INSECTA MUNDI, Vol. 9, No. 1-2, March - June, 1995 17

Additions to the checkered fauna ofBelize with the description ofa new species (Coleoptera: ) and a nomenclatural change

Jacques Rifkind 11322 Camarillo Sf. #3114 North Hollywood, CA 91602 USA

Abstract: New information on the distribution and ecology of Cleridae in , Central America is presented. Enoclerus (E.) gumae, ne.. slleeies, is deseribed frem Ctl) 0 Distriet, Belize and Cyma/otiera pal1idipennis Cltewolat 1843 is Illaeed as a junior synon)m of C. plolixa (Klug 1842).

Introduction Until the early part of this century, the only Tbe clerid fauna ofBelize (formerlyBritish Hon- means of transportation from Belize Cit.y, on t.be duras) is very poorly known. In the most recent coast, to Cayo (now San Ignacio), near the western checklist for Central America, Barr (1975) specifi border '/lith , was by a small, manuallY cally mentions only Cymatodera pallidipennis propelled, hand-carved boat -the "pitpan"- up ChevlOlat 1843 and Enoclel us x-album (Gorham the Belize River (Stephens 1841, HOI widr & Lyon 1882) as occurring in Belize, while recording 86 1990[?]). Rapids complicated the journey, which species from neighboring Guatemala. In addition could take from "8-21 days, traveling both day and to the speCIes lIsted by Barr, Gorham (1883), III the nIght" (HorWIch & Lyon 19911 [1], p 47) We know Biologia Centrali-Americana, records Cregya that F. Blancaneaux, the French chicle and ma- quadrisignata (Spinola 1844) (as Pelonium hogany contractor and amateur naturalist who quadrisignatum Spinola 1844) and the two cosmo- collected all the Belizean clerids examined by politan korynetines, Nec70bia 7u{icollis (Fabr. Gorham, didvisit Cayo, hisseveralcollectingrecords 1775) and N rn,fipes (deGeer 1775), from Belize ofScarabaeidae from there are listed in the B7:ologfa Finally, Ekis (1977) published several Belizean volume devoted to that family (Bates 1888,1889). records of Perilypus distinctus (Chevrolat 1874). All of Blancaneaux's clerid records, hmvever, are Thus, a total of only six species of clerids is previ- limited to the lowland region of the Rio Sarstoon, ously recOIded from Belize, a country encompass­ , which fOIms the southernmost ing a diverse range of habitats and whose neigh- border of Belize with Guatemala oors, MeXiCO and Guatemala, are home to hun­ Although there have been several substantIal dreds of elend species botanicalandherpetolOgical investigatiOnsinBelize The reason for our poor knowledge of Belize's (1923 Marshall Field Expedition to Central Cleridae relates to the lack of attention paid that America;1936 Michigan Carnegie Botanical Expe­ country by 19th and 20th century beetle collectors. dition to British Honduras; Cambridge Expedition This is probably tire result of the difficulty of inte- to British Honduras, 1959-60, Henderson and nor travel in Belize Selander and Vaurie (1962) Hoevers ] 975 ), and workers in other disciplines found only eight collecting localities in Belize listed such as mammology (Kirkpatrick and Cartwright in the whole Insecta portion ofthe Biologia. Com 1975 and references therein) and ornithology pare this with Guatemala from which they totaled (Russell 1964) have focused their efforts there, 128 separate collecting locations, and we can see recent research on the coleopteran fauna has some ofthe reason for the dearthofBelizeanclerids apparently been limited to a few species descrip- reported in scientific literature. tions (e.g. Chemsak 1983; ..Tohnson 1984). I hope that this paper, which presents the results of a 18 Volume 9, nos. 1-2, March-June, 1995, INSECTA MUNDI brief, geographically limited collecting trip, will Classification and sequence of taxa accord with inspire others to focus their attention on this beau- Barr (1975) tlful, relatlvely unspOIled country whIch no doubt possesses many unknown sources ofentomological CLERlNAE,Tillini. interest. In the follow ing discussion of clerid distribu- Cymutodel u pIolixu (KIng 1842) tion. the reader will not fail to notice that repeated references to the confused and uncertain status of Tillus prolixus Klug 1842:272. ~ny taxa b:com: a rathe~ an~oYing ieit~ott Cymatodera prolixa (Klug). Chevrolat 1876.11, Schenkling IS uncertamty IS primar y i e resu tot e 1903:21, 1906:251; Wolcott 1927:17; CorporaaI1950:51. unavailability ofsufficient material for study. Fur CJ'matodcrapallidipgn.nis Chevrolat 1843'8; Gorham 1882'132; ther intensive collecting in Belize and neighboring COrporaa1I950:51. New synonym. areas is clearly needed in order to solve these ComparIson of these speCIes' types by Barr eize con ams dIverse habItats, rangmg from (pers. comm.) shows them to be conspecific. I have mangrove swamp to cohune palm forest and "rain studied a transparency of Chevrolat's pale colored forest." Most of the discussed in this paper ty pe specimenofC. pallidipennis and conclude that were taken in three types of plant community. it is well within the observed variability of the limestone associated closed canopy broadleaved broadly distributed C. prolixa. hardwood forest (known as "High Ridge" in Belize); This species was previously recorded in Belize broken canopy mixed broadleaved forest ("Broken (as C. pallidipennis) from the Rio Sarstoon, Toledo RIdge"); and the upland pme--pimenta palm District. The following represents a new district fore st which grows in granitic soils and is locally record. One specimen, Belize, , 5.1 referred to as "Mountain Pine Ridge." The mi N Hattieville, 200m, June 21, 1992, swamp

"lower montane rain forest" and "evergreen sea- sonal foresr' of Beard (1944). It should be noted that the distribution ofthese forest types in Belize can be dependent as much on edaphic conditions as on elevation (Brokaw & Lloy d-E"ans 1987). Phyto- CI.ERINAE, CJerini: geographic terminology employed here is taken from Bartlett (1936), Lunden (19411), Beard (1944), Priocera clavipes Gorham 1882 and Kricher' (1989) Nevertheless, even a brief exammation of the literature will show that there New country record. One specimen, Belize, Cayo is no consensus on the use of terms; I believe that District, off of ChiquibUl Rd., 8 mi S Georgeville, each ofthese authors' attempts to redescribe what 420 m, broadleaved hardwood forest, June 21-24, he has observed reflects the subtle complexity of 1992, beating tree branch. Known pre"iously nom these forests and the general refusal of tropical Guatemala, Costa Rica, and Panama. nature to "SIt still" enough fOr us to characterize any part of it with a simple label. The purpose ofthis paper is to describe anewly Priocera stictica Gorham 1882 discovered clerid species, to propose a nomencla- tural change, and to present new information on tbe distribution of cJerids in Belize IstrIct,an IgnaclO,an gnaclO ote m, Abbreviations are as follows: California Acad- ,9 emy of Sciences, San Francisco, CA, (CASC); FIeld June 26, 1992, riparian hardwood forest, MV and Museum ofNatural History, Chicago, IL, (FMNH); fluorescent lights. This species is broadly distrib- The Natural History Museum, London, England, (BMNH); and Jacques RiiKind collection, Nor th Hollywood, CA, (JNRC). Unless otherwise noted, all specimens listed below as representing new records were collected by J. Rifkind and P. A. Gum and are housed in JNRC. INSECTA MUNDI. Vol. 9. No. 1-2. March - June. 1995 19

Perilypusdistinctus(Chevrolat 1874) (GOIham 1882). In these Belizean specimens the posterior pronotal slope abruptly and deeply falls Ekis (1977) examined several specimens ofthis awayfrom a relativelyconvexpronotaldIsk, and the species flOm Corozal and Belize DistIicts and "M- pronot~lSitElS arEl arcuately constrictEld to a 'Nell set tee Dist" (his quotation marks). The latter is most certainly an abbreViation for "Manatee Districf' Vllhich rElwrs either to the area arollnd the coastal pronotalcharactersdiscussedherearequite different site of Manatee in southern Belize District, or to from thatofanyotherAphelocerus I have examined, Belize Disbictitself. Allthe PerilYPu8 Spinola 1841 and I believe that this species will prove to be new specimens listed below were taken on low vegeta­ Members of the mttdus "group" greatly re­ tion in shaded forest areas near clearings and semble and are probably invohed in mimiery roadcuts; similar experience collecting Perilypus complexes that include weevils of the genera ill Mexico suggests that these low, relatively cool ltfy1lnex StUlm 1826 and Lissodeles Champion areas are a preferred microhabitatfor atleast some 1906 as well as members ofother beetle families as of the species in the genus. The following is a new district record. One specimen, Belize, Cayo District, Vaca Plateau, Cbiquibul Rd ,between and the Guacamallo possible models fOI this mimicly, Ianging from ants Bridge, 620-650 m, June 28, 1992, closed canopy to spiders (O'Brienpers. comm., Barr, pers. comm.), broadleaved hardwood forest, beating low shrubs; there has been no compelling anecdotal or experi­ one specimen, Belize, Cayo District, Jlaca Plateau, m~mtal evidence to support any particular hypoth- Caracol, 680 m, June 28, 1992, closed canopy eSls. broadleaved hardwood forest, sweeping shrubs. Species "B" has the pronotum densely punc This species has been recorded from Mexico and tate, the elytra shallowly, densely punctate and G'ultemala. weakly, tIansveIsely I ugose. Each elytIOn bears a small, dense patch ofwbite setae justin front ofthe mIddle and a hne of whIte setae along the suture Perilypus quadrilineatus (Chevrolat 1843) extending from baSEl to mid elytra. The posterior pronotal slope is much shallower than in species New country record. Four specimens, Belize, "A" and the basal pIonotal collal is nanowel and less distinctly set off from the pronotum. Ca~o District Mountain Pine Ridge Pine Rid~e chi.o Line atLittle Vaqueros Cleek,ij25 m, June~ Thefollowing represent new countryrecords & 28, 1992, hroken canopy hroadleaved hardwood forest, beating and sweeping low vegetation in p eocerus sp. "A." Two examples, Behze, slash area; one specimEln, Belize, Cayo District, Cayo District, Mountain Pine Ridge, Pine Ridge , 340 m, June 29, 1992, broadleaved Chito Line atLittleVaqueros Creek, 525 m, June 26 haldwood fOlest, sweeping low shlub. Known also & 28, 1992, blOken canopy blOadleaved haIdwood from Mexico and Guatemala. forest, beating and sweeping in slash area. Aphelocerzts sp "B" One specimen, Belize, Cayo District, San Ignacio, San Ignacio HotEll, 290 m, June 26, 1992, riparian hardwood forest, MY Aphelocerus Kirsch 1870 and fluorescent lights. Tbis genus, broadly distributed in Central AmerIca but preVIOusly unrecorded from BelIze, IS Enoclerus (E.) fugitivus Wolcott 1927 in a chaotic state taxonomically. Neither generic nor inters ecific definin characters have been Panama, also rangesintoMexico (Chiapas; Quintana

from theGuatemalanborder, indicatesthelikelihood ofits presence inthe Peten region ofthat country as well. WolcottdescribedE. (E.)fugitivus from a black coloredspecimen; anothercolormorph, withthehead, 20 Volume 9, nos. 1-2, March-June, 1995, INSECTA MUNDI pIOnotum, basal halfofthe ely tra, legs, and ventral

notedthatthe reddishcolorvarietyofE. (E.) fugitivus

Cayo District, Mountain Pine Ridge, Pine Ridge

Vaca Plateau, Caracol, 680 m, June 28,1992, closed shrubs.

gumae new speCieS

bust, elytra flattened above. Color: integument black except antennae, reddish; head with longitudinal, triangular red narrowbandconsistingofparallel, unevenly spaced, subbasal black marking; each elytron with set of 3 m 1 just attaining marginal bead. Head: slightlY narrower across eyes than maxi- mum width of pronotum; surface shining, moder- Figure 1. Enoclerus (E.) gumae Ritkind. Dorsal habitus. ately densely punctate, punctmes moderate sized, cially on vertex, composed of short, fine, pale, suberect setae and fewer erect darker setae; front. feebly bi-impressed; clypeus with front margin transverse; eyes medium sized, depth of emargin ation slightly shallower than length of antennal scape; antennal club (Fig. 2) large, consisting of 3 n n m m at apical 1/3, apex acute; scape and funicular cute urn with finer and more sparsely arranged setae. atapex. Pronotum:narrowerthanelytraatbase(35:42), Elytra: little more than 1.5X as long as wide, slightlybroaderthanlong(70:65); surfaceunevenand widest just behind middle, disk moderately flat- ratherfinely, shallowly, butsubconfluently, calculose- INSECTA MUNDI, Vol. 9, No. 1-2, March - June, 1995 21

2

Figure 2 Enoclerus (E) gumae Rjfkjnd Antenna Variation. The paratype specimen is 5.80 mm in length, has the reddish frontal spot latitudinally, tened; SUIface velY coalsely, densely, puncturedand ~:~:~~a~h~:rJo~~~~1:nr~~~~ei~~::~It~::,~~: rugose,somewhatmorefinelypunctateandsmoother on eithersideofelytral suture, ativorymarkings, and elytra bear a few indistinct longitudinal carinae at apicall/5; ivory elytral markings with scattered which become obsolete at apicall/4. The female is puncturescontrastmgly darkened; vestiture conSIst­ unknown. ing of dense, inconspicuous patches of short, fine, suberect, black and white setae (correspondingly DIagnoSIS. The new speCIes possesses tIbIal cari­ arranged on black and ivory colored areas of the nae and a compact antennal club, placing it in the elytra), scatteredlonger erectblack setae, and more subgenus Enoclerus Barr 1976. Because of its salient coarse, suberect, white setae, latter concen- unique combination of coloration, elytral patteln- trated at base, along suture, and then expanded ing, and integumental sculpturing, Enoclerus (E.) obliquely towards sidestocover apicall/5 of elytra; subbasaltUIllesceneeBBlightly ele vated, sidesslightly constricted behind moderately prominent humeri, .. sag£ttanus IS 6, w C I ers m then graduallyexpanded tobehind middle wherethey elytral pattern, in having the surface of the elytra broadlycurve toseparatelyrounded apices. much smoother, with finer, shallower punctation, Legs: stout;profemora noticeablYmoreexpanded and is overall a more robust . Enoclelus (E.) thanmeso andmetafemora;surfaceshining, sparsely nigromaculatus (Chevrolat ]843), which likely oc- punctate on first pair of legs, densely punctate on curs m BelIze, IS SImIlar m SIze and general confIgu­ second and third pairs, moderately covered with ration to E. (E.) gumae, but it too has a completely intermixedshort, suberect, andlong, erect,palesetae different arrangement of markings on the elytra. and occasional long, erect, black setae; protarsal pulvilli broader than those of meso and metatarsi. Relationships. The placement of any new species Tibialcarinae present. within EnQclerns is problematic since, as Barr Mesosternum. densely, shallowly, punctate and asperous, moderately covered with recumbent pale setae; anterior median process conspicuous, projecting well forward of prothoracic foraminal tematically studied. For example, many species flange, rounded, and with longitudinal ovoid de- described in Enoclel us may indeed belong in pression at middle; posterior median process not Caestron Spinola 1844 and the generic distinctness elevated. Metasternum. densely, finely, and shallowly punctate and rugulose, covered with recumbent pale setae, especiallyatsides; basal1i2with narrow, ships, E. (E.) gumae belongs to the (most likely) longitudinal area at midline impunctate and gla­ polyphyletic group ofEnoclerus species which lack brous; anteriormedian process not elevated. strongly produced meso and/or metasternal projec- Abdomen:surface shining, moderatelydensely, tions and distinctly elevated elytral subbasal tu- coarsely punctate, moderately covered with erect mescences. At thIS pomt m our understandmg of 22 Volume 9, nos. 1-2, March-June, 1995, INSECTA MUNDI

EPIPHLOEINAE

PhlOgistosternus near erythrocephalus (Gorham 1882)

head and pronotal anterior margin of this species arc extremelYvariable in othercharacteristicssuch Etymology.Itiswithgreatdelightthat I name this as size and elytral color and patterning. They may IepIesent eitheI a single poly typic species OI mOle species for Patricia Ann Gum, my linda compaiiera ~han one speciefh Further work is needed to un- de campo y "ogUt, co-coHeeWI of the ty pe. angle these relatIOnships. In any case, the mate- DistributIon. Known only from the type localIty. rial collected In Belize displays some of thiS varia- tion, with four darker specimens ranging from 3.50 mm to 5.35 mm in length and one paler, some,,"vhat broader~ecimen"dwarfing" the others at 7.0 mm Enoclerus (EJ pilatei (Chevrolat 1874) in length:

New country record. One specimen, Belize, Cayo New country record. One specimen, Belize, Cayo District, Mountain Pine Ridge, Pine Ridge Cbito District, Chaa Creek, 340 m, June 29, 1992, Line atLittleVaqueros Creek, 525 m, June 26 & 28, broadleaved hardwood forest, on tree branch; four 1992, broken canopy broadleaved hardwood forest. specimens, Belize, Cayo District, Mountain Pine beating and sweeping low vegetation in slash area. Rid~e, Cooma Cairn Rd. 0-1.5 mi NW Jct. Rd. to ThiS species was extremelY wary and fast; although Hid en Valley Falls, 812-900 m, June 24, 1992, several beetles were seen, only one was success- beatingdeadPin II scaribaea Mor Ph logistosternlls fully netted. Previously recorded from Mexico and erythrocephalus is also known from Mexico, Gua- Guatemala. temala, Honduras, and Panama.

PHYLLOBAENINAE khnea near marginella (Klug 1842)

Isohydnocer a clyptocel ina (Gorham 1883) The systematics of this genus ale in need of revision; many ofthe described species arc variable As varIOus workers have pomted out (Chapm m coloratIOn and the status ofsome IS questIOnable. 1922, Wolcott 1927), this species, and possibly some New country record for the genus. One speci- others now described in Phyllobaenus Dejean 1837 men, Belize, Cayo District, Mountain Pine Ridge, and lsohvdnocera Chapin1917, should be included Pine Ridge Cbito Line at Little Vaqueros Creek, in a new genus. Given our current insufficient 525 m, June 26 & 28. 1992, broken canopy understanding of the speciose and variable genus broadleaved hardwood forest, beating and sweep­ Phyllobaenlls, an adequate definition of this new ing in slash area taxon IS not yet pOSSible. The follo',ving data represent a new eountry KORYNETINAE, Enopliini. record. One specimen, Belize, Cayo District, offof Chiquibul Rd., 8 mi S Georgeville, 420 m, Neorthopleura purpurea (Gorham 1883) broadleavedhardwoodforest,June 21-24, 1992, beat- mg and sweepmg; one speCimen, BelIze, Cayo DIS­ trict, "La Vega", 1-2 mi S Mountain Equestrian Until now, this species has been known only Trails, offof Chiquibul Rd., June 23, 1992, beating. from the type series: nine specimens collected at Chacoj, AltaVerapaz, GuatemalabyChampion. Barr Isohyd1tocera cryptocerina has been previously re corded from Guatemala and Costa Rica. (1976a) reported thatspecimenshe examinedhad tire dorsal coloration rusty rather than the "rubra " of INSECTA MUNDI, Vol. 9, No. 1-2, March - June, 1995 23

GOlham's(1882)originaldesCIiption.Ban speculated Acknowledgments t~~ di~cre~ancy the might be the result of a color I wish to thank D. Kempster and M. Kerley (BMNH) for the loan of type material. I greatly benefited from discussions on mimicry with HA Hespenheide (University of California, Los Ange­ specimen of the lycid Plateros ochraceus Gorham les), C. w. O'Erien (Florida A&M University, 1880, remarkably similarinsize, shape, and color to Tallahassee), and VI. F. Barr (Moscow, ID). Richard T lv • pUTpurea and possibly its model, was collected S. Miller (Montana State University, Bozeman) ~::;~~nt~~l~e::~~:,tci~~~~~~~~s~l~l:~~;:r~~ kindly provided identification ofthe lycid specimen and information on its distribution. As usual, W. F. comm.). Barr gracIousLy made available specimens in his care and information from his bountiful cranial New country record. One specimen, Belize, Cayo storehouse of cleridology; I thank him and C. L. District off of Chi uibul Rd. 8 mi S Geor eville thoughtful comments on this manuscript. I am grateful to Chief Forestry Officer E Green (, Belize), who extended permission to collect in the Mountain Pine Ridge and to Mark to differ from beetles in the type population. Bevis ofMountain Equestrian Trails (Cayo, Belize) for able and cheerful guidance on horse trail and jeep track. I am also indebted to J. D. Beierl (North Chariessa vestita (Chevrolat 1835) Hollywood, CA) for providing the excellent habitus drawmg Finally, I thank Patricia A Gum (North Barr (pers. comm.) believes that this species, Hollywood, CA) for assistance in the field and with its metallic bIlle, apically expanded elyfra and constant encouragement on the "bug stuff." red legs, is probably mimicking one or more species ofchemically protected chrysomelids. Itis interest­ ing to note then, that two specimens of this insect References were collected as they lay fully exposed on a fallen, Barr, W. F. 1962. A key to the genera and a classifica­ sunlit tree trunk where they would have been tion of the North American Cleridae (Coleoptera). conspicuous to any potential predator. Further­ Coleopts. Bull. 16:121-127. more, one insect, '!pon bemg pIcked up, exuded a Barr, W. F. 1975. Family Clerid~. V~l. 4. Family 7i. [ reddish fluid from its underside, much like many of North and Central America and the West Indies chrysomelids are observed to do. Ifthis fluid is not Flora and Fauna Publications, Gainesville, Florida. noxious, then its secretion could represent a behav- Barr, W E' ] 9763 Taxonomy of the new clerid genus ioral component added to mimetic shape and col­ Neorthopleura (Coleoptera). Melanderia, 24: 1-14. oration in patterning after the aposematic model Barr, W E' 19'16b Descriptions and taxonomic notes of insect; on the other hand, if the fluid is indeed Enoclerus and some allied genera (Coleoptera: noxious to predators, C. vestita may be involVed in Cleridae) Melanderia,24·17.35 a mutually reinforcing MUllerian mimicry relation Bartlett,H. H. 1936. A methodfor field workin tropical ship with its putative chrysomelid model, rather American phytogeography based upon a botanical reconnaissance in Garts of British Honduras and than being a simple Batesian mimic as previously the Peten forest of uatemala. [palt 1, pp. 1-25, 14 suspected. Chariessa vestita has been recorded ~IS.~ In: Botan; of the Maya area: miscellaneous from the (ImtedStates (Texas), Mexico, Guatemala, atyls. Carnegi lnst. 'Nash~, 'Nashington, D. C. Nicaragua, and Costa Rica. Bates,H.W. 1888. BiologiaCentrali-Americana, Insecta, Coleoptera, Melolonthidae, Rutelidae, Dynastidae, New countIy I CCOI d. Three specimens, Belize, 2(2):161 336, pIs. Cayo District, Mountain Pine Ridge, Pine Ridge Bates,H.W. 1889. BiologiaCentrali-Americana, Insecta, ChitoLine atLittle Vaqueros Creek, 525 m, June 26 Coleoptera, Dynastidae, Cetoniidae, suppl. 2(2):337- & 28, 1992, broken canopy broadleaved hardwood 416, pIs. forest, onfall@ntree tr'mkand b@atinginslash area Bea~~~to~ ~:~:~~~::;egetationintroPicalAmerica 24 Volume 9, nos. 1-2, March-June, 1995, INSECTA MUNDI

Chemsak, J. A. 1983. Anew CentIalAmerican species KricheI,J. C. 1989. A neotropicalcompanion. Princeton of Derancistrus (Coleoptera: Cerambycidae). Proc. University Press, Princeton. 436 pp. Entomol. Soc. 'Hash. 85(4):714 716. Lundell, C. L. 1940. The 1936 Michigan Oarnegie bo Chevrolat, A. 1843. Coleopteres du Mexique. Magazin. tanical expedition to British Honduras. [522(14):1- de Zoologie, pp. 1 37, pIs. 57, figs.]. In: Botanyofthe Maya area: miscellaneous Chevrolat, A. 1876.Memoire surlafamilIe des Clerites. papers. Carnegie Inst. Wash., Washington, D. C. L. Buquet, Paris. 51 pp. Russell, S. M. 1964. A distributional study ofthe birds Corporaal, J. B. 1950. Cleridae. Coleopterorum ofBritishHonduras.AmericanOrnithologists'Union, Catalogus Supplementa. 23: 1-373. Ornithological Monographs No. 1. 195 pp. Ekis, G 1977 Classification, phylogeny, and zoogeog- Schenkling,S 1903 ColeopteraMalacodermata' Fam raphy of the genus Penlypus (Coleoptera:Clendae). Clendae. FaSCIcle 13.1n: P. Wytsman (edItor), Gen­ Smjthson Contr Zool 227'1-138 era Insectorum Bruxelles 124 pp , 5 tables Gorham, H. S. 1882. BlOlogla Centrah-Amencana, Schenkhng, S. 1906. DIe Clenden des Deutchen Insecta, Coleoptera, Cleridae 3(2)'129-168 Entomologischen National.Museums, nebst Gorham, H. S. 1SS3. Biologia Centrali-Americana, BeschreibungenneuerArten. DeutscheEnt. Zeitschr., Insecta, Coleoptera, Cleridae 3(2): 169-193. pp. 241-320. Henderson, R.W., and L. G. Hoevers. 1975. A Selander, R. B., and P. Vaurie. 1962. A gazetteer to checklist and key to the amphibians and reptiles of accompany the "Insecta" volumes of the "Biologia Belize, Central America. Cont. BioI. Geol. Milwau­ Centrali-Americana." Amer. Mus. Nov. 2099:1-70. kee Pub. Mus. 5.1-63. Spinola,M. 1844. Essaimonographiquesurles Clerites, Hespenheide, H. A. 1987. A revision of Lissoderes Insectes Coleopteres. 2 vols.,Genes, 10 + 602 pp., 47 . Ohampion (Coleoptera. Cmculionidae. Zygopinae). pIs. Coleopts Bull. 41(1):41-55. Stephens, J. L. 1841. Incidents of travel in Central HOFwieh, R. H., and J. Lyon. 1990 [1]. .\Belizean Fain f.merica, Chiapas and Yucatan. 1988 edition forest: the community baboon sanctuary. Orang­ (abridged), Century Hutchinson Ltd., London. 332 utan Press, Gays Mills, Vlisc. 420 pp. pp. Johnson, C. D. 1984. Acanthoscelides belize, N. Sp. Wolcott, A. B. 1927. A review of the Cleridae of Costa (Coleoptera: Bruchidae). J. Kans EntomoL Soc. Rica Col. Contr., 1(1):1 104. 57(2):339·343. KirkpatrICk, R. D., andA. M. CartWrIght. 1975. LISt of mammalsknown tOOCCllrin Belize Biotropica 7'136- 140. Klug, J. C. F. 1842 Versuch einer systematischen Bestimmung und Auseinandersetzung der Gattungen und Arten der Clerii, einer Insecten familie aus der Ordnung der Coleopteren. Abh. Preussische Akad. Wiss., pp. 259-397.