Boletín de la Sociedad Geológica Mexicana ISSN: 1405-3322 Sociedad Geológica Mexicana, A.C.

Mannani, Maryam Late scleractinian corals from Nayband Formation, southwest Ardestan, Central Iran Boletín de la Sociedad Geológica Mexicana, vol. 72, no. 2, 00002, 2020, May-August Sociedad Geológica Mexicana, A.C.

DOI: https://doi.org/10.18268/BSGM2020v72n2a090619

Available in: https://www.redalyc.org/articulo.oa?id=94366150002

How to cite Complete issue Scientific Information System Redalyc More information about this article Network of Scientific Journals from Latin America and the Caribbean, Spain and Journal's webpage in redalyc.org Portugal Project academic non-profit, developed under the open access initiative This is an open access article under the CCBY-NC-SAThis Universidad NacionalAutónoma de México. Peer Reviewing under the responsibility of Manuscript May accepted: 24,2019 Corrected manuscript received: March 11,2019 Manuscript received: December 19,2018 BSGM2020v72n2a090619 http://dx.doi.org/10.18268/A090619. Sociedad Geológica Mexicana, 72(2), Ardestan, Central Iran: Boletín de la corals from Nayband Formation, southwest Mannani, M.,2020,Late Triassic scleractinian How tocitethisarticle: [email protected] * Corresponding author:(M.Mannani) Iran. I. Rof Payame NoorUniversity, 19395-4697Tehran, 1 Maryam central Corales escleractínidosdelTriásicotardíodelaFormaciónNayband,suroesteArdestán,Irán Iran Late TriassicscleractiniancoralsfromNaybandFormation,southwestArdestan,Central (https://creativecommons. org/licenses/by-nc-sa/4.0/) sitn rfso,Dprmn fGeology,Assistant Professor, of Department Mannani 1,* tion, Ardestan, Iran. Late Triassic, Nayband Forma Keywords: Scleractinian corals, Central Asia. Calcareous Alps and Pamirs,ern the Late Triassic coralsfrom North area bear taxonomic resemblance to the studied chiopods). coralsof The gastropods,echinoderms, andbra ( dwellers bivalves, andother reef with hydrozoan these biostromes, andoccurtogether the main reef-building in organisms Formation. Spongesare andcorals Nayband Howz-e-Khan of Members bioconstructions, inBidestan and 24 . They occurinbiostromal attributedforms, to 14 genera and thamnasterioid, and plocoid) growth and colonial(cerioid,meandroid, represented by solitary, phaceloid Ardestan town (Central Iran), is Nayband Formation, southwest of Late Triassiccoral faunafrom the ABSTRACT Boletín de la Sociedad Geológica Mexicana Geológica Sociedad la de Boletín Heterastridium spp., e.g ., - - - colonial (cerioides, meandroides, thamnaste decrecimiento solitario,formas faceloide y tan (Irán Central) está representada por Formación Naybandel suroeste en deArdes La fauna coralina del Triásico tardío de la RESUMEN Irán. mación Nayband, Ardestan, actinios, Triásico tardío, For Palabras clave: Corales escler Pamir, AsiaCentral. tardío de los Alpes Calcáreos delNorte y litud taxonómica con corales del Triásico Los corales del área de estudio tienen simi gasterópodos y echinodermos, braquiópodos). (e.g., yotrosbivalvos habitantes del arrecife asociados al hidrozoo deestos biostromas,tructores yseencuentran cons representan los principales organismos Formación Nayband. Esponjas y corales de la miembros Bidestan y Howz-e-Khan de biostromas,en bioconstrucciones en los y 24especies. Dichos corales se encuentran rioides y plocoides), atribuidas a 14géneros /72(2)2020 Heterastridium spp., 1 ------

Late Triassic corals, Nayband Formation, southwest of Ardestan ABSTRACT INTRODUCTION / GEOLOGICAL Late Triassic corals, Nayband Formation, southwest of Ardestan SETTING 2. Geologicalsetting 1. Introduction 2 2 band Formation (nametakenfrom the Nayband brate fauna is known from the Late Triassic Nay and taxonomically diverse Triassic marine inverte Iran. Particularlycentral and northern abundant Upper Triassic sediments are widely exposed in h abn omto,i 0k otws fthe Nayband Formation,southwest in 40km of coralsdescribed (TableThe 1) were collected from Ardestan town. southwest of locatedsections Bagher in Abad andDizluinthe fauna from Nayband Formation recognized in two Triassic(Norian toRhaetian) scleractinian coral Late is to provide more extensive description of thisstudy in the Late Triassic. of purpose The the TethysrelationshipsOcean with other parts of distributionThis indicates paleobiogeographic also reported from Tajikistan (Melnikova, 2001). thesetaxa were Ardestan town.Many of west of thesouth area wereof alsodescribedin40Km the Ferdows the scleractinian coralsof species of and Howz-e-Khan members. Remarkably, nine lected the Bidestan in fromthe Ferdows patch reef Nayband Formation (14taxa), which were col Norian toRhaetian scleractinian coralsfrom the Hautmann, 2001;Nützel and Senowbari-Daryan, 1999; Repine, 1996; tion ( and bivalvefrom mollusks theNaybandForma published authors studiesonsponges, gastropod, sich 2003), Zahedi (1976), Kluyver byBrönnimann unithavelithostratigraphic this been outlined of Sedimentology, biostratigraphy, andpaleontology e-Sheikh, (ii) Nayband,and (iii)Howz-e-Khan. band Formation intothree (i) Howz- members: out by(1929), who the Naysubdivided Douglas study onNayband Formation strata was carried Tabas). pioneering The Naybandan (southof of thesmallvillage Mountain, 3008m.a.s.l.), eastof / Boletín de la Sociedad Geológica Mexicana Geológica Sociedad la de Boletín et al. e.g. To date, onlyShepherd , Douglas, 1929;Fallahi (2005),andHautmann (2001).Several et al. (1971),Seyed-Emami (1971, et al et al. ., 2003,2010). et al. et al. http://dx.doi.org/10.18268/BSGM2020v72n2a090619 (2012)described , 1983;Nützel (1983), Für ------/72(2)2020 the Howz-e-Khan member. (Brönnimann Qadir is member thatNayband overlayformation sponges, andcorals).Inaddition, upper part of (reefal members limestoneswithalgae, calcareous and Howz-e-KhanSheikh (shalesandsandstones), algae, calcareous sponges, andcorals),Howz-e- sandstones, and reefal limestones withnumerous Bidestan (oolithic limestones,and sandstones), the following (from bottom to top): Gelkan (shales (Figure 1B). Bagher Abad village (33°05’28’’N, 52°03’27’’E) of 1C), and inthe locality situated northeast 15 km Dizlu village (33°04’10’’N, 52°01’40’’E) (Figure of Ardestan, from the outcropseast 5km the town of horizonsin theQadirmember contain bivalves browncomprises and marls. sandstones Two key Qadir Member, uppermost The upto200mthick, the Howz-e-Khan member. the age of chiopods) (Figure corals(Table 4). The 2) indicate ( dwellers isms, includingsponges, corals, andreef with athicklimestonebed(Figure 2F). rests ontheHowz-e-Sheikh Member and starts Howz-e-Khan Member, which isupto 15 mthick, brownunfossiliferous (Figure sandstones 2E).The light mainly of 44mandconsists of a thickness by the Howz-e-Sheikh member, which reaches fauna (Figure Bidestan Member is overlain 4). The blage indicates the late Norian–Rhaetian age of corals is presented in Table assem 2. This list of spp. andbivalves are alsorelatively abundant. A reef-building hydrozoanorganisms; In the biostromes,and coralsare sponges the main biostromal bioconstructions (Figure 2C and 2D). with oolithiclimestoneat the basemember, and member, the Bidestan member is about 11 m thick, ber (Biostromethe Gelkan on conformably 1)lying (Middle Triassic) (Figure Bidestan Mem 2B). The m thick),which onthe Shotori formation deposits is GelkanMember (up to 400 Nayband formation of part Lowermost 2A). (Figure members five has 1971; Kluyver e.g. h ebrcnan bnatre organ abundant member contains The reef , echinoderms, bivalves,, echinoderms, gastropods, andbra The NaybandFormationThe into subdivided is In the Dizlu area, the Nayband Formation et al. , 1983;Fürsich et al. Heterastridium Article A090619 , 2005). et al. - - - - ,

Article A090619 Table 1. Scleractinian corals from the Upper Triassic of Nayband coralsfromtheUpper Triassic of FormationinthesouthwestTable 1.Scleractinian Ardestan. Procyclolitidae VAUGHAN and WELLS, 1943 Procyclolitidae VAUGHANandWELLS, Cyclophyllidae RONIEWICZ,1989 Stylophyllidae FRECH, 1890 Stylophyllidae FRECH, Pamiroseriidae MELNIKOVA,1984 Actinastraeidae ALLOITEAU,1952 Astraeomorphidae FRECH,1890 Margarophylliide VOLZ,1890 Reimaniphyllidae MELNIKOVA,1975 Coryphylliidae Beauvais,1981 gastropods, andbrachiopods)(Figure corals 5).The ( dwellers als, andreef organisms,sponges, including: cor abundant reef reefal limestone (Figures 3Band3C),contains byformed is up to25mthick,cream tolight-gray beds (Figures Howz-e-Khan 3A).The Member, Qadir overlainMember is byEarly mation startswiththeHowz-e-Khan Member. The byunconformity Early Cretaceous deposits. NaybandFormation overlain is withanangular indicate a Rhaetian age (Figures 2Gand2H).The ( Indopecten glabra In the Bagher Abad area, the Nayband For ) and flora ( Family e.g. , echinoderms, bivalves,, echinoderms, Clathropteris http://dx.doi.org/10.18268/BSGM2020v72n2a090619 Boletín de la Sociedad Geológica Mexicana Mexicana Geológica Sociedad la de Boletín spp.), which - - Astraeomorpha confusa Astraeomorpha crassisepta Procyclolites triadicus Kompsasteria oligocystis Distichomeandra austriaca Margarophyllia Margarophyllia Margarophyllia Oedalmia norica Retiophyllia clathrata Retiophyllia norica Distichophyllia Cuifia elliptica Cuifia gigantella Cuifia columnaris Pamirophyllum iranicum Stylophyllopsis lindstroemi Stylophyllopsis rudis Stylophyllopsis polyactis Crassistella juvavica Pamiroseris multiseptata Pamiroseris rectilamellosa Pamiroseris meriani Chondrocoenia schafhaeutli 3. Materials oal eescindt hc rsraino coralla were sectioned to check preservation of represented by eight scleractinian families. Several 88specimens, materialThe examinedof consists byunconformity Early Cretaceous deposits. isoverlainNayband formation, withanangular thekeythe QadirMember,beds in the One of m thick,containsbrownand marls. sandstones e-Khan Member. Qadir Member, The upto 125 (Table 2) indicate a Rhaetian age for the Howz- (MELNIKOVA,1975) cf. sp. cf. cf. (FRECH,1890) (MELNIKOVA,1975) (RONIEWICZ,1996) norica (FRECH,1890) crenata capitata (EMMRICH,1853) (STOPPANI 1858-1860) (EMMRICH,1853) Genus andspecies (FRECH, 1890) (FRECH, 1890) (WINKLER, 1861) (FRECH,1890) (FRECH,1890) (MELNIKOVA) (MELNIKOVA, 1972) (FRECH,1890) (FRECH,1890) (WINKLER, 1861) (MÜNSTER,1841) (REUSS, 1854) (FRECH, 1890) (MÜNSTER,1841) (WINKLER, 1861) / 72(2)2020 3 3

Late Triassic corals, Nayband Formation, southwest of Ardestan GEOLOGICAL SETTING / MATERIALS Late Triassic corals, Nayband Formation, southwest of Ardestan MATERIALS Figure 1 4 4 recognizable, well preserved and major diagnostic features are patterns. Macroscopically, coralla are generally order Sciences)in diagenetic toassess Academyof Paleobiology,microscopy (at Polish the Institute of were also examined by cathodolumienscence composing the skeletal structures. A few samples pletely recrystallized withblockycalcite crystals the original structure. Coralla are typicallycom / Geological map and location of the studied sections in the southwest of Geologicalmapandinthesouthwest Ardestan,B.Bagher locationofthestudiedsections Abad,and C.Dizlusection. Boletín de la Sociedad Geológica Mexicana Geológica Sociedad la de Boletín i.e. , septa, axial structures,, septa, walls. http://dx.doi.org/10.18268/BSGM2020v72n2a090619 - /72(2)2020 University. museumPayame samples in Shahinshahr Noor (8378–8467) are codenumber shown:The of Coral) (8378–8467). (Shahin shahr Payame Noor University Iran Payame-Noor, under the “SH-PNUICO” symbol location repository housed at is the Univeristy of sometimes visible ( features micromorphological are Some traces of e.g. , septal granulations).The granulations).The , septal Article A090619 Article A090619 rocks assigned totheNayband Formation. (B)Alternatedbrown sandstoneand marl, GelkanMember. (C)Reefal limestone,Bidestan Figure 2 Reefal limestone, Howz-e-KhanMember.(G)Bivalve keybed, QadirMember. (H)Flora keyhorizon, QadirMember. in limestone, BidestanMember. (E)Alternated brownsandstones corals Member. and (D)Scleractinian marls,Howz-e-Sheikh Member. (F) Exposures oftheNayband ofLateTriassic deposits Formation,Dizluarea,southwestof Ardestantown.(A)Fieldviewof the http://dx.doi.org/10.18268/BSGM2020v72n2a090619 Boletín de la Sociedad Geológica Mexicana Mexicana Geológica Sociedad la de Boletín / 72(2)2020 5 5

Late Triassic corals, Nayband Formation, southwest of Ardestan MATERIALS Late Triassic corals, Nayband Formation, southwest of Ardestan MATERIALS Table 2.DistributionofCoralsintheNorianof toRhaetianofthesouthwest Ardestan. 6 6 wall. S/mm S H LCD GCD D d c-c following: Abbreviations usedinthe text includethe / imtro juvenile corallite, : diameterof : number of septa; septa; : number of imtro adultcorallite, : diameterof : height of the solitary coralor colony, the solitary : heightof Boletín de la Sociedad Geológica Mexicana Geológica Sociedad la de Boletín : distancebetween thecalicular axes, Crassistella juvavica Pamiroseris multiseptata Pamiroseris rectilamellosa Pamiroseris Stylophyllopsis Stylophyllopsis rudis Stylophyllopsis polyactis Chondrocoenia Astraeomorpha confusa Astraeomorpha crassisepta Procyclolites triadicus Kompsasteria oligocystis Distichomeandra austriaca Margarophyllia Distichophyllia Cuifia elliptica Cuifia gigantella Cuifia columnaris Pamirophyllum iranicum Margarophyllia Margarophyllia Oedalmia norica Retiophyllia clathrata Retiophyllia norica : smallercaliculardiameter, : greater caliculardiameter, : septal density measured: septal at the calicular meriani Taxon schafhaeutli cf. cf. sp. cf. c cf. lindstroemi capitata norica S1 renata , S2 …etc., http://dx.doi.org/10.18268/BSGM2020v72n2a090619 ▄ ▄▄ ▄ ▄▄ ▄▄ ▄ ▄ ▄ ▄ ▄ ▄▄ ▄▄ 22 12 biostromes Dizlu Area Section C /72(2)2020 4. Systematicpaleontology ▄▄ ▄▄ ▄▄ ▄▄ ▄▄ ▄ ▄ ▄ ▄ ▄ ▄ ▄ ▄ ▄▄ Type species: Family Coryphylliidae BEAUVAIS, 1981 1996 Cuifia columnaris Order ScleractinaBOURNE,1900 Genus Cuifia columnaris Cuifia Figures 7Aand7B Cuifia gigantella Bagher Abadarea MELNIKOVA, 1975 biostrome Section B RONIEWICZ, 1996 : RONIEWICZ, Melnikova, 1975 Article A090619 Article A090619 Heterastridium Hydrozoan(C) Howz-e-Khan member. in reefallimestone, colony coral Naybandexposures.ofFormation rockScleractinian (B) Figure 3 HPUC80 56 3 1 5 110 135 65×67 SH-PNUICO8408 elements are numerous, differentiated into differentiated numerous, more than 3 size orders. Endotheca is formed are elements faces. and granulation on septal septa Radial a considerable diameter, thick calice of and deep sub-cylindrical shape, circular corallum of itary Description: surements (inmm): fromBiostrome 2,Section area.C, Dizlu Mea Material: ubrDHSS/5mm S H D Number Exposures of ofLateTriassic theNayband deposits Formation, Bagher Abadarea,southwest of Ardestan.(A)Fieldview p. 21,pl.3,figs. 1-3; pl. 4,figs. 1,4,5 2001 sp.inlimestoneofHowz-e-Khanmember. (D)Bivalvekeybed, Qadir member. MELNIKOVA, p. 43,pl.11,fig. 1 One specimen (SH-PNUICO8408) Cuifia columnaris The coral is characterized The by sol RONIEWICZ: http://dx.doi.org/10.18268/BSGM2020v72n2a090619 Boletín de la Sociedad Geológica Mexicana Mexicana Geológica Sociedad la de Boletín - - Tethys. the the European and Asianparts of Rhaetian of species isknownThe from the upper Norian and Central Iran. Ardestan, e-Khan Member, Dizlu, southwest of Occurrence: conditions. paleoenvironmental in succeedingchanges of ular diameters, which resulted from development posed growth stageswithsmaller or larger calic destroyed.surface is corallumshowssuper The byvesicular dissepiments. Wall epithecal thick, 1975 Cuifia gigantella Cuifia gigantella Late Triassic, Rhaetian, Howz- Figures 7Gand7H pl. 14,figs. 1,2 MELNIKOVA, 1975 : MELNIKOVA, p. 83, / 72(2)2020 7 7 - -

Late Triassic corals, Nayband Formation, southwest of Ardestan SYSTEMATIC PALEONTOLOGY Late Triassic corals, Nayband Formation, southwest of Ardestan SYSTEMATIC PALEONTOLOGY Figure 4 8 8 / Biostratigraphic rangeoffauna Late Biostratigraphic chart Triassic from theNayband Formation,of Dizluarea,southwest Ardestan. Boletín de la Sociedad Geológica Mexicana Geológica Sociedad la de Boletín http://dx.doi.org/10.18268/BSGM2020v72n2a090619 /72(2)2020 Article A090619 Article A090619 Figure 5 Biostratigraphic range of Late Triassic faunarangeTriassic Biostratigraphic ofLatefrom theNayband Formation, Bagher Abadarea,of southwest Ardestan. http://dx.doi.org/10.18268/BSGM2020v72n2a090619 Boletín de la Sociedad Geológica Mexicana Mexicana Geológica Sociedad la de Boletín / 72(2)2020 9 9

Late Triassic corals, Nayband Formation, southwest of Ardestan SYSTEMATIC PALEONTOLOGY Late Triassic corals, Nayband Formation, southwest of Ardestan SYSTEMATIC PALEONTOLOGY triadicus PNUICO8429, C,Dizluarea. (F) Colonyincalicular view.(G) Colony insideview.(H,I) Biostrome1,Section B,Bagher Abadarea.(D)Colonyincalicularview.(E) Section side view.(F,G) Biostrome2, (D, E)SH-PNUICO8428, view. (C)Colonyintransverse section. Colony incalicular Dizlu area.(B) C, Section Biostrome 2, PNUICO8424, C, Dizlu Biostrome2,Section area. Colony in calicular view.(B–E) (Frech, 1890). SH-PNUICO8432, Biostrome 2, Section C,Dizluarea.Colony incalicularview. Biostrome2,Section (Frech, 1890).SH-PNUICO8432, C,Dizluarea.(H)Corallum incalicularview.(I)Corallum Biostrome1,Section (Frech, 1890)SH-PNUICO8426, insideview.(J) Figure 6 10 10 / Scleractinian coralsfrom Scleractinian theNayband Formation,southwest of Ardestan,Iran. (A) (Frech,1890).SH-PNUICO8401, C,Dizluarea.Corallum incalicular view.(K) Biostrome1,Section Boletín de la Sociedad Geológica Mexicana Geológica Sociedad la de Boletín http://dx.doi.org/10.18268/BSGM2020v72n2a090619 /72(2)2020 Retiophyllia norica Retiophyllia clathrata (Frech,1890).(B,C)SH-PNUICO8466, Oedalmia norica Distichomeandra Distichophyllia (Emmrich, 1853).SH- (Frech, 1890)SH- Article A090619 cf. Procyclolites cf. austriaca norica

Article A090619 HPUC80 34×45 SH-PNUICO8406 SH-PNUICO8405 HPUC80 3×34 67 96 41 35×43 SH-PNUICO8407 ubrDH D Number The species is knownis species from The the Norian–Rhaetian Central Iran. Ardestan, e-Khan Member, Dizlu, southwest of Occurrence: thick,partlydestroyed.epithecal, all. Wall arethe S4 septa thin andthe shortest of blades andseptal of and shorter than the S2 septa, in thickness,the S1 septa are the S3 septa thinner of cavity, are the S2 septa half blades septal of continue the S1 septa to axial bladesthe septal of faces. into4sizeseptal arranged orders: Septa on granulation high, and thick order size first the shape, calice circular, of septa slightly deepened, Description: surements (inmm): fromBiostrome 2,Section area.C, Dizlu Mea Material: 2001 Measurements (inmm): 8407) from Biostrome 2,Section C, Dizlu area. Material: 2001 1989 1975 thePamir Mountains. of ubrDHSS/5mm S H D Number ueie1×63.5 12×16 juvenile (adult) RONIEWICZ, p. 65,pl.15,figs. 1,2,7 MELNIKOVA, p. 43,pl.11,figs. 2–4 Cuifia elliptica Cuifia gigantella Cuifia Cuifia elliptica Cuifia Two(SH-PNUICO8405, specimens One specimen (SH-PNUICO8406) Coryphyllia elliptica Coryphyllia Corallum solitary, low-conical in Late Triassic, Rhaetian, Howz- 12 79 9 92 47 21×24 Cuifia elliptica

Figures 7Cto7F 14, figs. 3–5 10, figs. 1,2 MELNIKOVA, 1975 8181+42+1)6 168(14+14+28+112) 28 : MELNIKOVA, p. 84,pl. MELNIKOVA, p. 42, pl. MELNIKOVA: http://dx.doi.org/10.18268/BSGM2020v72n2a090619 S (Melnikova): Boletín de la Sociedad Geológica Mexicana Mexicana Geológica Sociedad la de Boletín S/5mm 11-12 - HPUC82 84 5167-8 7-8 156 156 45 59 40 29 48 51 SH-PNUICO8427 SH-PNUICO8426 ubrGDLDHSS/5mm S H LCD GCD Number The species is knownis species from The the Norian–Rhaetian Central Iran. Ardestan, e-Khan Member, Dizlu, southwest of Occurrence: the rejuvenated stage, thewallisvery thick. rejuvenation. At stagesof superposed mation of environmental conditions, which resulted in for destroyed. corallum developed This in variable are ontheorder. shorter depending Epitheca is other septa shorter and thinner than the S1 septa; are the S2 septa slightly fossa, blades septal of continueto the center: the S1 septa to the axial elliptica is characterized by typical of size and form, Description: Grosser (holotype), Hallstätter Zlambachgraben Frech (1890)described Central Iran. Ardestan, Bidestan Member,southwest Dizlu, of Occurrence: fossa,Endotheca is vesicular. Epitheca is destroyed. from which S1 reach the septa to the calicular orders, size 3 least at into differentiated are ments maySepta reach to theaxialcavity. the skeleton. Radialele of features specific of examination preservation does notallow for detailed state of tures to Description: Measurements (inmm): 8427) from Biostrome 1,Section C, Dizluarea. Material: thePamir Mountains. of Family Reimaniphylliidae MELNIKOVA, 1975 Type species: Distichophyllia h ogs et ftwo size ordersreach of longest septa . The Distichophyllia norica Genus Two(SH-PNUICO8426, specimens The specimenSH-PNUICO8405 The Late Triassic, Rhaetian, Howz- Late Triassic, Norian–Rhaetian, The coral is similaringeneral fea The Figures 6Hand6I Distichophyllia Montlivaltia norica cf. norica / 72(2)2020 Montlivaltia norica (FRECH,1890) (Frech, 1890),but its CUIF, 1974 FRECH,1890 from 11 11 C. - - -

Late Triassic corals, Nayband Formation, southwest of Ardestan SYSTEMATIC PALEONTOLOGY Late Triassic corals, Nayband Formation, southwest of Ardestan SYSTEMATIC PALEONTOLOGY Kompsasteria oligocystis (I) Corallumview. in proximal(H) view. in calicular Corallum (G) Dizlu area. C, Section 2, Biostrome 8406, SH-PNUICO 1975). (Melnikova, H) (G, view. in calicularCorallum Dizlu area. C, Section 2, Biostrome SH-PNUICO8407, (F) section. in transverse Corallum (Melnikova, 1975).(C–E)SH-PNUICO8405, C, Dizluarea.(C)Corallum Biostrome2,Section in calicular view. (D)Corallum insideview.(E) C,Dizluarea.(A)Corallum1996). SH-PNUICO8408, incalicularview.(B)Corallum Biostrome2,Section in side view.(C–F) Figure 7 12 12 / Scleractinian corals from the Nayband Formation, southwest of Ardestan,CentralIran, corals from Iran. theNaybandsouthwest (A,B) Formation, Scleractinian Boletín de la Sociedad Geológica Mexicana Geológica Sociedad la de Boletín (Frech,1890).SH-PNUICO8433, C,Dizluarea. Colonyintransversesection. Biostrome2,Section http://dx.doi.org/10.18268/BSGM2020v72n2a090619 /72(2)2020 Cuifia columnaris Cuifia gigantella Cuifia elliptica Article A090619 (Roniewicz,

Article A090619 HPUC86 1.×15 14×13 51 13.5×21 SH-PNUICO8466 HPUC82 53 5712×15 35.7 15×33 SH-PNUICO8428 SH-PNUICO8467 ubrDHd H D Number from Biostrome 2,SectionB, BagherAbadarea. Dizlu area and onespecimen(SH-PNUICO8428) SH-PNUICO8467) fromBiostrome 2,Section C, Material: 1989 1890 NorthAmerica. of according toStanley(1986), present the Norian in Tethys,the European andAsiatic partsof and moos,in Species common andSchneckengraben. Fischerwise, Kesselwand-Rohr ZlambachBeds of Calcareous Alps:inthe the Northern tian of Roniewicz (1989) the species ispresent in the Rhae Salzberg andScharitzkehlalp. According to otenClaeu ls lmahBd f Calcareous Alps: Zlambach Beds of Northern the speciesisknown fromThe the Rhaetian of Ardestan, CentralIran. west of Khan Member, DizluandBagher Abad, south Occurrence: the order. to thecenter, the remaining are septa shorterin continue S1 septa into 3–4 orders. blades Septal of differentiated fusiform, Septa angle. obtuse with SH-PNUICO8426 shows corallites bifurcating Description: Measurements (inmm): p. 52,pl.8,figs. 5–8; pl. 9,fig. 2;pl.14, figs. 1,2 Type species: Retiophyllia norica Thecosmilia norica Thecosmilia 33. 3312.7×17 43.3 13×37.8 Retiophyllia norica Genus Two(SH-PNUICO8466, specimens 14–24 (nonpl.10,fig. 6) Corallum phaceloid. The specimen Corallum phaceloid. The Late Triassic,Rhaetian, Howz-e- Retiophyllia frechi Figures 6Bto6E Retiophyllia (FRECH):RONIEWICZ, FEH p 9 p. , figs. 1, pl. 9, p. :FRECH, (FRECH,1890) 0(91+03)8 105(19+16+40+30) 7(51+63)7juvenile 7 78(15+12+16+34) CUIF, 1966 68 79 84 90 S http://dx.doi.org/10.18268/BSGM2020v72n2a090619 Roniewicz, 1989 /m Observations S/3mm Boletín de la Sociedad Geológica Mexicana Mexicana Geológica Sociedad la de Boletín 7-8 - juvenile 6-7 7 bifurcation bifurcation bifurcation juvenile before before before - - - HPUC83 .×242. .×. 014 8 70 30 6.4×6.4 5.4×8 21.5 18.4 7.4×12.4 SH-PNUICO8430 7.4×9.6 SH-PNUICO8429 ubrDHdSS/3mm S d H D Number and SH-PNUICO8430) from Biostrome 1,Sec Material: 1974 1890 NorthAmerica. species intheNorianof Roniewicz, 1989). Stanley (1986) mentioned this Fischerwise (Frech, andZlambachgraben 1890; ( (Reuss) in Smith,1927,andfrom South America sis North America ( Norian of the from described were names specific different the Alps and Carpathians. Similar corals under characteristic speciesis The for the Rhaetian of Central Iran. Ardestan, Bidestan Member,southwest Dizlu, of Occurrence: size orders. are vesicular. Dissepiments 2–3 into differentiated are granular. Septa is tion apparatus. septal Ornamenta of simple division bifurcating or trifurcating. species shows The a is characterized by phaceloid corallum, corallites Description: tion C,Dizluarea. Measurements (inmm): Thecosmilia Type species: 1853 inClapp&Shimer, 1911; Retiophyllia clathrata Retiohyllia clathrata Thecosmilia clathrata Thecosmilia Lithodendron clathratum p. 108,pl.4,fig. 1,pl.5,figs. 3,7 Oedalmia norica Two specimens (SH-PNUICO8429 Genus sp. indetinMaeda 15, pl.4,figs. 1–5,7–11 The specimenSH-PNUICO8430 The Late Triassic, Norian–Rhaetian, Thamnastraea norica Thamnastraea Figures 6Fto6G Oedalmia Figure 6A (Emmrich): RONIEWICZ, (FRECH,1890) (EMMRICH,1853) EMMRICH: FRECH, p.FRECH, EMMRICH: / 72(2)2020 CUIF, 1976 : EMMRICH,p. 78 Calamophyllia suttonen Thecosmilia fenestrata Thecosmilia et al. FRECH,1890 , 1974). 13 13 - - -

Late Triassic corals, Nayband Formation, southwest of Ardestan SYSTEMATIC PALEONTOLOGY Late Triassic corals, Nayband Formation, southwest of Ardestan SYSTEMATIC PALEONTOLOGY 14 14 S-NIO443×4991. - 81+01)5 38(10+10+18) 6-7 9×11.6 9 34×54 SH-PNUICO8424 Number from Biostrome 2,Section C, Dizluarea.Measure Material: 1989 1890 specimens (SH-PNUICO8421 and 8422) from 8415, 8417,8420)from Biostrome 1,and2 Material: up tonow from thelate Carnian. with areserve to the genus Remarks: Roniewicz, 1989). Fischerwise, Kesselwand-Rohrmoos(Frech, 1890; Calcareous Alps:Zlambach Bedsof Northern the speciesisknown fromThe the Rhaetian of Central Iran. Ardestan, e-Khan Member, Dizlu,Southwest of Occurrence: are shortandthin. theS3septa blades septal of shorter thanS1 septa, to axialcavity, are the S2 septa thinnerandslightly continue the S1 septa size blades orders: septal of cavity is fissure-like.The are septa arranged into 3 linked by blades. simplebiseptal Inthe calice, axial the series,In and meandroid calicesare form. thamnasteroid colony margin. There is mixingof colony, withsimplecalicular series paralleling the Description: ments (inmm): / Type species: Boletín de la Sociedad Geológica Mexicana Geológica Sociedad la de Boletín Margarophyllia Family Margarophylliidae CUIF, 1977 Thamnastraea norica Thamnastraea RONIEWICZ, p. 62,pl.15,figs. 3–6 colony Genus Dof Oedalmia norica Four(SH-PNUICO8411, specimens One specimen (SH-PNUICO8424) Three taxa have been here attributed colony The specimen represents The a laminar Late Triassic, Rhaetian, Howz- H of Margarophyllia Montlivaltia capitata Figures 8Hand8I cf. pl. 17,figs.1–6 capitata c-c d nov. sp.: FRECH,p. 63, (FRECH,1890): (MÜNSTER,1841) VOLZ, 1896 Margarophyllia http://dx.doi.org/10.18268/BSGM2020v72n2a090619 Münster, 1841 S known S/2mm - /72(2)2020 HPUC82 51 61614 10 116 15 108 9 16 84 19 13-14 13 96 15 11 84 17 15 21 96 6 18 18 SH-PNUICO8422 12 28 10 SH-PNUICO8421 12 SH-PNUICO8420 13.7 10 12 SH-PNUICO8417 16 SH-PNUICO8415 SH-PNUICO8411 HPUC81 . 41430 17 144 12 144 14 11 120 18 13 120 7.6 22 24 132 7 28 9 200 21 18 12 SH-PNUICO8419 13 8 26 SH-PNUICO8412 11 19 SH-PNUICO8418 11 18 SH-PNUICO8414 12 SH-PNUICO8410 SH-PNUICO8409 ubrGDLDHSS/5mm S H LCD GCD Number ubrGDLDHSS/5mm S H LCD GCD Number Septa spindle-like, differentiated into several size several into differentiated spindle-like, Septa short calicular fossa.Calice ellipsoid anddeep. curved proximally, subcircular in section,with Description: ments (inmm): Biostrome 2, Section C, Dizlu area. Measure with shortcalicular fossa. thin, spindle-like, Septa to ellipsoid, from to shallow, slightlydeepened shape slightly curved proximally. Calice circular Description: Measurements (inmm): Section C,Dizluarea. 8410, 8414,8418,8412,8419)from Biostrome 1, Material: Ardestan, CentralIran. Dizlu, southwest of Bidestan Member and Howz-e-Khan Member, Occurrence: the bestpreserved proximally. dissepiments. Epithecal wall pellicular and fragile, be observed. small Endotheca vesicular, built of granulation and denticulation ondistaledge can flat lateral features: septal Other septa. S1 the of the length S3 are the septa longer than half septa, aresepta shorter and slightlythinner than the S1 theS2 dissociate into fragments. blades Septal of the calicular fossa with their axial margins, which orders. are S1septa the thickest, reach The to Margarophyllia Six specimens(SH-PNUICO8409, Corallum solitary, conical,slightly Corallum solitary, conicalin Late Triassic, Norian–Rhaetian, Figures 8Ato8G cf. crenata (MÜNSTER,1841) Article A090619 - Article A090619 PNUICO8416, C, DizluArea. (J)Corallum Biostrome 1,Section incalicular view.(K)Corallum insideview. PNUICO C,Dizluarea.(H)Corallum in calicularview.(I)Corallum 8415,Biostrome1,Section insideview.(J,K) Corallumin sideview. (H,I) Dizlu area.(F)Corallumincalicularview.(G) C, Section 1, Biostrome SH-PNUICO8412, G) (F, section. Corallumin transverse (E) view. in side Corallum (D) view. in microstructureCorallum (C) C,Dizlu area.(A)Corallum(Münster, 1841).SH-PNUICO8414, incalicular view. (B)Corallum Biostrome1,Section inmicrostructure view. Figure 8 Scleractinian corals fromtheNayband Scleractinian Formation, southwest of Ardestan,CentralIran, Iran. (A–G) http://dx.doi.org/10.18268/BSGM2020v72n2a090619 Boletín de la Sociedad Geológica Mexicana Mexicana Geológica Sociedad la de Boletín Margarophyllia cf. / 72(2)2020 capitata Margarophyllia Margarophyllia (Münster, 1841) SH- (Münster,1841)SH- cf. crenata sp.SH- 15 15

Late Triassic corals, Nayband Formation, southwest of Ardestan SYSTEMATIC PALEONTOLOGY Late Triassic corals, Nayband Formation, southwest of Ardestan SYSTEMATIC PALEONTOLOGY 16 16 HPUC81 691 23+6 12 72(36+36) 16 9 16 SH-PNUICO8416 ubrGDLDH LCD GCD Number nohc eiua,md fsmalldissepiments. Endotheca vesicular, made of margin distal septal canbe observed. ulation of GranulationS2 septa. faces anddentic septal on the length of arethe S3 septa longer than half dissociate are into fragments; the S2 septa shorter; tinue to the calicular fossa andtheir axial margins differentiated into 3 size orders.The S1 septa con Measurements (inmm): 8432) from Biostrome 2,Section C, Dizluarea. Material: Central Iran. Ardestan, Bidestan Member,southwest Dizlu, of Occurrence: is destroyed. corallum, epitheca theca but in the upper part of the corallum is covered with epi Lower part of dissepiments. ofsmall made Endotheca tifiable. other orders are hardly iden of Septa S1 septa. andthinner from the S2 are shorter at about a half reachingsepta toaxial cavity, the blades septal of the S1 lation on the distal edge. blade Septal of calice ellipsoid, deep. thick, withdenticu Septa Description: surements (inmm): fromBiostrome 1,Section area.C, Dizlu Mea Material: Central Iran. Ardestan, Bidestan Member,southwest Dizlu, of Occurrence: quently destroyed. Epitheca thin,generally preserved proximally, fre / Type species: Boletín de la Sociedad Geológica Mexicana Geológica Sociedad la de Boletín Distichomeandra austriaca Genus Two(SH-PNUICO8431, specimens One specimen (SH-PNUICO8416) Corallum solitary,shape,in conical Late Triassic, Norian–Rhaetian, Late Triassic, Norian–Rhaetian, Distichomeandra Distichomeandra spinosa Margarophyllia Figures 8Jto8K Figure 6K (FRECH,1890) CUIF, 1976 sp. http://dx.doi.org/10.18268/BSGM2020v72n2a090619 S Cuif, 1976 S/5 mm ------/72(2)2020 HPUC83 52 48956(51+0 7 60(15+15+30) 7 5 58(14+14+30) 8×9 9.5 18×12.5 14 50 20 40 25 60 SH-PNUICO8432 SH-PNUICO8431 HPUC83 352 17 1+04)8 76(16+20+40) 11 13.5×20 SH-PNUICO8433 Number ubrdc-c d Number septal faces,septal they are round or elongate. Endoth in length. Granulesthe S1 septa are present on the of are the S3 septa half thinner than the S1 septa; are the S2 septa shorter and ity, blades septal of arethe S1 septa thick andcontinue to axial cav into 3 sizethin, arranged orders. blades Septal of the adult. Septa center in the extended portion of calicular series.a new as new corallite A appears colonywith short parts of centric calicesandsome is characterized by polygonal, shallow andmono Description: Measurements (inmm): (SH-PNUICO8433), Dizlu area. Material: RONIEWICZ, p. 30, pl. 2, figs. 2, 3, pl. 3, figs. 1–5 1989 (Roniewicz, 1989). the Gosaukamm region, and in Fischerwise of Calcareous Alps,Northern inthe Zlambach Beds the known speciesis in the Rhaetian of The Central Iran. Ardestan, Khan Member, Dizlu, southwest of Occurrence: Late Triassic,Rhaetian, Howz-e- corallites. but from differs it in larger of diameters colony, budding, septa, andthick of modeof form Remarks: eca vesicular, concave. 1890 Family Cyclophyllidae RONIEWICZ, 1989 Genus GCD of GCD Kompsasteria oligocystis colony Kompsasteria oligocystis Isastraea oligocystis OnesamplefromBiostrome 2 The coral resembles The LCD of LCD Kompsasteria colony The specimenSH-PNUICO8432 The figs. 8,8A, b, 8B colony H of Figure 7I RONIEWICZ, 1989 : FRECH,p. 24,pl.5, d c- c c- d (FRECH,1890) (FRECH,1890): D. austriaca S Article A090619 S inthe S/3 S/3 mm S/3 - - - Article A090619 HPUC80 62 61410 144 36 21 26 SH-PNUICO8401 ubrGDLDHSS/3mm S H LCD GCD Number the Northern Calcareous Alps (Roniewicz,the Northern 1989). and Kesselwand-RohrmoosSchneckengraben in known speciesis from The the Zlambach Bedsof Central Iran. Ardestan, e-Khan Member, Dizlu, southwest of Occurrence: of large corallites,to descriptions the coral answers Remarks: cal surfaceisconcave. round or elongate. Endotheca vesicular, endothe Granules are present faces, on the septal theyare in length. the S1 septa of are S3 septa half septa, are S2 septa shorter and thinner than S1 blades of continue S1 septa to axial cavity,blades of septal theadult. extended portionof A newcorallite appear as anewcenter in the tric calices andshort,monolinear calicular series. characterizedis byshallow, polygonal, monocen Description: faces covered by menianes. ments thin, numerous, equal in thickness. Lateral Description: Measurements (inmm): from Biostrome 1,SectionC,Dizluarea. Material: 1989 1890 Family Procylolitidae Vaughan, andWells, 1943 Type species: K. oligocystis Septa thin, arranged into 3size thin,arranged orders.Septa Septal Procyclolites triadicus Procyclolites triadicus p. 85,pl.23,figs. 1–3,pl.24,figs1,2 Procyclolites triadicus Genus One specimen (SH- PNUICO 8401) With itsvery and thinwall and septa, figs. 1–16,text-fig. p.65 . Calice slightly concave. Radial ele specimenSH-PNUICO8433 The Late Triassic, Rhaetian, Howz- Procyclolites triadicus Procyclolites Figure 6J FRECH:RONIEWICZ, : FRECH,p. 64,pl. 18, FRECH,1890 FRECH,1890 http://dx.doi.org/10.18268/BSGM2020v72n2a090619 FRECH,1890 Boletín de la Sociedad Geológica Mexicana Mexicana Geológica Sociedad la de Boletín - - - HPUC82 52 81521621 3-4 10 1.6-2 1.5×2 28 15×25 SH-PNUICO8425 Number berg (Frech, andZlambachgraben 1890). (Roniewicz, 1989); reported from Halstatter Salz Fischerwise and Kesselwand-Rohrmoos Beds of Calcareous Alps:inthe Zlambach Northern the known speciesis in the Rhaetian of The Central Iran. Ardestan, Bidestan Member,southwest Dizlu, of Occurrence: eso Fshrie iiiyo Gosaukamm, Fischerwise, of vicinity Beds of Calcareous Alps:Zlambach the Northern tian of present speciesis The in the Norian and Rhae Central Iran. Ardestan, Bidestan Member, Dizlu, Southwest of Occurrence: dissepiments. low, by endotheca formed thin and vesicular shape,nar in notabundant, septa density septal Description: Measurements (inmm): from Biostrome 1,SectionC,Dizluarea. Material: 1989 MELNIKOVA, p. 56, pl. 20, figs. 1–3, pl. 21, fig. 2 1980 1890 1854 Family FRECH,1890 Astraeomorphidae 66, text-fig. on p. 70,pl.19,figs. 14–18 Astraeomorpha crassisepta Astraeomorpha crassisepta RONIEWICZ, p. 94,pl.28,figs. 3–5 1986 Astraeomorpha crassisepta Type species: Genus BARI-DARIAN, p. 38,pl.2,fig. 3 Astraeomorpha crassisepta colony Dof Astraeomorpha crassisepta One specimen (SH-PNUICO8425) Astraeomorpha crassisepta Colony thamnasteroid, colum Late Triassic, Norian–Rhaetian, Late Triassic, Norian–Rhaetian, Figures 10Aand10B Astraeomorpha colony Hof pl.16,figs. 4–7 REUSS, 1854 Astraeomorpha crassisepta - S/1mm S c-c d / 72(2)2020 REUSS, 1854 REUSS: p.FRECH, REUSS, 1854 REUSS: SENOW : REUSS, p. 127, REUSS: REUSS:

17 17 - - - -

Late Triassic corals, Nayband Formation, southwest of Ardestan SYSTEMATIC PALEONTOLOGY Late Triassic corals, Nayband Formation, southwest of Ardestan SYSTEMATIC PALEONTOLOGY calicular view.(F) Colonyintransversesection. in calicularview. (E,F) B,Bagher C,Dizluarea.Colony AbadBiostrome 2,Section area.Colonyincalicular Biostrome 2, Section view.(D)SH-PNUICO8458, B,Bagher (C, D) Abadarea.Colonyintransversesection. 2, Section B,Bagher(Winkler, Abad area.Colonyincalicular Biostrome 2,Section view. (B)SH-PNUICO8450, 1861).(A)SH-PNUICO8449, Biostrome Figure 9 18 18 / Scleractinian coralsfrom theNayband Scleractinian Formation,of Ardestan, southwest Central Iran, Iran. (A,B) Boletín de la Sociedad Geológica Mexicana Geológica Sociedad la de Boletín Pamiroseris meriani http://dx.doi.org/10.18268/BSGM2020v72n2a090619 (Stoppani, 1858-1860). SH-PNUICO8462, Biostrome 2, Section C, Dizluarea.(E)Colonyin (Stoppani,1858-1860). SH-PNUICO8462, Biostrome 2,Section /72(2)2020 Pamiroseris multiseptata (Melnikova, 1975). (C) SH-PNUICO8457, (Melnikova,1975).(C)SH-PNUICO8457, Pamiroseris rectilamellosa Article A090619

Article A090619 HPUC84 02 518221521 4-5 18 1.5-2 1.8×2.2 15 20×22 SH-PNUICO8445 Number RONIEWICZ, p. 96, pl. 28, figs. 1, 2; pl. 29, figs. 29, pl. 2; 1, figs. RONIEWICZ,28, pl. p.96, 1989 KRISTIAN-TOLLMANN, TOLLMANN and 1979 1890 1861 North America. to Stanley (1986), it is present in the Norian of Timor andNEAsia(Roniewicz, 1989).According the Caucasus, Central Asia; Norian–Rhaetian of Alps(Lombardy); the Southern Rhaetian of Tyrol; Scharitzkehralp; Rhaetian limestones of Hallstätter SalzbergZlambachgraben, and oino NorthAmerica. Norian of According toStanleypresent(1986) itis inthe al. Timor (Roniewicz, 1989). Kristian-Tollmann theAlps, Caucasus, Central and Asia, tian of in the Norian andRhae common speciesis The Central Iran. Ardestan, e-Khan Member, Dizlu, southwest of Occurrence: synapticulae numerous, thin. dissepiments zone, are septa subparallel. Columella present, in severalarranged series. Inthe intercalicular Description: surements (inmm): fromBiostrome 2,Section area.C, Dizlu Mea Material: MELNIKOVA, p. 57,pl.20,fig. 4,pl.21,fig. 1 (90 eotdi rmteRata fIran. (1980) reported it from the Rhaetian of 1986 1980 Astraeomorpha confusa Thamnastraea confusa Thamnastraea Astraeomorpha confusa p. 67,pl.19,figs. 2,3, 5,6,8,10,13 HAMEDANI, p. 171,pl.4,fig. 3 colony Astraeomorpha confusa Dof Astraeomorpha confusa SCHÄFER, p. 46,pl.10,fig. 3 One specimen (SH-PNUICO8445) Astraeomorpha confusa Astraeomorpha confusa Figures 10Cand10D Thamnasterioid, fungiform; calices fungiform; Thamnasterioid, Late Triassic, Rhaetian, Howz- colony Hof 8, fig. 10 1–3 - S/1 mm S c-c d (WINKLER,1861) : WINKLER,p. 488,pl. WINKLER:FRECH, (WINKLER): (WINKLER): http://dx.doi.org/10.18268/BSGM2020v72n2a090619 (WINKLER): WINKLER: Boletín de la Sociedad Geológica Mexicana Mexicana Geológica Sociedad la de Boletín et - - HPUC84 63 021722 7 6 24-25 5-6 24 5 2.1 25 2 2.1×2.2 24-25 2.2 2×1.7 16 2.5 2.3×2.5 20 45 2.5×2 31 32 21 52 21 36 SH-PNUICO8446 16 38 SH-PNUICO8445 20 SH-PNUICO8443 SH-PNUICO8442 Number 1980 1980 two and B) (A, 36 p. on text-fig. 37, p. FRECH, 1890 1861 Type species: e-Khan Member, Dizlu andHowz-e-Khan Occurrence: wallthin. than theS1.Endothecadissepimental, arenear to thecollumela; the S2septa shorter into 2sizearranged orders. approach S1septa The faces granular. the septal Septa of Ornamentation New corallites appear between adult corallites. donotcontinueSepta tothecolumella. form. in andovalor rounded, columella thin, styliform diameters.in mm Calicesare shallow, polygonal be large but hemispherical coloniesare about 30 or hemisphericalinshape. Lamellate colonies may Description: Measurements (inmm): 8446) from Biostrome 2,Section C, Dizluarea. Abad area, andtwo(SH-PNUICO8444, specimens 8443, 8445)fromBiostrome 2,Section B, Bagher Material: 1989 KRISTIAN-TOLMANN, TOLLMANN and Chondrocoenia schafhaeutli RONIEWICZ, p. 104,pl.33,figs. 1–3,5 Genus Prionastraea Stephanocoenia schafhaeutli Stephanocoenia HAMEDANI, p. 169,pl.1,figs. 1–4 GCDof colony BARI-DARYAN, p. 41,pl.5,fig. 1 Chondrocoenia schafhaeutli Stephanocoenia schafhäutli Stephanocoenia Astrocoenia schafhaeutli Tree(SH-PNUICO8442, specimens Chondrocoenia LCDof colony Colonies of plocoid type, plocoid lamellate Coloniesof Late Triassic, Rhaetian, Howz- Prionastraea schafhaeutli Figures 10Eto10G ? text-figs. on p. 37 Schafhaeutli pl. 8,fig. 11 colony Hof 1861 RONIEWICZ, 1989 / 72(2)2020 (WINKLER,1861) -cSS/1mm S c-c d : WINKLER,p. 488, FRECH:SENOW (WINKLER): (WINKLER): WINKLER, WINKLER: 19 19 -

Late Triassic corals, Nayband Formation, southwest of Ardestan SYSTEMATIC PALEONTOLOGY Late Triassic corals, Nayband Formation, southwest of Ardestan SYSTEMATIC PALEONTOLOGY Biostrome 2, Section C,Dizluarea. Colonyintransversesection. Biostrome 2,Section Crassistella juvavica I) (H, view. in calicular Colony Dizlu area. C, Section 2, Biostrome SH-PNUICO8446, (G) view. in calicular colony of Detail (F) view. calicular proximal view.(E–G) Astraeomorpha confusa crassisepta Figure 10 20 20 / Boletín de la Sociedad Geológica Mexicana Geológica Sociedad la de Boletín Scleractinian coralsfrom theNayband Scleractinian Formation,southwest ofArdestan,Central Iran, Iran. (A,B) (Reuss, 1854). SH-PNUICO8425, Biostrome 1, Section C,Dizlu area.(A)Colonyincalicular view. (B)Colonyinsideview.(C,D) Biostrome 1,Section (Reuss,1854).SH-PNUICO8425, (Frech, 1890). (H) SH-PNUICO8436, Biostrome 1, Section C,Dizluarea.Colonyincalicular view.(I) SH-PNUICO8438, Biostrome1,Section (Frech,1890).(H) SH-PNUICO8436, Chondrocoenia schafhaeutli (Winkler,1861). SH-PNUICO8445, Biostrome 2, Section C,Dizlu area.(C)Colonyincalicularview.(D) (Winkler,1861). Biostrome2, Section SH-PNUICO8445, http://dx.doi.org/10.18268/BSGM2020v72n2a090619 (WINKLER,1861). E, F. SH-PNUICO8444, Biostrome 2, Section C, Dizluarea.(E)Colonyin (WINKLER,1861). Biostrome 2,Section E,F.SH-PNUICO8444, /72(2)2020 Astraeomorpha Article A090619 Article A090619 SH- PNUICO8465 H NIO443 65566454 4 4-5 48 40 4.5 5-6 4-5 5.5×6.6 3.5-4.5 5 31 42-44 3×3.6 16 4 2.5-3.8 3.5-4.5 40 4.5×3.5 30 3×4 44-46 2 3.5-4 16 32 4-5 4.4×5 21 33 50 4.8×4.5 36 13 SH- PNUICO8464 60 19 60 SH- PNUICO8463 20 69 SH- PNUICO8462 40 SH- PNUICO8461 22.3 49 SH- PNUICO8460 PNUICO8459 SH- Number juvenile adult 1858-1860 Type species: North America. to Stanley (1986), it is known from the Norian of Iran by Kristian-Tollmann et al. (1980). According species wasdescribed fromThe the Rhaetian of Afghanistan. the Pamirsand Norian–Rhaetian of Carpathians; the (Lombardy); Alps Rhaetian of the Southern and Voralpe near to Altenmarkt; Rhaetian of KothalpKesselwand-Rohrmoos, Kössen Bedsof Fischerwise and eous Alps:Zlambach Beds of Calcar the Northern presentthe Rhaetian in of According toRoniewiczis (1989), thespecies Central Iran. Ardestan, Member, Bagher Abad, southwest of tion C,Dizluarea. Measurements (inmm): (SH-PNUICO8462, 8465) from Biostrome 2,Sec Section B, Bagher Abad area. Two specimens 8460, 8461,8463,8464)from Biostrome 2, Material: MELNIKOVA, p. 74, pl. 19, figs. 2–4, pl. 24, fig. 4 Family Pamiroseriidae MELNIKOVA, 1984 Genus 2001 1971 Pamiroseris meriani GCDof colony MELNIKOVA, p. 34,pl.2,fig. 2 62 9545635 5 55 6.3 5×4.5 19 20 26 Five specimens (SH- PNUICO 8459, Thamnastraea meriani Thamnastraea Pamiroseris meriani Pamiroseris meriani Pamiroseris LCDof colony 108, pl.26,figs. 3–6 Thamnastraea meriani Thamnastraea Figures 9Eand9F colony Hof MELNIKOVA, 1971 1860 (STOPPANI, 1860) -cSS/1mm S c-c d (STOPPANI): (STOPPANI): http://dx.doi.org/10.18268/BSGM2020v72n2a090619 : STOPPANI, p. . 04 40 4.2 STOPPANI, Boletín de la Sociedad Geológica Mexicana Mexicana Geológica Sociedad la de Boletín - - HPUC85 36 389563-43 3 32-44 3 5.6 18-23 4-5 22-26 3.8 8×9 28 5 3 4.8 4×3.4 33 30-37 5-6 3 3.5×3.3 21 20 5.2 5×4.5 63 3 1 3 28 5.4 28 4 3.9×4.5 40 63 28-30 4.8 4.5×5 15 27 28 36 28 3.5 SH-PNUICO8455 38 4.5×4.3 25 19 4.2 SH-PNUICO8454 33 3.8×5.3 19 6 50 4.8×5 SH-PNUICO8453 43 27 42 7×6.7 SH-PNUICO8452 31 17 48 SH-PNUICO8451 36 30 38 SH-PNUICO8450 26 36 SH-PNUICO8449 43 28 SH-PNUICO8448 44 SH-PNUICO8447 SH-PNUICO8446 Number 1861 theAlpsandPamirs (Roniewicz, 1989). tian of specieswasdescribed fromIran. This the Rhae Ardestan, Central ber, Bagher Abad, southwest of Khan Member, DizluandHowz-e-Khan Mem Occurrence: Columellawell septa. developed. distal edgeof faces,wall, granulationseptal on denticulation on witha small trunk, shallow calices,fungi-form thick Description: ments (inmm): from Biostrome 2,Section C, Dizluarea.Measure area. Six specimens(SH-PNUICO8450-8455) 8449) from Biostrome 2,Section B, Bagher Abad Material: 1989 MELNIKOVA, p. 63, pl. 25, figs. 1, 2, pl. 26, fig. 1 KRISTAN-TOLMANN, TOLLMANN and 1980 1979 1890 FRECH, p. 60,text-fig. on p. 61,pl.16,figs. 1–15 Pamiroseris rectilamellosa 1986 Figures 9A,9BandFigures 14Ato14F RONIEWICZ, p. 111,pl.34,figs. 3–5 Thamnastraea rectilamellosa Thamnastraea HAMEDANI, p. 169,pl.2,figs. 1–3 GCDof Thamnasteria rectilamellosa Thamnasteria colony Thamnastraea rectilamellosa Thamnastraea Thamnasteria rectilamellosa Thamnasteria Pamiroseris rectilamellosa Pamiroseris rectilamellosa SCHÄFER, p. 46,pl.1,fig. 3 Four specimens(SH- PNUICO 8446- LCDof colony Thamnasterioid colonies, flat or flat colonies, Thamnasterioid Late Triassic,Rhaetian, Howz-e- 487, pl.8,fig. 7 colony Hof / 72(2)2020 (WINKLER,1861) -cSS/1mm S c-c d (WINKLER): : WINKLER, p. (WINKLER): (WINKLER): WINKLER: WINKLER: 21 21 - - -

Late Triassic corals, Nayband Formation, southwest of Ardestan SYSTEMATIC PALEONTOLOGY Late Triassic corals, Nayband Formation, southwest of Ardestan SYSTEMATIC PALEONTOLOGY 22 22 onies, corallitesare series but in arranged at the col of margin the At granules. flat and circular are faces arethe S2 septa short. Septal covered by into 2–3orders. are S1septa longandthick, The trabecular denticles are present. arranged Septa edge,On theinternal or straightform. in sigmoid Description: et al. IranbyKristian-Tollmann from theRhaetian of Pamirs (Roniewicz, species isknown 1989). The the Caucasus and and inthe Norian–Rhaetian of the Tatra Mountains,Kothalp; the Rhaetian of in the Voralpe near Altenmarkt,feln, Rhaetian of the Donnerkogels, Dachsteinkalkand the Hoch Fischerwise, Dachsteinkalk of Zlambach Beds of berg, (Frech, Oedalm, Zlambachgraben 1890),in Calcareousin HallstätterSalz Alps: Northern the in the speciesiscommon Rhaetian of The Iran. Ardestan, Central ber, Bagher Abad, southwest of Khan Member, DizluandHowz-e-Khan Mem Occurrence: tal spaces(Figures 14Ato14F). strong red-luminescence) inintersep were formed altered, secondary, Mn-enriched(showing deposits structureAlthough septal is clearly diagenetically (strong deposits redto interseptal luminescence). regionsthe septal are much darker incomparison in the skeleton. However, incathodoluminescence, recognizable are centers/fibers) (calcification ture microstruc primary vex. Nodistinctfeatures of corallite it iscon depressed while at the center of are organized.the periaxial In ring, endotheca is newcorallites apparatusesces andthenseptal of homogeneous. appear between New centers cali colonies, is thecorallite arrangement center of / 1967 Boletín de la Sociedad Geológica Mexicana Geológica Sociedad la de Boletín Pamiroseris multiseptata (1980). 1975 MELNIKOVA, p. 131,pl. 30, figs. 1,2 2001 Fungiastraea multiseptata Pamiroseris multiseptatum Pamiroseris multiseptata Late Triassic,Rhaetian, Howz-e- Thamnasteroid colonies, are septa Figures 9Cand9D pl.2,fig. 3 (MELNIKOVA, 1967) : Melnikova, p. 24, Melnikova: http://dx.doi.org/10.18268/BSGM2020v72n2a090619 Melnikova: ------/72(2)2020 SH-PNUICO 8457 HPUC85 35 2465754 4-5 49 5 4.6×5.7 22 50 63 SH-PNUICO8458 SH-PNUICO8456 Number juvenile juvenile ments (inmm): Biostrome 2, Section C, Dizlu area. Measure Two specimens(SH-PNUICO8456, 8458) from fromBiostrome 2,Section B, Bagher Abad area. Material: Abad area. Measurements (inmm): 8440, 8441)fromBiostrome 2,Section B, Bagher tion C,Dizluarea. Two(SH-PNUICO specimens (SH-PNUICO8438, 8439) from Biostrome 2,Sec -8437) from Biostrome 1andtwo specimens Material: 1989 1980 1890 Pamirs (Roniewicz, 1989). the speciesisknown fromThe the Rhaetian of Ardestan, CentralIran. west of Khan Member, DizluandBagher Abad, south Occurrence: in andthey areor straight form, larger than corallites sigmoid wall. Corallites thick have many of septa withasmalltrunk,shallow calices,fungi-form and Description: Type species: Pamiroseris rectilamellosa MELNIKOVA, p. 75,pl.19,fig. 5,pl.22, p. 113,pl.34,figs. 1,2,pl.35,figs. 1,2 Stephanocoenia juvavica Stephanocoenia Genus Crassistella juvavica Actinastraea juvavica GCDof colony BARI-DARYAN, p. 42,pl.5,fig. 3 Crassistella juvavica 23 83.8×5.5 28 6×5 32 15 62 24 28 One specimen (SH-PNUICO8457) Four specimens (SH-PNUICO8434 Crassistella LCDof colony Stephanocoenia juvavica Stephanocoenia Thamnasterioid colonies, flat or flat colonies, Thamnasterioid Late Triassic,Rhaetian, Howz-e- Figures 10Hand10I fig. 3,pl.24,fig. 3 text-fig. on p. 38 colony Hof RONIEWICZ, 1989 FRECH:RONIEWICZ, . (FRECH, 1890) : FRECH,p. 38, the left (FRECH): SENOW -cSS/1mm S c-c d . 60 5.5 54 5.5 . 50 3.5 50 4.2 FRECH,1890 Article A090619 5 5 - - - - Article A090619 HPUC84 12 2192163 7-8 6-7 30 7-8 22 6-7 1.6 25 6-7 1.4 1.9×2 22 6 1.9×1.7 1.7 24 12 6 1.4 1.7×2.6 8 24-25 2.1 2.1×2.6 23 6-7 22 22 1.9 2.3×2 28 5 22 31 1.3 1.7×2 33 12 37 SH-PNUICO8441 25 1.6×1.9 1.9 22 35 SH-PNUICO8440 22 2.2×1.6 12 36 SH-PNUICO8439 24 18 30 SH-PNUICO8438 31 36 SH-PNUICO8437 38 38 SH-PNUICO8436 39 SH-PNUICO8435 SH-PNUICO8434 Number Occurrence: thecorallite. center of walled vesicles, sub-horizontal or convex inthe abundant, thin- of Endotheca isformed septa. the S1 the length of of are S3septa half The aresepta subequal,theycontinue tocolumella. and the S2 the S1 septa orders, blades septal of size 2–3 into differentiated wedge-like, are Septa with round faces are granules.Septal ornamented increased byinto divisiontwo juvenile corallites. deep, columella small andwall is thick.Corallites nies. Mamillae grow uplikecolumns. Calices are colo mamillate and massive or lamellate and flat Description: 1890 Type species: 1989). the Pamirs and NE Asia (Roniewicz, Rhaetian of the Gruber Reef; dubious inFischerwise; Norian– and Kesselwand-Rohrmoos; Schneckengraben of Calcareous Alps:inthe Zlambach Beds Northern the known speciesis in the Rhaetian of The Ardestan, CentralIran. southwest of Rhaetian, Howz-e-Khan Member, Bagher Abad, Bidestan and Howz-e-Khan Members, Dizluand fig. on p. 49,pl. 12, fig. 3,pl.15, figs. 17–23 1972 Stylophyllopsis polyactis Family Stylophyllidae FRECH,1890 GCDof Stylophyllopsis polyactis colony Figures 11Ato11Iand12A to12F Genus (designated by DIENER,1921). Stylophyllopsis polyactis LCDof colony Stylophyllopsis polyactis p. 239, text-fig. 14a–c The speciesischaracterized The by Late Triassic, Norian–Rhaetian, Stylophyllopsis colony Hof : FRECH, p. 48, text- FRECH,1890 FRECH,1890 -cSS/1mm S c-c d FRECH:CUIF, http://dx.doi.org/10.18268/BSGM2020v72n2a090619 FRECH,1890 Boletín de la Sociedad Geológica Mexicana Mexicana Geológica Sociedad la de Boletín - HPUC80 61 11010-11 6 160 9 188 9 21 168 7 13 111 9 14 19 124 8 38 7 16 96 8 20 12 3-4 38 176 SH-PNUICO8400 21 8 12 161 24 SH-PNUICO8393 24 72 31 26 SH-PNUICO8388 22 23 96 27 SH-PNUICO8389 18 9 25 SH-PNUICO8394 38 19 24 15 SH-PNUICO8398 38 23 SH-PNUICO8395 16 SH-PNUICO8391 21 SH-PNUICO8385 SH-PNUICO8383 ubrGDLDHSS/5 S H LCD GCD Number mm): 2, Section B, Bagher Abad area.Measurements (in (SH-PNUICO8388, imens 8395) fromBiostrome Biostrome 2, Section C, Dizlu area and two spec 8385, 8389, 8391, 8393, 8394, 8398, 8400) from Material: 1890 1989). and HallstätterSalzberg (Roniewicz,engraben Fischerwise, Kesselwand–Rohrmoos, Schneck of Calcareous Alps:inthe Zlambach Beds Northern the in the speciesiscommon Rhaetian of The Ardestan, CentralIran. Abad, southwest of Member,Khan and Howz-e-Khan Dizlu Bagher Occurrence: all.Wall thick. are S4septa theshortestof The continue theS1 andS2 septa tothe axial cavity.of into 3–4sizethin, arranged orders. blades Septal Septa spines. septal with filled cavity Axial calice. to conicalinshape, withcircular andshallow Description: RONIEWICZ, p. 124,pl.36,figs. 6,7, Stylophyllopsis rudis Stylophyllopsis rudis 1853 Figures 13Ato13Gand 15A to15F 1989 Eight specimens (SH-PNUICO8383, Eightspecimens pl. 37,fig. 1,pl.38, figs. 8,9 Fungia rudis Stylophyllopsis polyactis 50, pl.12,figs. 1, 4-14 Corallum solitary, sub-cylindrical Late Triassic,Rhaetian, Howz-e- -EMMRICH, p. 378 (EMMRICH,1853) EMMRICH: FRECH,p. / 72(2)2020 FRECH: 23 23 - -

Late Triassic corals, Nayband Formation, southwest of Ardestan SYSTEMATIC PALEONTOLOGY Late Triassic corals, Nayband Formation, southwest of Ardestan SYSTEMATIC PALENTOLOGY Corallum incalicular C,Dizluarea. view.(H)Corallum Biostrome2,Section inproximal view.(I) SH-PNUICO8389, calicular view.(E) Septalspinesincorallum. (F)Corallum C,Dizluarea. (G) inproximalview.(G,H)SH-PNUICO8398, Biostrome2,Section Corallum inproximalview,septa C,Dizluarea.(D)Corallum subequal in Biostrome 2,Section inthickness,thin.(D–F)SH-PNUICO8391, C,Dizluarea.(A)Corallum in calicular Biostrome 2,Section view.(B)Corallum(French, 1890).(A–C)SH-PNUICO8383, insideview.(C) Figure 11 24 24 / Boletín de la Sociedad Geológica Mexicana Geológica Sociedad la de Boletín Scleractinian corals from the Nayband Formation, southwest of Ardestan, Central Iran, Iran. (A–I)Iran. Iran, CentralArdestan,of the Naybandfromsouthwest Formation, corals Scleractinian http://dx.doi.org/10.18268/BSGM2020v72n2a090619 /72(2)2020 Stylophyllopsis polyactis Article A090619

Article A090619 transverse section. (F) Corallum in side view. (G, H) (G, view. side in Corallum (F) section. transverse Corallum inproximalview.(D–F)SH-PNUICO8394, Biostrome2,Section C,Dizluarea.(D)Corallum incalicularview. (E)Corallum in C,Dizluarea.(A)Corallum in calicular Biostrome 2,Section view.(B)Corallum(French, 1890).(A–C)SH-PNUICO8393, insideview.(C) Figure 12 S3 red, S4blue. yellow, S2 white, S1 colors: following in the dots with fourmarked ofsize orders septa (H) view. in calicular Colony H) (G, Dizlu area. C, Scleractinian coralsfromNayband Formation,southwestof Ardestan,Central Scleractinian Iran, Iran. (A-F) http://dx.doi.org/10.18268/BSGM2020v72n2a090619 Boletín de la Sociedad Geológica Mexicana Mexicana Geológica Sociedad la de Boletín Pamirophyllum iranicum (Melnikova, 1972) SH-PNUICO8423, Biostrome 1, Section Section 1, Biostrome SH-PNUICO8423, 1972) (Melnikova, / 72(2)2020 Stylophyllopsis polyactis 25 25

Late Triassic corals, Nayband Formation, southwest of Ardestan SYSTEMATIC PALEONTOLOGY Late Triassic corals, Nayband Formation, southwest of Ardestan SYSTEMATIC PALENTOLOGY 26 26 HPUC87 11 29 9 10 96 15 96 8-9 12 96 10 40 96 8 17 11 87 8 18 36 21 56 8 13 27 21 80 SH-PNUICO8378 10 15 14 14 72 SH-PNUICO8382 11 14 32 17 72 SH-PNUICO8399 9 18 10 16 72 SH-PNUICO8397 15 42 20 72 SH-PNUICO8396 10 45 27 SH-PNUICO8390 13 11 13 SH-PNUICO8387 15 20 SH-PNUICO8392 15 17 SH-PNUICO8380 17 SH-PNUICO8379 SH-PNUICO8384 ubrGDLDHSS/5mm S H LCD GCD Number 1989 TOLLMANN, TOLLMANN andHAMEDANI, 1980 Bidestan and Howz-e-Khan Members, Dizluand Occurrence: 15A to15F). (Figures red-luminescence strong their by firmed original microstructure) that is also con of signs wall thick. Coralla are entirely recrystallized (no the last order are veryshort, of the septa septa; shorter and slightly thinner than the S1 S2 septa continuingS1 septa near to the axial cavity; the into 4sizeRadial elements arranged orders:the (SH-PNUICO8392). morphotype thick-septal (SH-PNUICO8387) and morphotype thin-septal in the spines axial cavity.septal species has This calice subcircular and shallow, in form, horn-like Description: Measurements (inmm): fromBiostrome 2,Section B, Bagher Abad area. specimens (SH-PNUICO8387, 8399and8382) strome 2,Section C, from Dizluarea, and three 8380, 8384,8390,8392,8396,8397)fromBio Biostrome 1,seven (SH-PNUICO8379, specimens Material: / Boletín de la Sociedad Geológica Mexicana Geológica Sociedad la de Boletín RONIEWICZ, p. 126,pl.39,figs. 2–6, Stylophyllopsis mojsvari Stylophyllopsis rudis One specimen (SH-PNUICO8378)fromOne specimen pl. 40,figs. 1,2,pl.42,fig. 7 Corallum solitary, conical to Late Triassic, Norian–Rhaetian, p. 173,pl.5,fig. 3 FRECH:KRISTAN EMMRICH: http://dx.doi.org/10.18268/BSGM2020v72n2a090619 - - /72(2)2020 HPUC88 71 2S-2 28 72 S1-S2: 13 144 S1-S3: 32 29 15 17 27 18 SH-PNUICO8381 SH-PNUICO8386 ubrGDLDHSS/5mm S H LCD GCD Number (SH-PNUICO8386 and 8381) from Biostrome 2, Material: Iran (Kristian-Tollmann species wasdescribed fromThe the Rhaetian of the Kössen Beds from Kothalp (Roniewicz, 1989). Fischerwise, Kesselwand-Rohrmoos, andin of In the Alps, it is commoninthe Zlambach Beds Calcareous Alps,Northern Iran,andthe Pamirs. the known speciesis in the Rhaetian of The tan, CentralIran. Ardes Howz-e-Khan Bagher Abad, southwest of Central Iran. Ardestan, e-Khan Member, Dizlu, southwest of Occurrence: thelatter species. of the septa large much calices andsepta exceeding in number whichit resembles Rhaetian Remarks: flattened calice.deformed, specimenSH-PNUICO8381 showsThe a short andvery thin.Wall thick. S4 are S1, the septa the septa the length of of half are the S3septa abouta shorter than theS1septa, S1 continuesepta tothe center, are the S2 septa the into 3–4sizearranged orders. of spines Septal extendspines to the axial cavity. very Septa thin, in shape, calicecircular, Septal slightlydeepened. Description: Dizlu area.Measurements (inmm): Pamirophyllum iranicum Stylophyllopsis Genus Two well preserved specimens The coral, having The spines in thinseptal Figures 12G and12H Pamirophyllum Corallum solitary, sub-cylindrical Late Triassic, Rhaetian, Howz- Figures 13Hto13L Roniewicz, 1990 cf. lindstroemi MELNIKOVA, 1972 et al. Melnikova and , 1980). FRECH,1890 S. lindstroemi Article A090619 , shows - Article A090619 (F) Corallum incalicular view.(G)Corallum inproximal view. (H–L) calicular and sideviews.(E)Corallum insideview,thin—septal morphotype. C,Dizluarea. Biostrome 1,Section (F,G)SH-PNUICO8378, B,Bagher Abadarea.(C)Corallumseptal morphotype. Biostrome2,Section (C–E) SH-PNUICO8387, incalicularview.(D) Corallum in C, Dizluarea.(A)Corallum incalicular view.(B)Corallum Biostrome 2,Section insideview, thick— FRECH, 1890.(A,B)SH-PNUICO8392, Figure 13 Biostrome 2, Section C,Dizluarea. (J)Corallum incalicular view.(K)Corallum (L)Biostrome 2,Section Corallum intransversesection. insideview. C,Dizluarea. (H)CorallumBiostrome 2,Section incalicular view.(I) Corallum insideview,septaarevery thin.(J–L)SH-PNUICO8386, Scleractinian corals from the Nayband Formation, southwest of Ardestan, Central Iran, Iran. (A–G) Iran.Iran,Central Ardestan, of southwest NaybandFormation, the from corals Scleractinian http://dx.doi.org/10.18268/BSGM2020v72n2a090619 Boletín de la Sociedad Geológica Mexicana Mexicana Geológica Sociedad la de Boletín Stylophyllopsis cf. lindstroemi Frech, 1890.(H,I) SH-PNUICO8381, / 72(2)2020 Stylophyllopsis rudis 27 27

Late Triassic corals, Nayband Formation, southwest of Ardestan SYSTEMATIC PALENTOLOGY Late Triassic corals, Nayband Formation, southwest of Ardestan SYSTEMATIC PALENTOLOGY Mn-enriched (showing deposits strong red-luminescence) were formedininterseptal spaces. secondary,altered, diagenetically clearly is Althoughluminescence).redstructure (strong septal deposits interseptal in comparisonto (calcification centers/fibers) arerecognizableintheskeleton.However, incathodoluminescence, theseptal regionsaremuchdarker (D, F)Enlargements ofthin-sectioned corallain cathodoluminescence microscopeview.No distinctfeaturesofprimary microstructure optical microscope.(C–F)Enlargements ofthin-sectioned coralla.(C,E)Enlargements of thin-sectioned corallainopticalmicroscopeview. in corallum Thin-sectioned (B) view. in calicular Colony (A) area. BagherAbad B, Section. 2, SH-PNUICO8449,Biostrome (Winkler,1861), Figure 14 28 28 / Boletín de la Sociedad Geológica Mexicana Geológica Sociedad la de Boletín Scleractinian corals from the Nayband Formation, southwest of Ardestan, Central Iran, Iran. (A–F) Iran. Iran, Central Ardestan, of southwest NaybandFormation, the from corals Scleractinian http://dx.doi.org/10.18268/BSGM2020v72n2a090619 /72(2)2020 Pamiroseris rectilamellosa Article A090619

Article A090619 longitudinally). (C–F) Enlargements of thin-sectioned coralla. (C,E)Enlargements of thin-sectioned coralla inopticalview.(D,F) (B) Transverselythin-sectioned corallum (juvenilespecimenattached totheadult corallum (upper partofthepicture)sectioned rudis C,Dizluarea. (Emmrich, (A)Corallum and Biostrome2,Section in calicular(left) 1853),SH-PNUICO8379, lateral(right) views. Figure 15 microstructure), whichisalsoconfirmed bytheir strong red-luminescence. original of signs (norecrystallized entirely are Coralla view. microscope in cathodoluminescence coralla thin-sectioned ofEnlargements Scleractinian coralsfromthe Naybandof Ardestan,Central Scleractinian Iran, Formation,southwest Iran. (A–F) http://dx.doi.org/10.18268/BSGM2020v72n2a090619 Boletín de la Sociedad Geológica Mexicana Mexicana Geológica Sociedad la de Boletín / 72(2)2020 Solitary Stylophyllopsis 29 29

Late Triassic corals, Nayband Formation, southwest of Ardestan SYSTEMATIC PALENTOLOGY SYSTEMATIC PALENTOLOGY/ CONCLUSIONS Late Triassic corals, Nayband Formation, southwest of Ardestan / ACKNOWLEDGEMENTS / REFERENCES 5. Conclusions 30 30 SH-PNUICO8423 SH-PNUICO8423 Number juvenile adult 1972 although showsome specimens weaker (Figures altered (no original microstructure is preserved) all collected corals are diagenetically Skeletons of which characterize the Norian–Rhaetian stage. actinian coralswere recognized inthese members, scler limestone. Fourteen genera and 24speciesof reefal Bidestan and Howz-e-Khan form members Howz-e-Sheikh, Howz-e-Khan, andQadir. Bidestan, The Gelkan, members: five includes estan Ard NaybandThe Formation in the southwest of Nayband region, CentralIran. theShurabe-Nagi, fromcies the upperNorianof Remarks: Central Iran. Ardestan, Bidestan Member,southwest Dizlu, of Occurrence: surface iscovered by wall. thin,epithecal in the order. are Dissepiments small. Corallum ordersare andshorter thinnerthantheS1septa continuesepta to the axial cavity, higher of septa theS1 into 3sizearranged orders. blades Septal of Septa columella. papillary with filled cavity Axial faces withsmallgranulations. edges; septal sharp oid, calicespolygonal withhighand anddeep Description: mm): fromBiostrome area. 1,Dizlu Measurements (in Material: / Boletín de la Sociedad Geológica Mexicana Geológica Sociedad la de Boletín 1990 1975 MELNIKOVA andRONIEWICZ, p. 5, MELNIKOVA, p. 77,pl.9:2,10:1 Stylophyllum iranicum colony 05 91×31 62+44)5 96(24+24+48) 12 19×23 19 30×51 Dof Pamirophyllum iranicum Stylophyllum iranicum One specimen (SH-PNUICO8423) Melnikova (1972) described this spe colony SH-PNUICO8423: colony ceri Late Triassic, Norian–Rhaetian, Hof pl. 21,fig. 1 pl. 10,fig. 3 11 21+83)5 72(18+18+36) 7 11×11 c-c d : MELNIKOVA, p. 59, MELNIKOVA: MELNIKOVA: http://dx.doi.org/10.18268/BSGM2020v72n2a090619 S S/2 mm - - - - /72(2)2020 Acknowledgements References Mexican the Geological Society) for granting the in-Chief, Dr. Antoni Camprubí (Bulletin of Sciences. I appreciate the Editor- Academyof Paleobiology, Polish theInstitute of of Stolarski Jarosław and Roniewicz Ewa thanks author The Calcareous AlpsandPamirs.Northern Formation resemblance bears to corals from the copy. Scleractinian coralfaunafrom the Nayband cathodoluminescencelumienscence in micros Mn-induced of intensity different by expressed is stronger14) andsome (Figures 15)alteration that Clapp, C.,Shimer, H.W., Sutton 1911,The Brönnimann,P., Zadinetti, L., Bozorghnia, Bourne, G.C., , 1900, The Beauvais, L.,1981,Sur la taxinomie des Alloiteau, J., 1952,Madréporaires post- comments, andtheirhelpfuladvice. thisarticle, their constructive the manuscript of upgrade to effort valuable their for reviewers the UNAM) forhelp inthe publishing his process, and tor, Prof. FranciscoJ. Geology, Vega (Institute of for the publication, the Associatepermission Edi Vancouver Canada, Proceedings Island, of the Vancouver group,of Vancouver Island: 14(5), 7−16. 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Late Triassic corals, Nayband Formation, southwest of Ardestan REFERENCES