 2002: fig. 2) ...... 13 upturned; elytral striae barely impressed, 13. Elytra longer and narrower (ratio length/ intervals depressed (Baehr 1995: fig. 9); ae- width > 1.42), more oviform (Baehr 1992, deagus see Baehr (1995: fig. 7). Sape Valley, fig. 1); base of pronotum distinctly wider north of Juliana Top (Gunung Mandala), than apex (ratio base/apex > 1.18); lower sur- eastern central Papua, at 3400 m . . sapei Baehr face of aedeagus in middle straight, apex less 9. Aedeagus with short, wide, laminate api- upturned (Baehr 1992: fig. 2). Kangine area, cal plate that is bent down (Fig. 1; Baehr Baliem Valley, central Papua, at 1900 m . . . 1995: figs 10, 11); basal angles of prono- ...... riedeli Baehr, 1992 tum very obtuse, without any perceptible – Elytra shorter and wider (ratio length/width sinuosity (Fig. 3; Baehr 1995: figs 13, 14); < 1.30), more quadrate (Baehr 2002: fig. 4); either very small species (length < 3.8 mm) base of pronotum distinctly narrower than with short elytra, or larger species (length apex (ratio base/apex < 0.92); lower sur- 4.6 mm) with elongate elytra ...... 10 face of aedeagus in middle evenly concave, – Aedeagus either with short, laminate apex apex more distinctly upturned (Baehr 2002: that is markedly falciform (Baehr 1998: fig. 2). Bulldog Rd. area, Morobe Prov., fig. 2) or with elongate, less wide apex that is eastern central Papua New Guinea, at 2550– slightly curved upwards at tip (Baehr 1992: 2600 m ...... loebli Baehr, 2002 fig. 2; 2002: fig. 2;); basal angles of pronotum obtuse, though commonly with per- Mecyclothorax lackneri sp. n. ceptible sinuosity (Baehr 1992: fig. 1; 1998: Figs 1, 3, 5 fig. 1; 2002: fig. 4;), medium sized species (length 3.8–4.4 mm) ...... 12 Type material. Holotype: ?, Indonesia Papua Keca- 10. Smaller and shorter species, length matan Abenaho Pass Valley 3°51’S – 139°05’E < 3.8 mm, ratio l/w of elytra < 1.37; basal 18.-25.II.2005, 1700–2250 m dist. mtn. rainforest; angles of pronotum almost rounded off; an- sifted, leg. T. Lackner (ZMAN). Paratypes: 1?, 2/, terior transverse sulcus of pronotum weak or same data (CBM, ZMAN). absent (Fig. 3; Baehr 1995: fig. 12); left paramere elongate, less curved (Fig. 1; Baehr 1995: Diagnosis fig. 10). Central Papua ...... 11 Small, black species, characterized by absence of the – Larger and more elongate species, length posterior lateral pronotal seta, very wide pronotum > 4.5 mm, ratio l/w of elytra > 1.50; basal bearing markedly obtuse basal angles and impunc- angles of pronotum distinct, though obtuse tate base, short, very convex, somewhat oviform (Baehr 1995: fig. 12); left paramere shorter, elytra with finely punctate striae, and glossy, though more curved (Baehr 1995: fig. 11). Langda not iridescent surface. According to external mor- area, eastern central Papua, at 2300 m . . . phological characters and to shape of aedeagus the ...... langdae Baehr, 1995 species probably is most closely related to M. jiwikae 11. Pronotum narrower, ratio l/w < 1.2, with Baehr, but is distinguished from this species: 1, by narrower base, ratio width of base/width of the wider pronotum having a wider base; 2, shorter apex < 1.0; elytra longer and narrower, ra- elytra; and 3, a much shorter, differently shaped tio l/w 1.37; aedeagus slender and elongate, aedeagus bearing a wider apex and differently shaped with triangular apex (Baehr 1995: fig. 10). sclerites in the internal sac. Jiwika area, at 2300 m . . . jiwikae Baehr, 1995 – Pronotum wider, ratio l/w > 1.3, with wider Description base, ratio width of base/width of apex > 1.1; Measurements. Length: 3.55–3.75 mm; width: elytra shorter and wider, ratio l/w < 1.32; 1.60–1.75 mm. Ratios. Width prothorax/head: aedeagus shorter and more compact, with 1.32–1.34; width/length of prothorax: 1.32–1.34; rounded apex (Fig. 1). Pass Valley, between width base/apex of prothorax: 1.11–1.14; width 1700 and 2250 m ...... lackneri sp. n. elytra/prothorax: 1.37–1.38; length/width of elytra: 12. Apex of aedeagus short, laminate, markedly 1.27–1.32. falciform (Baehr 1998: fig. 2). Bilai area, Colour. Very dark piceous to almost black (when

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Table 1. Measurements and ratios of the Mecyclothorax species of New Guinea.

For better comparison of the species the measurements and ratios are compiled in the following table. For M. toxopei ratios were taken from the description, though this species is quite easily identified.

Body ratio ratio ratio width ratio ratio length width width/length base/apex width length/width (mm) pronotum/head of pronotum pronotum elytra/pronotum elytra

toxopei 4.7 1.32 1.29 0.98 1.50 1.41 sedlaceki 3.4–4.2 1.28–1.39 1.28–1.38 0.9–1.0 1.45–1.65 1.28–1.37 kubor 4.4 1.41 1.30 0.99 1.42 1.34 julianae 4.6 1.45 1.25 1.11 1.42 1.47 eipomeki 5.5 1.45 1.18 1.06 1.38 1.49 eliti 5.8–6.3 1.45–1.54 1.15–1.19 1.09–1.12 1.38–1.40 1.42–1.45 sapei 5.25 1.33 1.13 0.96 1.50 1.39 cuccodoroi 3.5–3.8 1.47–1.56 1.21–1.27 1.04–1.07 1.33–1.35 1.30–1.32 jiwikae 3.5–3.75 1.52–1.56 1.18–1.20 0.95–0.98 1.42–1.43 1.37 lackneri 3.55–3.75 1.67–1.72 1.32–1.34 1.11–1.14 1.37–1.38 1.27–1.32 langdae 4.6 1.54 1.17 1–05 1.38 1.52 bilaianus 4.0 1.54 1.17 1.01 1.45 1.42 riedeli 4.0–4.4 1.69 1.18–1.21 1.18–1.19 1.35–1.38 1.42–1.44 loebli 3.8–4.2 1.54–1.58 1.19–1.25 0.87–0.92 1.46–1.50 1.26–1.30

mature, three specimens), lateral margin and suture ened towards base, marginal channel shallow. Apex of elytra inconspicuously paler. Clypeus and labrum and base not margined. Anterior transverse sulcus reddish-piceous, palpi and antennae dark yellow, complete, well impressed, posterior transverse sulcus though antenna from 4th antennomere slightly faint. Median line well impressed, but anteriorly and darker. Legs dark yellow, tibiae with a slightly darker posteriorly abbreviated. Basal grooves short, linear, ring in middle. Lower surface dark piceous to black, rather deep. Basal area not explanate, almost on same elytral epipleura reddish. level with disk, impunctate. Anterior marginal seta Head. Rather narrow in relation to prothorax. Eyes situated slightly in front of middle, slightly removed moderately large, convex, laterally rather protruded, from lateral margin, posterior marginal seta absent. orbits small, oblique, c. ¼ of length of eye. Paraor- Surface impunctate, without any microreticulation, bital sulci deep, completely encircling the eye. Fron- very glossy. tal furrows deep, sinuate, rather elongate. Frons Elytra. Short and wide, dorsally markedly convex, without any distinct grooves. Posterior supraorbital widest diameter slightly behind middle. Humeri seta situated slightly behind posterior margin of obtusely rounded, lateral margin evenly curved. Basal eye. Clypeal suture distinct, well impressed, apex margin distinct, oblique, slightly sinuate, connected of clypeus straight. Labrum transverse, truncate, to scutellar striole. Striae except sutural stria slightly 6-setose. Mandibles moderately elongate, apically abbreviated at humerus, all striae except sutural stria suddenly curved. Mentum with distinct, apically also abbreviated at apex. Four median striae well rounded tooth. Submentum with very elongate setae. impressed, lateral striae increasingly weaker. Five Antenna rather short, almost attaining posterior median striae moderately punctate. Median five border of pronotum, median antennomeres slightly intervals distinctly convex. 7th stria impressed only < 1.5  as long as wide. Surface impunctate, with- near base, punctate. Scutellar striole fairly elon- out microreticulation, very glossy. gate, situated mediad of the outturned sutural stria. Pronotum. Large, wide, circular, considerably wider Marginal channel fairly wide. 3rd interval with 1 than long, disk fairly convex, laterally strongly and setiferous puncture in middle, situated in center of evenly curved, not excised in front of the basal interval. Punctures distinct, setae moderately elon- angles. Widest diameter about at middle or slightly gate. Near apex with a single setiferous puncture at in front. Base distinctly wider than apex. Apex end of 3rd stria. Marginal pores conspicuous, about straight or faintly concave, apical angles distinctly 13 in a row that is slightly interrupted in middle, projecting, at tip rounded off. Base almost straight, marginal setae very elongate. Intervals impunctate, only laterally faintly oblique. Basal angles widely without any traces of microreticulation. Surface very

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Mecyclothorax sedlaceki Darlington Lower surface. Largely impunctate, though Fig. 2 mesepisternum with a row of very coarse punctures. Metepisternum about as long as wide. Sternum VII in Mecyclothorax sedlaceki Darlington, 1971: 265. – Baehr male bisetose at apical margin, in female quadrisetose, 1995: 4, 5; 2002: 702. also with 4 deeply impressed punctures and setae in middle slightly removed from apical margin. This species was described from the female holo- Legs. Without striking features. Three basal tar- type sampled on Mt. Wilhelm at very high altitude someres of male anterior tarsus expanded and bise- (4250 m). A few additional specimens were now riately squamose. detected, likewise from Mt. Wilhelm, but from lower Male genitalia (Fig. 1). Genital ring short and wide, altitudes. The newly recorded specimens rather differ fairly asymmetric, with very narrow lateral arms, in their external morphological characters, e. g. body probably due to recent emergence of the specimen. size, degree of microreticulation of elytral intervals, Aedeagus short and stout, conspicuously curved and number of punctures in 3rd interval which can down towards apex, lower surface rather triangular. be asymmetrically three. The male genitalia, how- Apex markedly compressed, foliaceous, conspicu- ever, are structurally quite similar in all dissected ously spatulate, with convex tip, strongly sclerotized, males, even when some minor differences in size and slightly curved to left side and slightly concave on shape have been noted. left side. Ostium almost completely situated on right side. Internal sac with several complexly folded New records. 1?, New Guinea Mt. Wilhelm, 3750 m. sclerites. Left paramere large, though with narrow, 24.VI. (RMNH); 1?, New Guinea / NE / Mt. Wilhelm elongate apex, unisetose at apex and without setae Field Stat. 3200 m 14.IX.1968 (HNMB); 1?, New Guinea at lower margin. Right paramere narrower, likewise / NE / Mt. Wilhelm 3900 m 13–24.IX.1968 (CBM). with elongate apex, bisetose at apex and with about 8 setae at lower margin, most of these situated in For better recognition and comparison with other middle. species, measurements and ratios are given and the Female genitalia (Fig. 3). Gonocoxite 1 compact, at male aedeagus is described and figured. apical rim with a single elongate seta. Gonocoxite 2 moderately elongate, dentiform, with obtuse apex, Partial redescription with two moderately large ventro-lateral ensiform Measurements. Length: 3.4–4.2 mm; width: 1.5– setae, one elongate dorso-median ensiform seta situ- 1.95 mm. Ratios. Width prothorax/head: 1.28–1.40; ated about in middle of gonocoxite, but apparently width/length of prothorax: 1.28–1.38; width base/ without any nematiform setae near apex. Lateral apex of prothorax: 0.9–1.0; width elytra/prothorax: plate multisetose at and below rim. 1.45–1.65; length/width of elytra: 1.28–1.37. Variation. Apart from some variation in colour that Male genitalia (Fig. 2). Genital ring short and wide, is due to incomplete pigmentation, only some minor laterally convex, with very asymmetric base. Aedea- variation in relative shape of elytra was noted. gus comparatively elongate, gently curved to the right side, lower surface very gently concave. Apex Distribution straight, laterally compressed and rather narrow, Central Papua, former Irian Jaya, western New somewhat foliaceous, with obtuse tip and a fine den- Guinea. Known only from type locality. ticle on upper surface where the internal sac meets the sclerotized apex. Ostium largely situated on right Collecting circumstances side. Internal sac with several moderately sclerotized Sifted, probably from leaf litter, in montane rainfor- folds but without any denticles or heavily sclerotized est of medium altitude between 1700 and 2250 m. plates. Left paramere large, with narrow, very elongate apex, bisetose at apex and without any setae at Etymology lower margin. Right paramere shorter and stouter, The name is a patronym in honour of the collector. with much shorter apex, trisetose at apex, with dense setosity of rather elongate setae along almost Relationships the whole lower surface, a number of short setae on According to external morphological characters apical half of upper surface and some short setae on and the shape and structure of the male genitalia, apical part of external surface. M. lackneri appears most closely related to M. jiwikae Baehr, which occurs not far from the type locality of Distribution M. lackneri. Bismarck Range, central Papua New Guinea. Known so far only from Mt. Wilhelm, between 3200 and

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1 5

3 4 6

Figs 1–6. Mecyclothorax species. – 1, M. lackneri, male genitalia: aedeagus, parameres, and genital ring; 2, M. sedlaceki; ditto; 3, M. lackneri, female gonocoxites; 4, M. kubor, ditto; 5, M. lackneri habitus, length 3.55 mm; 6, M. kubor habitus, length 4.4 mm. Scales 0.25 mm (1–2), 0.1 mm. (3–4).

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4250 m. The smallest specimen recorded has been rectangular. Marginal channel rather narrow, little found at the lowest altitude. widened near basal angles, fairly deep. Apex and base not margined. Anterior transverse sulcus extremely Mecyclothorax kubor sp. n. faint, posterior transverse sulcus fairly deep, oblique. Median line incomplete and feebly impressed, ante- Fig. 4, 6 riorly and posteriorly abbreviated. Basal grooves Type material. Holotype: /, Papua New Guinea: about circular, deep. Basal area rather convex, below “Museum Leiden TERR PAPUA & N GUINEA level of disk, very coarsely punctate. Anterior mar- Kubor Range Mt. Kinkain. 3590 m. 15.VII.1963 W ginal seta situated slightly in front of middle, slightly Vink” (RMNH). removed from lateral margin, posterior marginal seta located right on basal angle. Disk impunctate, with- Diagnosis out any microreticulation, glossy. Medium sized, black, convex species, character- Elytra. Short and wide, dorsally convex, widest diam- ized by presence of the posterior lateral pronotal eter about at middle. Humeri obtusely rounded, seta, wide pronotum bearing a distinct sinuation lateral margin evenly curved. Basal margin distinct, near base and rectangular basal angles. According oblique, slightly sinuate, connected to scutellar to its body shape the species is closely related to striole. Striae except for sutural stria abbreviated at M. sedlaceki Darlington from the neighbouring base, all striae except for inner two also abbreviated Bismarck Range, but is distinguished from that spe- at apex. All striae except for 7th distinct though lit- cies by larger size and presence of five very distinct tle or not impressed, very coarsely punctate. Inter- setiferous punctures near 3rd elytral stria. vals barely convex. 7th stria barely recognizable, consisting of spaced and extremely fine punctures. Description Scutellar striole fairly elongate, situated mediad of Measurements. Length: 4.4 mm; width: 2.0 mm. the outturned sutural stria, consisting of mostly Ratios. Width prothorax/head: 1.41; width/length 4 coarse punctures. Marginal channel narrow. 3rd of prothorax: 1.30; width base/apex of prothorax: interval with 5 conspicuous setiferous punctures, 0.99; width elytra/prothorax: 1.42; length/width of all near 3rd stria, setae comparatively elongate. Near elytra: 1.34. apex with a single setiferous puncture at end of 3rd Colour. Black, palpi dark with pale apex, antennae stria. Marginal pores conspicuous, about 14 in a row reddish, legs piceous. Lower surface piceous, elytral that is barely interrupted in middle, marginal setae epipleurae reddish. very elongate. Intervals impunctate, with extremely Head. Rather narrow in relation to prothorax. Eyes superficial traces of about isodiametric microreticu- comparatively large, convex, moderately protruded, lation which laterally becomes slightly more distinct. orbits about half as long as eyes, obliquely convex. Surface glossy, though not iridescent. Posterior wings Paraorbital sulci moderately deep, slightly surpass- absent. ing the posterior margin of the eye. Frontal furrows Lower surface. Largely impunctate, though moderately deep, sinuate, rather short. Frons with- mesepisternum with a row of very coarse punctures. out any grooves. Posterior supraorbital seta situated Metepisternum about as long as wide. Abdominal slightly behind posterior margin of eye. Clypeal sterna in parts microreticulate, the posterior ones suture barely recognizable, clypeus anteriorly gen- also somewhat rugose. Sternum VII in the female tly concave. Labrum transverse, truncate, 6-setose. quadrisetose on apical margin, and with 4 deeply Mandibles moderately elongate, apically suddenly impressed punctures and setae in middle slightly curved. Mentum with distinct, apically rounded removed from apical margin. tooth. Submentum with very elongate setae. Antenna Legs. Without striking features. short, attaining but not surpassing posterior border Female genitalia (Fig. 6). Gonocoxite 1 compact, at of pronotum, pilose from mid of 3rd antennomere, apical rim with 4–5 rather stout setae. Gonocoxite median antennomeres c. 1 ⅓  as long as wide. Sur- 2 moderately elongate, dentiform, with acute apex, face impunctate, without microreticulation, glossy. with two large and stout ventrolateral ensiform setae, Pronotum. Large, wide, considerably wider than one elongate dorsomedian ensiform seta situated long, distinctly cordiform, disk fairly convex, lat- about in middle of gonocoxite, and two short api- erally evenly curved, with a short but conspicuous cal nematiform setae originating from a groove quite excision in front of the basal angles. Widest diameter removed from apex. Lateral plate multisetose at and in front of middle. Base as wide as apex. Apex almost below rim. straight, apical angles slightly projected, rounded Variation. Unknown. off. Base slightly convex throughout. Basal angles Male. Unknown

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Distribution Darlington, P.J. Jr.. 1962: The carabid beetles of New Kubor Range, central Papua New Guinea. Known Guinea. Part I. Cicindelinae, Carabinae, Harpalinae only from the type locality. through Pterostichini. – Bulletin of the Museum of comparative Zoology 126: 321–565. Collecting circumstances Darlington, P.J. Jr., 1971: The carabid beetles of New Largely unknown. Holotype sampled at 3730 m. Guinea. Part IV. General considerations; analysis and history of fauna; taxonomic supplement. – Bulletin of the Museum of comparative Zoology 142: 129–337. Etymology Deuve, T., 1987. Descriptions de deux Carabiques nou- The name refers to Kubor Range, home of this spe- veaux de Nouvelle-Calédonie et de Thailande (Cole- cies, and should be treated as a noun in apposition. optera, Caraboidea, Psydridae, Trechidae). – Revue française d’Entomologie (N. S.) 9: 143–146. Relationships Jeannel, R., 1944. Un Carabique nouveau de la Nou- Closely related to M. sedlaceki Darlington of neigh- velle-Calédonie (Coleoptera). – Revue française bouring Bismarck Range. d’Entomologie 10: 84–86. Larochelle, A. & M.-C. Larivière, 2001. Carabidae (Insecta: Coleoptera): catalogue. – Fauna of New Zea- Acknowledgements land 43: 1–284. My sincere thanks are due to Jan Krikken and Fred Liebherr, J.K., 2005. New species of Mecyclothorax (Co- van Assen of the National Museum of Natural leoptera: Carabidae, Psydrini) from Polipoli, Maui History Naturalis, Leiden, and Rob de Vos of the define an area of endemism on Haleakala Volcano. – Journal of the New York Entomological Society 113: Zoological Museum of the University of Amsterdam, 97–128. for the loan of material. Liebherr, J.K., 2007. Taxonomic revision of the Mecy- clothorax beetles (Coleoptera: Carabidae, Psydrini) of Molokai, Hawaii and recognition of areas of en- References demism on Kamakou Volcano. – Journal of the New Baehr, M., 1992. A new Mecyclothorax Sharp from New York Entomological Society 114: 179–281. Guinea (Insecta, Coleoptera, Carabidae, Psydrinae). Liebherr, J.K., 2008. Revision of Mecyclothorax Sharp of – Spixiana 15: 249–252. Hawaii Island: abundant genitalic variation in a nas- Baehr, M., 1995. The genus Mecyclothorax Sharp in New cent island radiation. – Deutsche Entomologische Guinea (Insecta, Coleoptera, Carabidae, Psydrinae). Zeitschrift: in press. – Mitteilungen der Münchner entomologischen Ge- Louwerens, C.J., 1949. Carabidae (Col.) from the Sunda sellschaft 85: 3–19. Islands. – Wissenschaftliche Ergebnisse der Sunda- Baehr, M., 1998. A further new species of the genus Expedition des Naturhistorischen Museums Basel: Mecyclothorax Sharp from western New Guinea (In- 303–325. secta, Coleoptera, Carabidae, Psydrinae). – Spixiana Moore, B.P., 1984. Taxonomic notes on some Australaian 21: 21–24. Mecyclothorax Sharp (Coleoptera: Carabidae: Psydri- Baehr, M., 2000. A new species of the genus Lestignathus nae) and description of new species. – Journal of the Erichson from Tasmania. With a note on the Tasma- Australian entomological Society 23: 161–166. nian species of Mecyclothorax Sharp (Insecta: Coleop- Moore, B.P., 1992. The Carabidae of Lord Howe Is- tera: Carabidae: Licininae, Psydrinae) – Records of the land (Coleoptera: Carabidae). – In: G.R. Noonan, South Australian Museum 33: 123–126. G.E. Ball & N. E. Stork (eds), The biogeography of Baehr, M., 2002. Two new species of the genus Mecy- ground beetles of mountains and islands. Intercept, clothorax Sharp from Papua New Guinea (Insecta, Andover: 159–173. Coleoptera, Carabidae, Psydrinae). – Revue Suisse Moore, B.P., T.A. Weir & J.E. Pyke, 1987. Rhysodidae d’Entomologie 109: 695–704. and Carabidae. – Zoological Catalogue of Australia 4: Baehr, M., 2003. Psydrine ground beetles (Coleoptera: 17–320. Carabidae: Psydrinae) excluding Amblytelini, of east- Perrault, G.G., 1978. La faune des Carabidae de Tahiti ern Queensland rainforests. – Memoirs of the Queens- II – genre Mecyclothorax (Sharp). – Nouvelle revue land Museum 49: 65–109. d’Entomologie 8: 27–36, 133–162. Baehr, M. & W. Lorenz, 1999. A revaluation of Loeffleria Perrault, G.G., 1992. Endemism and biogeography among globicollis Mandl from Borneo (Insecta, Coleoptera, Tahitian Mecyclothorax species (Coleoptera: Carabidae: Carabidae, Psydrinae). – Spixiana 22: 263–267. Psydrini). – In: G.R. Noonan, G.E. Ball & N.E. Stork Britton, E.B., 1948. A revision of the Hawaiian species (eds), The biogeography of ground beetles of moun- of Mecyclothorax (Coleoptera: Carabidae). – Occa- tains and islands. Intercept, Andover: 201–215. sional Papers of the Bishop Museum Honolulu 19: 107–166. Received: 28 September 2007 Accepted: 10 January 2008

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