The Journal of Neuroscience

EDITOR-IN-CHIEF Diana Bautista William C. Stacey Dora E. Angelaki Heinz Beck Raymond Swanson Paul Buckmaster Donna M. Wilcock SENIOR EDITORS Anis Contractor Dianna Willis The Journal of Neuroscience (ISSN-0270-6474) is the offi- Mark Bothwell Liset M. de la Prida cial journal of the Society for Neuroscience, 1121 14th Valina Dawson Sulayman Dib-Hajj Systems/Circuits Street, NW, Suite 1010, Washington, DC 20005. It is pub- Cheryl Dreyfus Greg DeAngelis Bruno Averbeck lished weekly by the Society, fifty weeks a year, one vol- Nancy Ip James Fawcett Lisa Goodrich Tallie Z. Baram ume a year. Sabine Kastner H. Tad Blair Jeanne Nerbonne Helmut Haas Subscriptions are on a volume basis beginning with the Artur Kania James Bisley David Perkel Kamran Khodakhah Matteo Carandini January 7 issue. The Volume 35 (2015) Institutional sub- Matthew Rushworth Carl Lupica Manuel Castro-Alamancos scription rates are available online at http://www. Yavin Shaham Olivier Manzoni Laura Colgin jneurosci.org/subscriptions/cost.dtl. Single copies for the REVIEWING EDITORS Lori McMahon Bevil Conway U.S. are $85, $105 elsewhere. Institutional subscribers Lin Mei J. Douglas Crawford Jose Manuel Alonso should direct subscription orders, back issue requests, Suzanne Moenter Alain Destexhe Christopher I. Baker Maiken Nedergaard claims, and changes of address to The Journal of Neurosci- Tracy L. Bale Abdeljabbar El Manira ence, Subscription Services, 1121 14th St., NW, Suite Bernd Nilius Andre Fenton Jeffrey Diamond Linda Overstreet-Wadiche 1010, Washington, DC 20005, [email protected]. Alyson Fournier Vincent Ferrera Matthew Rasband Paul Frankland Claims will be honored if received within three (3) Fabrizio Gabbiani Christian Rosenmund Vittorio Gallo Patricia Gaspar months of the month of issue. A maximum of three (3) Jürgen Sandkühler single-issue claims will be honored per volume per sub- Michael Gold Laura Schrader Asif Ghazanfar Jay A. Gottfried Mark Shapiro Michael Hasselmo scriber. Society for Neuroscience members interested in Ege T. Kavalali Joshua Singer Andrew Huberman subscribing to The Journal of Neuroscience at a reduced rate Christoph Kayser Wayne S. Sossin Christoph Kayser should contact the Society’s Membership Department Paul J. Kenny Henrique Van Gersdorff Andrew King (phone 202-962-4000, [email protected]). Society Eric Klann Bettina Winckler Richard Krauzlis members should also direct back issue requests to James Knierim Danny G. Winder Rohini Kuner Gary E. Landreth Mark Laubach [email protected]. Daeyeol Lee Development/Plasticity/ Mary Kay Lobo Joseph LoTurco Repair Advertising inquiries should be addressed to Marci Julio Martinez-Trujillo Leach, The Society for Neuroscience, 1121 14th St., NW, Mark P. Mattson Heather Cameron Lisa Monteggia Jonah Chan Suite 1010, Washington, DC 20005, phone 202-962-4000, Diane Merry Gabe Murphy Farzan Nadim Jean-François Cloutier [email protected]. Anna Christina Nobre Donna Fekete Guy Orban Sam Pleasure Roman Giger Richard Palmiter Reprints of a single copy of an individual article are avail- Dan H. Sanes Robert Hevner Anitha Pasupathy able from Infotrieve at http://www.infotrieve.com. Helen E. Scharfman Jenny Hsieh Gina R. Poe Geoffrey Schoenbaum Matthew Kelley Gregg Recanzone Photocopies: The Journal of Neuroscience is registered Charles Schroeder Marlies Knipper Margaret E. Rice with the Copyright Clearance Center, 222 Rosewood Ruth S. Slack Dimitri Krainc Dorit Ron Drive, Danvers, MA 01923. Authorization to photocopy Qiufu Ma Hans Straka Jane M. Sullivan items for the internal or personal use of specific clients is Ivan Toni Keith Murai Alessandro Treves Catherine S. Woolley Lucas Pozzo-Miller Doris Tsao granted by the Society for Neuroscience provided that the Edward S. Ruthazer Matthjis van der Meer copier pay to the Center the $15.00 copy fee stated in the ASSOCIATE EDITORS Carlo Sala Lori Zeltser code on the first page of each article. Special requests, Behavioral/Cognitive Jorg B. Schulz such as for general distribution, resale, advertising, and Michael Sofroniew FEATURES EDITOR Mark Baxter Carla Taveggia Teresa Esch promotional purposes or for creating new works, Michael Beauchamp Terrence Town should be directed to Journal Permissions, Society for Leonardo Cohen Flora Vaccarino EDITORIAL STAFF Neuroscience, 1121 14th St., NW, Suite 1010, Wash- Lila Davachi Teresa L. Wood Kari Black ington, DC 20005, [email protected]. Floris de Lange Lawrence Wrabetz Zachary Dourafei Ralph Dileone Fengquan Zhou Michael Eppenberger Postmaster: Send address changes to The Journal of Stan Floresco Chanelle Grannum Neuroscience, Society for Neuroscience Membership Alex Fornito Neurobiology of Disease Justin Martinez Department, 1121 14th St., NW, Suite 1010, Washington, Karl Friston Josef Anrather Matthew McCabe Jack Gallant DC 20005. Periodicals postage paid at Washington, DC, and Douglas Bayliss Jacqueline Perry additional mailing offices. Canada Post International Publi- Biyu He Monica Carson Suzanne Rosenzweig Bruce Hope Alain Chedotal cations Mail (Canadian Distribution) Sales Agreement No. Sheena A. Josselyn William Dauer 0895369. The Journal of Neuroscience is indexed by BIOSIS, Donald Katz Patrick Dougherty Brain Information Service, Chemical Abstracts, Current Ifat Levy Joseph El Khoury Contents, /Excerpta Medica, /Med- Gavin McNally David Eidelberg Micah Murray line, Index to Scientific Reviews, PubMed, PubMed Central, Frank M. Faraci ⅜ϱ Uta Noppeney Jonathan Godbout and PsycINFO. The Journal of Neuroscience is John O’Doherty Todd Golde printed on acid-free paper that meets the minimum Luiz Pessoa Christian Haass requirements of ANSI Standard Z39.48-1984 (Perma- Paul Phillips Giles Hardingham nence of Paper). Copyright © 2015 Society for Neuro- Cathy Price Markus Heilig science. All rights reserved. Greg Quirk Uwe Heinemann Antonio Rangel Eric Huang The ideas and opinions expressed in The Journal of Neuroscience Matthew R. Roesch Frances Jensen do not necessarily reflect those of the Society for Neuroscience or Gordon Shulman Robert Kalb the Editors of The Journal of Neuroscience. Publication of an ad- Harwig Siebner Michael S. Levine vertisement or other product mention in The Journal of Neurosci- Garret Stuber Andrew Lieberman ence should not be construed as an endorsement of the Gregor Thut Timothy H. Murphy manufacturer’s claims. The Society for Neuroscience does not Philippe N. Tobler David Pleasure assume any responsibility for any injury and/or damage to per- Ulf Ziemann sons or property arising from or related to any use of any material Erik D. Roberson contained in The Journal of Neuroscience. Krish Sathian Cellular/Molecular Barry Setlow Anne Baron-van Tara Spires-Jones