Hymenoptera: Chalcidoidea: Mymaridae) in India and Sri Lankapages with Descriptions of Eleven New Specieses

ISSN 0044-5088 Original Contributions to Zoology, founded in 1888 Ed. H.F. Paulus, Vienna

Volume 165

Prince Tarique Anwar, Shahid Bin Zeya and Kamalanathan Veenakumari Fairyfly genus Camptoptera Foerster (Hymenoptera: Chalcidoidea: Mymaridae) in India and Sri Lankapages with descriptions of eleven new specieses

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104. Karl Strenzke: Untersuchungen über die Tiergemein- 121. Hajo Schmidt: Prodromus zu einer Monographie der me- schaften des Bodens: Die Oribatiden und ihre Synusien diterranen Aktinien. in den Böden Nordddeutschlands. 2 Lfg. 1972. IV , 146 Seiten. ISBN 978-3-510-55005-0, € 50.– 1952. V , 173 Seiten. ArtNo. ES169010400, € 58.– 122. Georg Scheer; C. S. Gopinadha Pillai: Report on the 105. Edwin Möhn: Beiträge zur Systematik der Larven der Ito- Scleractinia from the Nicobar Islands. Results of the Xar- nididae (Cecidomyiidae, Diptera). 1. Teil: Porricondylinae ifa Expedition 1957/58 of the International Inst. for Sub- und Itonidinae Mitteleuropas. 2 Lfg. marine Research, Vaduz/Liechtenstein. 1955. VIII , 247 Seiten. ArtNo. ES169010500, € 95.– 1974. IV , 91 pages. ISBN 978-3-510-55006-7, 86.– 106. M. A. Basir: Oxyuroid Parasites of Arthropoda. A mono- 123. Anneliese Strenger; Herbert Schubart: Zur Kopfmor- graphic study. 1. Thelastomatidae. 2. Oxyuroidae. phologie der Ephemeridenlarven Ephemera danica. 1956. V, 79 pages. ArtNo. ES169010600, € 37.– Morphologische Grundlagen für die Klärung der Ver- 107. Otto Zur Strassen: Neue Beiträge zur Entwicklungs- wandtschaftsbeziehungen innerhalb der Milbenfamilie mechanik der Nematoden. Ameronothridae (Acari, Oribatei). 1959. V , 142 Seiten. ArtNo. ES169010700, € 67.– 1975. VI , 91 Seiten. ISBN 978-3-510-55007-4, € 91.– 108. Sigbert Wagener: Monographie der ostasiatischen For- 124. Maria-Anna Tiedemann; Franz Tiedemann: Vergleichend men der Gattung Melanargia Meigen (Lepidoptera, Saty- anatomische Untersuchungen an Schulter- und Beckengür- ridae). Erschienen in 3 Lieferungen. tel verschiedener südafrikanischer Skinkarten. Mit beson- 1961. VI , 222 Seiten, ArtNo. ES169010800, brosch., € 72.– derer Berücksichtigung von Reduktionserscheinungen. 1975. IV , 80 Seiten. ISBN 978-3-510-55008-1, € 81.– 109. August Wilhelm Steffan: Die Stammes- und Siedlungsge- schichte des Artenkreises Sacchiphantes viridis (Ratze- 125. Walter Sudhaus: Vergleichende Untersuchungen zur burg 1843), (Adelgidae, Aphidoidea). Phylogenie, Systematik, Ökologie, Biologie und Etholo- 1961. VIII , 113 Seiten. ArtNo. ES169010900, € 52.– gie der Rhabditidae (Nematoda). 1976. IV , 229 Seiten. ISBN 978-3-510-55009-8, € 147.– 110. Wolfgang Buchmann: Die Genitalanhänge mitteleu- ropäischer Dolichopodiden. 126. C.S. Gopinadha Pillai; Georg Scheer: Report on the 1961. V , 51 Seiten. ArtNo. ES169011000, € 37.– Stony Corals from the Maldive Archipelago. Results of the Xarifa Expedition 1957/58 of the International Institute 111. Gert M. Andres: Eine experimentelle Analyse der Ent- for Submarine Research, Vaduz, Liechtenstein. wicklung der larvalen Pigmentmuster von fünf Anuren- 1976. IV , 83 pages. ISBN 978-3-510-55010-4, € 86.– arten Bombina variegata L., Discoglossus pictus Otth, Hyla arborea L., Rana exculenta L. und Xenopus laevis 127. Anneliese Strenger; Albert Melbert; Gerhard H. Schmidt: Daudin. Zur Kopfmorphologie der Ephemeridenlarven Probosci- 1963. VII , 112 Seiten. ArtNo. ES169011100, € 49.– diplocia skorai. Sozialphänomene bei Heteropteren. 1977. VI , 53 Seiten. ISBN 978-3-510-55011-1, € 50.– 112. Inge Richter-Vollert: Untersuchungen zur Morphologie und Ökologie von Asphondylia sarothamni H. Lw. 128. Luitfried vonpages Salvini-Plawen: Antarktische und subark- 1964. V , 54 Seiten. ArtNo. ES169011200, € 35.– tische Solenogastres. Eine Monographie: 1898-1974. 113. M. L. Roonwal; Geeta Bose: Termite fauna of Rajasthan, (Lieferung 1). India. 1978. ISBN 978-3-510-55012-8, € 87.– 1964. VI , 58 pages. ArtNo. ES169011300, € 35.– Luitfried von Salvini-Plawen: Antarktische und suban- tarktische Solenogastres. Eine Monographie: 1898-1974. 114. Johannes Dönges: Der Lebenszyklus von Posthodiplos- (Lieferung 2). tomum brevicaudatum (Trematoda), eines Parasiten in 1979. ISBN 978-3-510-55013-5, € 90.– den Augen von Süßwasserfischen. 1965. V , 39 Seiten. ArtNo. ES169011400, € 27.– 115. 129. Hajo Schmidt; Dieter Zissler: Die Spermien der Antho- August W. Steffan: Evolution und Systematik der Adelgi- zoen und ihre phylogenetische Bedeutung. dae (Homoptera: Aphidina). 1979. VIII , 97 Seiten, ISBN 978-3-510-55015-9, € 84.– 130. Martin Baehr: Vergleichende Untersuchungen am Skelett 1968. XII , 139 Seiten. ArtNo. ES169011500, € 92.– und an der Coxalmuskulatur des Prothorax der Coleop- 116. Hermann Weber: Die Elefantenlaus Haematomyzus ele- tera. fantis Piaget 1869. Versuch einer konstruktionsmorphol- 1979. IV , 76 Seiten. ISBN 978-3-510-55016-6, € 65.– ogischen Analyse. Hrsg. und teilweise erg. von: Peter 131. Heinrich Schönmann: Zur Kopfmorphologie der Eph- Wenk. emeridenlarven. (Siphlonurus aestivalis Eaton und Lep- 1969. VI , 154 Seiten. ISBN 978-3-510-55000-5, € 99.90 erorus Goyí Peters). 117. Anneliese Strenger; Heinz Splechtna: Zur Kopfmorpholo- 1981. IV , 51 Seiten. ISBN 978-3-510-55017-3, € 45.– gie der Ephemeridenlarven Palingenia longicauda. Zur 132. Johann W. Wägele: Zur Phylogenie der Anthuridea Struktur und Gefäßversorgung der Reptilienniere. (Crustacea, Isopoda). Mit Beiträgen zur Lebensweise, 1970. 88 Seiten. ArtNo. ES169011700, € 63.– Morphologie, Anatomie und Taxonomie. 118. Wolfgang Scherzinger: Zum Aktionssystem des Sperlings- 1981. IV , 127 Seiten. ISBN 978-3-510-55018-0, € 91.– kauzes (Glaucidium passerinum L.). 1970. IV , 120 SampleSeiten. ISBN 978-3-510-55001-2, € 87.– 133. Georg Scheer; C. S. Gopinadha Pillai: Report on the stony corals from the Red Sea. 119. Ragnar Kinzelbach: Morphologische Befunde an Fächer- 1983. V , 198 pages. ISBN 978-3-510-55019-7, € 137.– flüglern und ihre phylogenetische Bedeutung (Insecta: Strepsiptera). 2 Teile. 134. V. B. Meyer-Rochow; K M. Tiang: The eye of Jasus Ed- 1971. XVII , 256 Seiten. ISBN 978-3-510-55903-9, € 164.– wardsii (Crustacea, Decapoda, Palinuridae). Electro- 1. Hälfte, 1971. XIII , 128 Seiten. ISBN 978-3-510-55002-9, € 86.– physiology, histology and behaviour. 2. Hälfte, 1971. IV , 128 Seiten. ISBN 978-3-510-55003-6, € 78.– 1984. IV , 61 pages. ISBN 978-3-510-55020-3, € 40.– 120. Gerhard Pretzmann: Die Pseudothelphusidae (Crusta- 135. R. Stützel: Anatomische und ultrastrukturelle Untersu- cea Brachyura). chungen an der Napfschnecke Patella L. Unter beson- 1972. IV , 182 Seiten. ISBN 978-3-510-55004-3, 87.– derer Berücksichtigung der Anpassung an den Lebens raum. 1984. VI , 54 Seiten. ISBN 978-3-510-55021-0, € 89.–

eschweizerbart_xxx Zoologica Original Contributions to Zoology

Founded 1888 by R. Leuckart, C. Chun, continued by W. Kükenthal, R. Hesse, W.E. Ankel

Edited by Hannes F. Paulus

Volume 165

Prince Tarique Anwar, Shahid Bin Zeya and Kamalanathan Veenakumari Fairyfly genus Camptoptera Foersterpages (Hymenoptera: Chalcidoidea: Mymaridae) in India and Sri Lanka with descriptions of eleven new species

With 241 figures

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Schweizerbart Science Publishers Stuttgart • 2020

eschweizerbart_xxx P.T. Anwar, S.B. Zeya & K. Veenakumari: Fairyfly genus Camptoptera Foerster (Hymenoptera: Chalcidoidea: Mymaridae) in India and Sri Lanka – with descriptions of eleven new species

Addresses of the authors: Prince Tarique Anwar (Corresponding author), Department of Zoology, Aligarh Muslim University, Aligarh 202 002, India, [email protected] Shahid Bin Zeya, Department of Zoology, Aligarh Muslim University, Aligarh 202 002, India, [email protected] Kamalanathan Veenakumari, National Bureau of Agricultural Insect Resources, Bengaluru 560 024, India

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eschweizerbart_xxx Zoologica Vol. 165, 1–89 Stuttgart, January 2020

Fairyfly genus Camptoptera Foerster (Hymenoptera: Chalcidoidea: Mymaridae) in India and Sri Lanka with descriptions of eleven new species Prince Tarique Anwar, Shahid Bin Zeya & Kamalanathan Veenakumari

With 241 figures

Abstract The Indian and already described Sri Lankan species of Camptoptera Foerster (Chalcidoidea: Mymaridae) are reviewed. Eleven species are described as new: C. alii Anwar & Zeya, sp. nov., C. anneckei Anwar & Zeya, sp. nov., C. bengalurensis Anwar & Zeya, sp. nov., C. fawnae Anwar & Zeya, sp. nov., C. huberi Anwar & Zeya, sp. nov., C. naseemi Anwar & Zeya, sp. nov., C. natalieae Anwar & Zeya, sp. nov., C. ogloblini Anwar & Zeya, sp. nov., C. scythe Anwar & Zeya, sp. nov., C. squama Anwar & Zeya, sp. nov. and C. usmanii Anwar & Zeya, sp. nov. Four species, C. fransciscae (Debauche), C. doptera Triapitsyn, C. okadomei Taguchi and C. papaveris Foerster, are recorded for the first time from India, albeit identifications of C. franciscae and C. okadomei are tentative. Camptoptera muiri (Perkins) and C. sakaii Taguchi recorded from India by Manickavasagam et al. (2011) and Rehmat & Anis (2014) are found to be misidentified and are described here as C. fawnae Anwar & Zeya, sp. nov. and C. ogloblini Anwar & Zeya, sp. nov., respectively. The other recorded species, C. dravida Subba Rao, from several Indian states by the same authors were incorrectly identified and are placed here with C. enocki (Howard). Males are described for the previously described females of C. assamensis Rehmat & Anis, C. dravida Subba Rao, C. franciscae and C. matcheta Subba Rao. The male paratype of C. dravida is here considered to belong to C. enocki (Howard), and the male paratype of C. matcheta is found to be an unidentified species of Eofoersteria Mathot. A key to females of 25 species from India and Sri Lanka is provided. Key words: Taxonomy, Chalcidoidea, fairyfly, India, Sri Lanka, biodiversity. pages

Contents Abstract ...... 1 Camptoptera kannada Subba Rao ...... 36 Introduction ...... 2 Camptoptera natalieae Anwar & Zeya, sp. nov...... 39 Material and methods...... 2 Camptoptera ?fransciscae (Debauche) ...... 42 Taxonomy ...... 3 Camptoptera dravida Subba Rao ...... 45 Genus Camptoptera Foerster, 1856 ...... 3 Camptoptera enocki (Howard) ...... 50 Key to species of Camptoptera Foerster in India and Camptoptera bengalurensis Anwar & Zeya, sp. nov...... 55 Sri Lanka, females ...... 4 Camptoptera ?okadomei Taguchi ...... 57 Synopsis of the species in India and Sri Lanka Camptoptera assamensis Rehmat & Anis ...... 59 (listed in the same order as keyed) ...... 6 Camptoptera squama Anwar & Zeya, sp. nov...... 63 Camptoptera serenellae Viggiani ...... 6 Camptoptera papaveris Foerster ...... 67 Camptoptera matcheta Subba Rao ...... 6 Camptoptera longifuniculata Viggiani ...... 71 Camptoptera protuberculata Viggiani ...... 14 Camptoptera doptera Triapitsyn ...... 71 Camptoptera ambrae Viggiani ...... 14 Camptoptera huberi Anwar & Zeya, sp. nov...... 75 Camptoptera tuberculata Viggiani ...... 18 Camptoptera brevifuniculata Subba Rao ...... 78 Camptoptera scythe Anwar & Zeya, sp. nov...... 18 Camptoptera alii Anwar & Zeya, sp. nov...... 83 Camptoptera anneckei Anwar & Zeya, sp. nov...... 22 Camptoptera usmanii Anwar & Zeya, sp. nov...... 83 Camptoptera ogloblini Anwar & Zeya, sp. nov...... 27 Acknowledgements ...... 87 Camptoptera naseemi Anwar & Zeya, sp. nov...... 31 References ...... 87 Camptoptera Samplefawnae Anwar & Zeya, sp. nov...... 33

eschweizerbart_xxx 2 P.T. Anwar, S.B. Zeya & K. Veenakumari

Introduction chi’s species from the Philippines and Taiwan, whose type material is lost, are not fully recognizable; for Members of Camptoptera Foerster (Hymenop- these, neotypes should be designated first based on tera: Mymaridae) are tiny parasitic wasps represented the fresh specimens collected at or near their type worldwide by 55 species (Noyes 2019). Triapitsyn localities (Triapitsyn 2018). The distribution in In- (2014) revised species of the entire Palaearctic region. dian states of all included species and their distribu- Ogloblin (1947) and Ogloblin & Annecke (1961) tion in other parts of the world (by country) are also described species of Camptoptera from Argentina and provided. All the studied taxa are illustrated. Many South Africa, respectively; Viggiani (1978) and Sub- more species likely remain to be described from the ba Rao (1989) described species from India and Vig- region. Males of most species are still unknown. giani (1978) also from Sri Lanka; Soyka (1953, 1961) They are similar to females except for sexually di- described many nominal species from Europe, mostly morphic features of genitalia and antenna (Triapit- synonymized by Triapitsyn (2014); Taguchi (1971, syn 2014), the number of flagellar segments in Ori- 1972, 1977) described 11 species from Japan, Taiwan ental species varies with either 7- or 10-segments. (Republic of China), and the Philippines (Bohol Is- The males we describe here are associated with fe- land). Although Taguchi’s species are valid his entire males of four previously described species. collection was lost, according to Triapitsyn (2014). Huber & Lin (1999) treated the Camptoptera-group genera and Huber (2011) showed that the type spe- Material and methods cies of Anaphes Haliday (at that time) was a species of Camptoptera. Manickavasagam et al. (2011) and The study is based on a large number of speci- Rehmat and Anis (2014), respectively, recorded C. mens of Camptoptera from several Indian states and muiri (Perkins) and C. sakaii Taguchi from India, but the union territory of Andaman & Nicobar Islands. we found their specimens to be misidentified and de- In addition to this, type and determined material scribe them here as new species. Triapitsyn (2017) housed in ZDAMU were also studied, and images of gave taxonomic notes on and illustrations of four spe- pages types located outside India were received on request cies of Camptoptera from India, and synonymized C. for providing brief diagnoses. kloptera Triapitsyn under C. matcheta Subba Rao. Collection and preservation techniques were Triapitsyn (2018) provided an annotated checklist of largely adapted from Noyes (1982). Slides were pre- Mymaridae in Taiwan, described C. jthuberi Triapit- pared following Noyes (1982) and Huber (2015), syn, recorded C. matcheta Subba Rao, and comment- with some modifications. After removing and mounted on other Camptoptera species occurring in Taiwan. ing the wings on a slide, the antenna and the head Kryger (1950: 47) stated [for the European fauna] was given a slight jerk to make them somewhat loose “until rearings have been performed there is no rea- but not completely broken off. The specimens were son to suppose that there is more than one species of then placed in KOH for 4–48 hours depending on Camptoptera, viz. C. papaveris Förster.” Triapitsyn the extant of sclerotization, i.e., highly sclerotized (2014) disproved that statement and further stated specimens were macerated for a longer duration and that the genus is speciose worldwide, particularly so light coloured specimens were macerated for about in the tropics from where many species remain to be 4–8 hours. The remaining steps followed were as giv- described. en by Noyes (1982), i.e., specimens were transferred A total of 26 species of Camptoptera are identi- sequentially to acetic acid, washed with distilled wa- fied and reviewed mainly from India and also from ter and then dehydrated in ascending grades of etha- Sri Lanka (types of the previously described taxa nol (all the steps for 10 minutes), when the specimen only). We provide a brief diagnosis of the genus, a is in absolute ethanol a drop of clove oil was added key to femalesSample of 25 species, redescriptions of the and after 10 more minutes the specimen was taken known species except diagnoses for those taxa where out of the cavity block and was placed for another 10 the types and determined material was not available, more minutes in a mixture of a drop of Canada bal- and descriptions of the new taxa. Providing a key to sam and clove oil in a cavity slide. Finally the speci- all species of the genus described from the Oriental men was transferred to a drop of Canada balsam on a region is currently not feasible because most of Tagu- microscope slide where it was dissected and its parts

eschweizerbart_xxx Fairyfly genus Camptoptera Foerster (Hymenoptera: Chalcidoidea: Mymaridae) 3

moved to separate drops in positions as indicated by Taxonomy Noyes (1982). The slides were dried for about two weeks in an incubator at 50 °C and then small cover Genus Camptoptera Foerster slips were placed onto small fragments of minute coverslips to prevent the specimens from being crushed (Figs 1–241) as the Canada balsam dries and shrinks. The slides were regularly monitored and the gaps created after Camptoptera Foerster, 1856: 119. Type species drying were refilled with Canada balsam. Camptoptera papaveris Foerster, by monotypy. Morphological terms mostly follow Zeya & Stichothrix Foerster, 1856: 117. Type species Sticho- Hayat (1995) and Gibson (1997). All measurements thrix cardui Foerster, by monotypy. Synonymy by were taken from the divisions of a linear scale of a Annecke & Doutt, 1961: 15. micrometer and converted into μm. Body lengths Pteroclisis Foerster, 1856: 144. Unnecessary replace- are taken from card-mounted specimen; all other ment name for Camptoptera, thought to have been measurements are from slide mounts made at 400× preoccupied by Camptopteris in Botany. magnification. Photographs of slide-mounted parts Eomymar Perkins, 1912: 26. Type species Eomymar were taken with a digital camera attached to a com- muiri Perkins, by monotypy. Synonymy by Huber & pound microscope “Leica DM 2500” and for habi- Lin, 1999: 27. tus view, insects as a whole were photographed un- Congolia Ghesquière, 1942: 320. Type species Con- der the stereo zoom binocular Nikon SMZ25. The golia sycophila Ghesquière, by original designation. photographs were then retouched using Adobe Pho- Synonymy by Debauche, 1949: 18. toshop® CS3. Photographic plates were prepared us- Sphegilla Debauche, 1948: 62. Type species Sphegilla ing Photoshop® 7.0. franciscae Debauche, by original designation. Syn- onymy by Yoshimoto, 1990: 32. Abbreviations used are: F = Funicle segment; mps = multi- Wertanekiella Soyka, 1961: 87. Type species Wer- porous plate sensillum or sensilla (= longitudinal sensilla of au- tanekiella brevicornis Soyka, by original designation. thors); MT = Malaise trap; SN = sweep net; YPT = yellow pan pages trap; PFT = pit fall trap; ANI = Andaman & Nicobar Islands. Synonymy under Sphegilla by Mathot, 1969: 2. Camptoptera (Zemicamptoptera) Ogloblin & An- The following acronyms are used for specimen necke, 1961: 24. Type species Camptoptera (Zemi- depositories: camptoptera) semialbata Ogloblin & Annecke, by original designation.; (as subgenus of Camptoptera). BMNH = Natural History Museum, London, U.K. BPBM = Bernice P. Bishop Museum, Honululu, Hawaii, Staneria Mathot, 1966: 214. Type species Staneria U.S.A. diademata Mathot, by original designation. Synony- ELNJ = Entomological Laboratory, Faculty of Agriculture, my by Huber & Lin, 1999: 27. Meijo University, Nagoya, Japan. INHS = Illinois Natural History Survey, Champaign, Illi- Diagnosis. Female. Body length less than 1000 μm. nois, USA Occiput with distinctly curved transverse suture LCXU = Life Science and Technology College, Xinjiang above foramen (Figs 11, 39) or usually with a medi- University, Urumqi, Xinjiang, China MHNG = Muséum d’Histoire Naturelle, Geneva, Switzer- an vertical suture above the transverse suture (Figs land. 94, 192, 227). Mandible variably shaped, with one NBAIR = National Bureau of Agricultural Insect Resources, pointed tooth (Figs 10, 45, 129, 189). Antenna with Bengaluru, Karnataka, India. 7 funicle segments; F2 usually ring-like (Fig. 9), NHMV = Naturhistorisches Museum, Vienna, Austria rarely quadrate to longer than broad (Figs 69, 75). USNM = United States National Museum of Natural His- Mesosoma with axilla slightly advanced into side tory, Washington D.C., USA. lobe of mesoscutum (Fig. 14); notaular line either ZDAMU = Insect Collections Department of Zoology, Aligarh MuslimSample University, Aligarh, Uttar Pradesh, India. well developed (Fig. 133) or reduced to less than an- ZIN = Zoological Institute, Russian Academy of Scien- terior one-third of mesoscutum (Fig. 150) or, rarely, ces, St. Petersburg, Russia. absent (Fig. 158); scutellum clearly divided in ante- ZLMU = Entomological Laboratory, Faculty of Agriculture, rior scutellum and frenum (posterior part of scutel- Meijo University, Shiogamaguchi, Tempaku-ku, lum, distinguished by different sculpture and, often, Nagoya, Japan. an anterior line of foveae); mesophragma not ex-

eschweizerbart_xxx 50 P.T. Anwar, S.B. Zeya & K. Veenakumari

fairly well with the original description and illustra- Additional materials examined (ZDAMU). 7 Θ, 2 tions given by Subba Rao (1989) and diagnosis and Η. INDIA: UTTARAKHAND: Ranikhet: Chauba- illustrations given by Triapitsyn (2017). tia, 3 Θ (each on slides under 3, 3, 4 coverslips), C. dravida was described by Subba Rao (1989) 27.x.2009, Coll. F.R. Khan; UTTAR PRADESH: from 4 females and 1 male collected by sweep net Aligarh, 1 Θ (on slide under 2 coverslips), Dept. of from Karnataka by J.S. Noyes in 1974. Triapitsyn Zoology, 4.iv.2011, Coll. T. Rehmat; 2 Θ, 2 Η (each (2017) remarked that the paratype male most likely on slide under 4, 2, 2, 3 coverslips), Shekha, 9. does not belong to this species because it has a broad- ix.2007; 1 Θ (on slide under 1 coverslip), Sharas- er fore wing with a complete median row of setae on wati, Chakra Bhandar, 3.x.2006 Coll. F.R. Khan. the disc. The male actually belongs to C. enocki (see (All determined as C. dravida Subba Rao by Rehmat comments under C. enocki). & Anis).

Camptoptera enocki Diagnosis. Camptoptera enocki (Howard) is distin- (Howard) guished from all other described Indian species of Camptoptera by having long seta between anterior (Figs 125–138) and posterior ocelli (Figs 129, 135). Camptoptera enocki is otherwise similar to C. Litus enocki Howard, in Howard & Ashmead, 1896: dravida Subba Rao (1989) in having more or less 643, females. Syntype, females, Ceylon [=Sri Lanka], same body length, and pattern of sculpture on head Pundaluoya (USNM), not examined. and thorax. It differs from C. dravida by having fol- Litus enocki Howard: Girault, 1911a: 132 (com- lowing features: head with transverse trabecula di- parison with Litus cynepsius); Girault, 1911b: 363 vided into three pieces (head with transverse trab- (descriptive note). ecula divided into five pieces in dravida); funicular Camptoptera enocki (Howard): Triapitsyn & Be- segments shorter and wider (longer and narrower in rezovskiy, 2004: 16 (combination under Camptop- dravida); frenumpages with transverse striations enclosed tera). in two cells medially (frenum with striations en- Camptoptera dravida Subba Rao: Subba Rao, closed in a single cell medially in dravida); wings 1989: 163, paratype male. Manickavasagam et al., broader (wings narrower in dravida); ovipositor 0.6– 2011: 395 (key), 399 (figs 7–10). Anis & Rehmat, 0.7× gaster (ovipositor 0.9× gaster in dravida). 2013: 3 (checklist). Manickavasagam & Rameshku- mar, 2013: 563 (checklist). Rehmat & Anis, 2014: Description. Female. Length 320–380 μm. Body 55 (list, key), 56 (females, males, Uttar Pradesh re- dark brown except gaster in distal half brown with cord, Uttarakhand record). [Misidentifications] yellowish tinge. Antenna pale brown. Wings subhya- line except area near venation and apex fumate. Legs, Material examined. 5 Θ, 1 Η. INDIA: HIMA- including coxae, pale brown. CHAL PRADESH: Shimla, 1 Θ (on slide under 4 Head (Fig. 129). Head, in frontal view, 1–1.6× coverslips, slide No. MYM.177), 2.viii.2014 (YPT), as broad as high, vertex with a pair long seta in be- Coll. K. Veenakumari. KARNATAKA: Mandya, 1 tween anterior and posterior ocelli, and with trans- Θ (on slide under 3 coverslips, slide No. MYM.73), verse striations, occiput with oblique striations, con- 3.v.2012 (MT), Coll. K. Veenakumari; ANDHRA verging anteriorly in middle. Mandibles pointed PRADESH: Visakhapatnam, Kailashpuram, 2 Θ meeting in middle. Antenna (Fig. 128) with scape (on slide under 4 coverslips, slide No. MYM.179), 2.2–2.6× as long as broad; pedicel 1.4–1.7× as long 2.ii.2014 (SN), Coll. M.T. Khan; UTTAR as broad, longer than all funicular segments individ- PRADESH: Aligarh, 2 Θ (each on slides under 4 ually except F3 which is sometimes subequal in coverslips,Sample slide Nos MYM.197, MYM.201), AMU length; funicular segments all longer than broad, ex- Camp., 12.iv.2014 (SN), Coll. F. Fatima; 1 Η (on cept F2 ring-like segment, F3–F7 decreasing in slide under 4 coverslips, slide No. MYM.178), length distad; clava 2.8–3.9× as long as broad, longer Akrabad 8.iii.2013 (SN), M.T. Khan. (NBAIR, than F5–F7 combined, with 4 mps. ZDAMU).

eschweizerbart_xxx Fairyfly genus Camptoptera Foerster (Hymenoptera: Chalcidoidea: Mymaridae) 51

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Figs 125–127. Camptoptera enocki (Howard), female (syntype, photographs courtesy of Michael W. Gates, USNM): 125, 126, slide; 127, habitus, lateral.

eschweizerbart_xxx 52 P.T. Anwar, S.B. Zeya & K. Veenakumari

pages

Sample

Figs 128–133. Camptoptera enocki (Howard), female: 128, antenna; 129, head, anterior; 130, fore wing; 131, hind wing; 132, mesosoma and gaster, with petiole; 133, mesosoma.

eschweizerbart_xxx Fairyfly genus Camptoptera Foerster (Hymenoptera: Chalcidoidea: Mymaridae) 53

Mesosoma (Figs 132, 133). Mesosoma 0.8–1.5× ments except F2 and F4 ring-like segments; F10 ob- as long as gaster; each side lobe of mesoscutum with long, longer than all flagellar segments individually, 1 seta anteriorly; mid lobe of mesoscutum with a and with a long spicule at apex. pair of setae in anterior margin; mesoscutum with Mesosoma (Fig. 138). Mesosoma as long as transverse striations, slightly convex anteriorly; no- gaster; setation and sculpture similar to females. Fore taular lines prominent, complete; scutellum with wing (Fig. 136) 13.8× as long as broad, disc with a transverse striations; each axilla with 1 seta, and with row of setae in middle extending to apex; longest longitudinal striations; sides of frenum with longitu- marginal seta 6.4× as long as maximum wing width. dinal striations, and medially with transverse and Hind wing (Fig. 137) 21× as long as broad; longest slightly curved striations, enclosed in two cells; pro- marginal seta 7.7× as long as maximum wing width. podeum with faint longitudinal striations and with Metasoma. Genitalia occupying nearly half peg-like sensilla, submedian carinae absent. Fore length of gaster. wing (Fig. 130) 12.6–14.5× as long as broad, disc Measurements (n=2): head width:height, 39:45; with one long seta slightly distal to venation; a single antennal segments length:width–radicle, 10:10; row of setae running from middle of the disc and scape, 38:16; pedicel, 33:23; F1, 30:15; F2, 3:8; F3, extending to apex, setae on anterior and posterior 30:15; F4, 3:6; F5, 30:13; F6, 35:13; F7, 35:13; F8, margins begin slightly proximal to middle of the 33:13; F9, 33:15; F10, 43:13; mesosoma, 153; me- disc; longest marginal seta 5–6.6× as long as maxi- soscutum, 40; scutellum, 15; frenum, 58; metano- mum wing width. Hind wing (Fig. 131) 21.2–24.8× tum, 5; propodeum, 33; fore wing length:width, as long as broad; longest marginal seta 7.1–8.8× as 380:28; longest marginal seta, 175; hind wing long as maximum wing width. length:width, 368:18; longest marginal seta, 135; Metasoma (Fig. 132). Petiole transverse, 0.6– protibia, 88; mesotibia, 113; mesobasitarsus, 23; 0.8× as long as broad; ovipositor slightly exserted metatibia, 110; petiole length:width, 25:28; gaster, beyond apex of gaster, 0.6–0.7× gaster, 0.7–0.9× 138; genitalia, 58. mesotibia and 0.7× –1.1× metatibia. pages Measurements (n=3): head width:height, 123– Host. Eriococcus sp. ? (Triapitsyn & Berezovskiy 143:90–103; antennal segments length:width–radi- 2004) cle, 5–8:8; scape, 33–40:13–18; pedicel, 30–33:20– 23; F1, 25–28:8; F2, 3:6–8; F3, 23–33:8; F4, Distribution. India: Andhra Pradesh, Himachal 23–28:6–8; F5, 20–28:8; F6, 20–25:10–13; F7, Pradesh, Karnataka, Uttar Pradesh Uttarakhand. 20–25:13–15; clava, 73–85:23; mesosoma, 125– 150; mesoscutum, 40–43; scutellum, 13; frenum, Comments. All the recorded specimens (7 females 50–58; metanotum, 6–8; propodeum, 30–33; fore and 2 males in ZDAMU, determined as C. dravida wing length:width, 323–385:23–28; longest mar- Subba Rao by Rehmat & Anis), figs 7-11 provided ginal seta, 138–140; hind wing length:width, 310– by Manickavasagam & Rameshkumar (2011), and 378:13–18; longest marginal seta, 110–125; proti- male paratype of C. dravida are found to be misiden- bia, 80–88; mesotibia, 110–113; mesobasitarsus, 23; tified. They are conspecific with C. enocki (Howard) metatibia, 113–120; petiole length:width, 18– in almost all the features, including patterns of stria- 23:30–33; gaster, 110–123; ovipositor, 80–90. tions on head and thoracic dorsum and structure of wings and distributions of microtrichia on fore wing Male. Length 270 μm. Head and mesosoma dark disc. brown. Antenna yellowish brown to brown. Gaster in basal three-fourth yellow, rest brown. Legs, including coxae, pale yellow. Head (Fig.Sample 135). Head, in frontal view, 1.2× as broad as high, with substrigulate striations. Antenna (Fig. 134) with scape 2.3× as long as broad; pedicel 1.4× as long as broad, slightly longer than F1, F3 and F4 individually, subequal to F6–F9 individually; flagellum 10-segmented, mps on all flagellar seg-

eschweizerbart_xxx 136. Jürgen Sieg: Neuere Erkenntnisse zum natürlichen Sys- ter besonderer Berücksichtigung des Dispersionsvermö- tem der Tanaidacea. Eine phylogenetische Studie. gens und des Reproduktionserfolgs der Arten. 1984. IV , 132 Seiten. ISBN 978-3-510-55022-7, € 100.– 2001. 147 Seiten. ISBN 978-3-510-55038-8, € 71.– 137. Hilke Ruhberg: Die Peripatopsidae (Onychophora). 152. Sigrun Bopp: Parasitismus oder Symbiose? Beziehungen Systematik, Ökologie, Chorologie und Phylogenetische zwischen einem parasitischen Bestäuber (Hadena bicru- Aspekte. ris Hufn., Lepidoptera: Noctuidae) und seinen Wirtspflan- 1985. VI , 184 Seiten. ISBN 978-3-510-55023-4, € 152.– zen (Silene-Arten, Caryophyllaceae). 138. Heinz-Hermann Reichenbach-Klinke: Zur Kenntnis der in 2003. X, 140 Seiten. ISBN 978-3-510-55039-5, € 98.– Mitteleuropa vorkommenden Polyopisthocothylea Odh- 153. Günther Fleck; André Nel: Revision of the Mesozoic fam- ner 1912 (Monogenea, Plathelminthes). ily Aeschnidiidae (Odonata: Anisoptera). 1986. VIII , 73 Seiten. ISBN 978-3-510-55024-1, € 66.– 2003. 172 pages. ISBN 978-3-510-55040-1, € 88.– 139. Harald Plachter; Johanna Plachter: Ökologische Studien 154. Joachim Haupt: The Mesothelae - a monograph of an ex- zur terrestrischen Höhlenfauna Süddeutschlands. ceptional group of spiders (Araneae: Mesothelae) (Mor- 1988. 67 Seiten. ISBN 978-3-510-55025-8, € 63.– phology, behaviour, ecology, taxonomy, distribution and 140. Johann-Wolfgang Wägele: Evolution und phylogene- phylogeny). tisches System der Isopoda. Stand der Forschung und 2003. 102 pages. ISBN 978-3-510-55041-8, € 78.– neue Erkenntnisse. 155. Bernhard A. Huber: High species diversity, male-female 1989. 262 Seiten. ISBN 978-3-510-55026-5, € 147.– coevolution, and metaphyly in Southeast Asian pholcid 141. Ernst Ebermann: Das Phänomen Polymorphismus in der spiders: the case of Belisana Thorell 1898 (Araneae, Milbenfamilie Scutacaridae (Acari, Heterostigmata, Tar- Pholcidae). sonemina, Scutacaridae). Friedrich Schaller: Tiere tropischer Regenwälder: struk- 1991. 76 Seiten. ISBN 978-3-510-55027-2, € 76.– turelle und funktionelle, vorwiegend qualitative Aspekte - Versuch einer Synopsis. 142. Barbara Kiesswetter: Stereotypes Verhalten. Begriff, Er- 2005. II, 167 pages. ISBN 978-3-510-55042-5, € 118.– scheinungsformen, auslösende Faktoren und Genese. Untersuchungen an Großkatzen in Zoologischen Gärten. 156. Hans Klaus Pfau: Functional Morphology and Evolution 1992. VI , 96 Seiten. ISBN 978-3-510-55028-9, € 96.– of the Male Secondary Copulatory Apparatus of the An- isoptera (Insecta: Odonata) 143. Martin Mahner: Systema Cryptoceratorum Phylogeneti- 2011. 103 pages. ISBN 978-3-510-55043-2, € 118.– cum (Insecta, Heteroptera). 1993. IX , 302 Seiten. ISBN 978-3-510-55029-6, € 244.– 157. Christine V. Schmidt; Stanislav N. Gorb: Snake Scale Microstructure: Phylogenetic Significance and Functional 144. R.V. Southcott: Revision of the larvae of the Microtrom- Adaptations bidiinae (Acarina: Microtrombidiidae), with notes on life 2012. 106 pages. ISBN 978-3-510-55044-9, € 119.– histories. Ed.: Werner Funke; F. Schaller. 1994. IV , 156 pages. ISBN 978-3-510-55030-2, € 151.– 158. Gerd Alberti; Jacek Dabert: Fine Structure of the Feather Mite Falculifer rostratus (Buchholz, 1869) (Acari, Falcu- 145. Friedhelm Erhard: Vergleichend- und funktionell-anato- liferidae): Gnathosoma, Digestive System and Supracox- mische Untersuchungen am Pleon der Oniscidea (Crus- al Glands tacea, Isopoda). Zugleich ein Beitrag zur phylogenetisch- pages 2012. 150 pages. ISBN 978-3-510-55045-6, € 154.– en Systematik der Landasseln. Hrsg.: Werner Funke; F. Schaller. 159. Esther Appel; Stanislav N. Gorb: Comparative functional 1995. IV , 114 Seiten. ISBN 978-3-510-55031-9, € 82.– morphology of vein joints in Odonata 2014. 104 pages. ISBN 978-3-510-55046-3, € 119.– 146. Porose integumental organs of oribatid mites (Acari, Oribatida) Ed.: Gerd Alberti; Roy A. Norton. 160. Gerd Alberti; Elliot W. Kitajima: Anatomy and Fine Struc- 1997. 143 pages. ISBN 978-3-510-55033-3, € 99.80 ture of Brevipalpus Mites (Tenuipalpidae) – Economically Important Plant-Virus Vectors 147. Peter Weygoldt: Revision of the Species of Phrynichus 2014. 192 pages. ISBN 978-3-510-55047-0, € 199.– Karsch, 1879 and Euphyrynichus Weygoldt, 1995 (Che- licerata, Amblypygi). 161. John D. Plant; Hannes F. Paulus: Evolution and Phylog- 1998. 65 pages. ISBN 978-3-510-55034-0, € 71.– eny of Bees – Review and Cladistic Analysis in Light of Morphological Evidence ( Hymenoptera, Apoidea) 148. Stanislav Gorb: Functional morphology of the head-ar- 2016. 364 pages. ISBN 978-3-510-55048-7, € 169.– rester system in Odonata. 1998. IV , 132 pages. ISBN 978-3-510-55035-7, € 74.– 162. František Gregor, Rudolf Rozkošný, Miroslav Barták & Jaromír 5N: Manual of Central European Muscidae 149. Stefan Bergleiter: Zur ökologischen Struktur einer zentra- (Diptera). Morphology, taxonomy, identification and dis- lamazonischen Fischzönose. Ethologische und morphol- tribution ogische Befunde zur Ressourcenteilung. 2016. 219 pages. ISBN 978-3-510-55049-4, € 169.– 1999. IV, 191 Seiten. ISBN 978-3-510-55036-4, € 102.– 163. Thomas Brockhaus: Die Eiszeitlibellen der Alten Welt. 150. Peter Weygoldt: Revision of the genus Damon C. L. Koch, Pleistozäne Biogeographie paläarktischer Libellen 1850 (Chelicerata: Amblypygi: Phrynichidae). Sperm 2018. 145 pages. ISBN 978-3-510-55050-0, € 109.– transfer, spermatophore morphology, and female genitalia of three species of whip spiders. Charinus seychella- 164. Thies H. Büscher, Constanze Grohmann, Sven Bradler & rum Kraepelin, 1898, Damon medius (Herbst, 1797), and Stanislav N. Gorb: Tarsal attachment pads in Phasmato- Phrynichius scaber (Gervais, 1844). (Chelicerata, Ambly- dea (Hexapoda: Insecta) Sample 2019. 94 pages. ISBN 978-3-510-55051-7, € 84.– pygi). 1999. 64 pages. ISBN 978-3-510-55037-1, € 50.– 151. Peter Kache: Die Besiedlung urbaner Lebensräume durch Kurzflügelkäfer (Coleoptera, Staphylinidae). Un-

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eschweizerbart_xxx Zoologica, Volume 165, 2020 P.T. Anwar, Fairyfly genus Camptoptera Foerster

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