BIOLOGICAL SCIENCES

The biology major provides a unified especially suitable for the distribution receives a formal review annually by the curriculum for undergraduates enrolled in requirement because they have no prerequi­ Cornell University Institutional Animal either the College of Agriculture and Life sites: BIO G 101-104, 105-106, 107-108, 109- Care and Use Committee (LACUC). Sciences or the College of Arts and Sciences. 110, 170, 202; BIOEE 154, 207, 264, 275; 4. Any concerns regarding the use of live Courses in biological sciences are integral to BIOGD 184; BIOMI 192; BIOAP 212; BIOPL animals in teaching should be many disciplines and are basic requirements 240, 241. Note that introductory biology can addressed first to the faculty member in many schools and colleges at Cornell. only count fo r distribution credit when taken responsible for that course. He or she as a two-semester sequence: BIO G 109-110, Graduate study in the biological sciences is is required to be in compliance with all 105-106, or 101 and 103 plus 102 and 104, administered by more than a dozen special­ applicable regulations and guidelines. or 107-108. Advanced placement in biology, ized fields within the Graduate School, as Alternatively, students may choose to with a score of 4 or 5 (6 or 8 credits, described in the Announcement of the address their concerns to the director respectively) may be applied to the Group 1 Graduate School. of the Cornell Center for Research distribution area in accordance with regula­ Animal Resources, Dr. Fred Quimby, at tions stipulated by the Arts College. 253-3520. The director may initiate In the College of Human Ecology, the natural discussion with the faculty member ORGANIZATION sciences distribution requirement is for at least responsible for a particular course 6 credits selected from BIO G 109-110, 101 without involving the student if he or Many different departments participate in the and 103 plus 102 and 104, 105-106 or 107-108 she would prefer to remain anonymous. biology major. or from specified courses in chemistry or 5. Enrollees in those courses in the physics. Advanced placement in biology with Student services are provided by the Office of biological sciences in which animal use a score of 4 or 5 (6 or 8 credits, respectively) Undergraduate Biology (OUB), is a component may, at the professor’s also satisfies the distribution requirement in www.bio.comell.edu, which includes the discretion, be asked to sign copies of the natural sciences. Behrman Biology Advising Center. Co-located this statement (USE OF ANIMALS...) at in Stimson Hall, the professional and student Switching from one introductory biology the first meeting of the course.” advisers provide academic and career sequence to another at midyear may not be advising, as well as help undergraduates find possible because of variation in presentation research opportunities on campus. Advisers in of topics. Students must receive permission of the OUB also follow the progress of biology the instructor to switch sequences. Taking ADVANCED PLACEMENT majors and work closely with faculty advisers. sequences in reverse order is strongly Additional services and resources of the discouraged. For information on credit for advanced Biology Center include tutoring, lecture tapes, placement in Biological Sciences, please see examination files, and extensive information the section on Advanced Placement in the on summer research opportunities and front of this publication. graduate programs. The center has comfort­ able areas for studying and relaxing. USE OF ANIMALS IN THE The Shoals Marine Laboratory, a cooperative BIOLOGICAL SCIENCES venture with the University of New Hamp­ CURRICULUM: CORNELL THE MAJOR shire, is located on Appledore Island in the UNIVERSITY The major of biological sciences is available to Gulf of Maine. Its base office in Stimson Hall students enrolled in either the College of provides academic and career advising for Students wishing to enroll in biology (“BIO”) Agriculture and Life Sciences or the College of students interested in the marine sciences and courses should know and understand the Arts and Sciences. The undergraduate administers the SEA Semester program for following criteria relative to the use of animals program is coordinated for students in both Cornell students pursuing studies at Woods in the teaching program, as passed by the colleges by the Office of Undergraduate Hole or aboard the schooner W estw ard or faculty of the Division of Biological Sciences Biology. By completion of the sophomore brigantine Corwith Cramer. in 1988, and reaffirmed in 1997: year, all students who intend to major in 1. “Live animals will be used for teaching biological sciences must declare the major and in certain courses in the biological a program of study through the Office of Undergraduate Biology, in 216 Stimson Hall. DISTRIBUTION REQUIREMENT sciences. Some animals will require humane euthanasia after they have Whenever possible, students should include In the College of Agriculture and Life been used for teaching. the introductory biology, chemistry, and Sciences, the biological sciences distribution 2. Courses bearing the “BIO” description mathematics sequences in their freshman requirement (Group B) is a minimum of 9 conform to the rules for the care of schedule and complete the organic chemistry credits, including at least 6 credits of such animals as outlined in Guiding lecture course in their sophomore year. introductory biology satisfied by Biological Principles in the Care and Use of Biology majors should regularly monitor their Sciences 109-110, 105-106, or 101 and 103 Animals (as approved by the Council of progress in the major, and should assess as plus 102 and 104, or 107-108. Advanced the American Physiological Society), realistically as possible the likelihood of placement in biology with a score of 4 or 5 the Guide for the Care and Use of achieving at a level that is consistent with their (6 or 8 credits, respectively) satisfies the Laboratory Animals. (DHEW publica­ academic and personal goals. Weak perfor­ requirement for introductory biology. The tion 86-23, revised 1996; see p. 14, mance in core courses, particularly after the additional credits may be satisfied by any Courses o f Study), the Animal Welfare freshman, year, may indicate a need to re­ biological sciences courses ex cep t BIO G 2 0 0 Act, and the New York State Public evaluate aptitude and genuine interest in the (unless permission is obtained), BIO G 2 09, o r Health Law. Within these regulations, major. Students with questions, particularly BIOSM 367. and in keeping with the principle of with concerns about their ability to complete For students in the College of Arts and Academic Freedom of the Faculty, the the major, are encouraged to consult with Sciences, all biology (“BIO”) courses can be use of animals to aid in teaching any their biology adviser, and to take advantage of used toward fulfillment of the biological biological sciences discipline is at the the advising and counseling resources of the distribution requirement except BIO G 2 0 0 discretion of the professor in charge. Office of Undergraduate Biology as well as (unless permission is obtained), BIO G 2 0 9 , o r those of the university and their college. 3- Each course, as well as research BIOSM 3 6 7 . The following courses are projects, in which animals are used The requirements for the biological sciences major are listed below. Requirements 1-8 148 BIOLOGICAL SCIENCES - 2EE 1-2 EE

must be taken for a letter grade. Courses high school or (b) achieving “qualifica­ ment; BIONB 322 Hormones and taken for the program of study should be tion” status in a language as-defined by Behavior; BIONB 325 taken for a letter grade unless the course is the College of Arts and Sciences or (c) Neurodiseases-Molecular Aspects’; offered for S-U only or if the student’s adviser successfully completing at least six BIONB 326 The Visual System; grants permission. college credits in a foreign language. BIONB 492 Sensory Function; NS Students registered in the College of 331 Physiological and Biochemical 1) Introductory biology for majors Arts and Sciences must satisfy the Bases of Human Nutrition. (one year): BIO G 101 and 103 plus language requirement as stated by that b) Laboratory courses: AN SC 301 102 and 104, or 105-106. BIO G 107- college. 108, offered during the eight-week Animal Reproduction and Develop­ Cornell Summer Session for eight •Since modem biology has an important ment; BIO G 401 Introduction to credits, also satisfies the introductory physical and quantitative orientation, students Scanning Microscopy; BIO G 403 biology requirement for majors. are advised to undertake basic science courses Transmission Electron Microscopy Students may choose to accept that emphasize this approach. Asterisks in the for Biologists; BIOAP 313 Histology: advanced placement credit if they have above list indicate the courses that provide The Biology of the Tissues; BIOAP received a score of 5 on the Advanced this orientation, but all courses listed are 319 Animal Physiology Laboratory; Placement Examination of the College acceptable. BIOBM 440 Experimental Molecular Biology; BIOBM 44lExperimental Entrance Examination Board (CEEB). Although not required for the biological Students with a score of 4 must fulfill Proteins and Enzymology; BIOBM sciences major, a course in statistics is 442 Experimental Cell Biology; the introductory biology requirement recommended for students planning graduate by taking BIO G 101-102, 101 and 103, BIOBM 443 Experimental Molecular study or a research career. Students should Neurobiology; BIONB 491 102 and 104, 103-104, or 105 or 106. consult their faculty advisers when choosing Principles of Neurophysiology. These students should consult appropriate courses in statistics. information available in the respective 2) Biochemistry: Chemistry 300, Quantita­ course offices and in the Office of Note: Core courses cannot count toward the tive Chemistry; six credits of organic Undergraduate Biology (216 Stimson Program of Study Requirements. chemistry (Chemistry 357-358 or 359- Hall) to determine which semester to 360); a minimum of four credits of complete the introductory biology Programs of Study and Requirements organic chemistry laboratory (Chemis­ requirement. For students in doubt, As noted in the list of requirements above, try 301-302 or 251-252-302 or 301 or completion of BIO G 101 and 103 is students accepted into the biological sciences 251-252); four credits of biochemistry advised. These students receive a total major must choose a Program of Study. laboratory courses (BIOBM 430); and of eight introductory biology credits Whereas the core requirements of the biology Physical Chemistry (Chemistry 389-390 (four AP credits plus four course curriculum provide the common foundation or 287-288 or 287-390 or 389-288). credits). deemed essential for all biology majors, the N ote: Chemistry 288 is designed for biologists. 2) General chem istry (one year); role of the Program of Study is to provide Five hours of Biochemistry are recommended either a concentration in a particular area of Chemistry 207-208,* or 206-208, or (331 and 332 or 330 and 334 or 333 and 334) biology or, in the case of the General Biology 215- 216.* and students interested in graduate work in Program of Study, a survey of biology that is biochemistry should take Physics 207-208 and 3) College mathematics (one year): one broad but not superficial. The Program of semester of calculus (Mathematics 106, consider taking a third semester of calculus in Study requirement can be met by taking 13 to preparation for CHEM 389-390. Be sure to 111, 191 or their equivalent) plus one 15 credit hours of courses chosen by the semester selected from the following: complete CHEM 207-208 or 215-216 during student in consultation with his or her biology the freshman year. a. a second semester of calculus adviser. Programs of Study for particular (Mathematics 112, 192, or their subject areas are designed by faculty N ote: Biology majors in the College of equivalents). specializing in the subject. Typically, the Agriculture and Life Sciences who select this Program of Study consists of one or more program of study are allowed to take up to 61 b. a course in finite mathematics courses that provide foundation in the subject credit hours in the endowed colleges due to (Biometry 101, 417, Mathematics and a list of optional courses from that area or the high number of required endowed courses 105, 231). related areas, many of which are at an for Biochemistry. c. a course in statistics (Biometry 261, advanced level (300 or higher). Because 3) Computational Biology: One course in Mathematics 171, Agriculture and biology is an experimental science, most computer programming (COM S 100, Resource Management 210, Programs of Study require one or more Introduction to Computer Programming Psychology 350, Industrial and laboratory courses. The laboratory require­ or ABEN 151, Introduction to Comput­ Labor Relations 210 and 211). ment in some Programs of Study can be met ing); one course in mathematics by participation in the independent research 4) Organic chemistry: Chemistry 257 (MATH 221, Linear Algebra and course (BIO G 499). The possible Programs of and 251, or 357-358 and 251, or 357- Calculus; MATH 231, Linear Algebra; Study and their requirements are listed below: 358 and 301, or 359-360 and 251, or MATH 294, Engineering Mathematics; 359-360 and 301. 1) Animal Physiology: BIOAP 311 Introduc­ MATH 420, Differential Equations and tory Animal Physiology, BIOAP 316 Dynamical Systems; BTRY 408, Theory 5) Physics: Physics 207-208,* 112-213,* Cellular Physiology, plus a minimum of 7 of Probability; or BTRY 421, Matrix or 101-102. Those who take Physics credit hours selected from the following Computation); a bridging course, i.e., a 112-213 are advised to complete lecture and laboratory courses, of which course in mathematical modeling Physics 214 as well. at least 4 credit hours must be a labora­ applied to biology (BIOEE 460, 6) Genetics: BIOGD 281. tory course. Theoretical Ecology; or BIOBM 321, Numerical Methods in Computational 7) Biochemistry: BIOBM 330, or 331 a) Lecture courses: ABEN 454 Molecular Biology); and one course Physiological Engineering; AN SC and 332, or 333. from the following list of advanced 300 Animal Reproduction and 8) Evolutionary Biology: BIOEE 278 or courses: Development; AN SC 410 Nutri­ BIOPL 448. Note: BIOPL 241, Botany, is tional Physiology and Metabolism; BIOBM 631, Protein Structure and a prerequisite course to BIOPL 448. AN SC 427 Fundamentals of Function; BIOGD 481, Population 9) A program of study selected from the Endocrinology; BIO G 305 Basic Genetics; BIOGD 484, Molecular outline below. Immunology; BIOAP 214 Biological Evolution; BIONB 422, Modeling Behavioral Evolution; BIOPL 440, 10) Foreign language: students registered Basis of Sex Differences; BIOAP 458 Phylogenetic Systematics; AN SC 420, in the College of Agriculture and Life Mammalian Physiology; BIOBM 407 Quantitative Animal Genetics; NTRES Sciences must satisfy the foreign Nature of Sensing and Response: 305, Wildlife Ecology; NTRES 340, language requirement for the biology Signal Transduction in Biological Quantitative Population Analysis; major by (a) presenting evidence of Systems; BIOBM 437 Eukaryotic NTRES 410, Quantitative Methods in successful completion of three or more Cell Proliferation; BIOGD 385 Wildlife Management; NTRES 670, years of study of a foreign language in Developmental Biology; BIOGD 483 Molecular Aspects of Develop­ THE MAJOR 149

Spatial Statistics; BTRY 451, Mathemati­ N ote: The Ecology and Evolutionary Biology Students may also choose from the cal Modeling of Populations; BTRY 662, program of study offers an undergraduate following courses to complete the 13- Mathematical Ecology; BTRY 408, specialization in Marine Biology and Oceanog­ credit requirement: BIOGD 682, Theory of Probability; MATH 420, raphy. A description of this specialization can Fertilization and Early Development; Differential Equations and Dynamical be found in the section entitled COURSES IN BIOGD 684, Advanced Topics in Systems; COM S 211, Computers and MARINE SCIENCE. Population Genetics; BIOGD 687, Programming; COM S 409, Data Note-. The Organization for Tropical Studies Developmental Genetics; BIOBM 633, Structures and Algorithms for Computa­ Biosynthesis of Macromolecules; tional Science; OR&IE 360, Engineering (OTS) offers an Undergraduate Semester Abroad Program, featuring two courses in BIOBM 639, The Nucleus; BIOEE 663, Probability and Statistics II; OR&IE 361, biology (Fundamentals of Tropical Biology Theoretical Population Genetics; BIOMI Introductory Engineering Stochastic 694, Genetics of Diverse Bacteria; Processes. and Field Research in Tropical Biology) and one course each in Environmental Policy and BIOPL 641, Laboratory in Plant N ote: MATH 112, Calculus should be used to Latin American Culture. Cornell biology majors Molecular Biology; BIOPL 644, Plant fulfill the core requirement for a second term in the Ecology and Evolutionary Biology Growth and Development; BIOPL 652, of math. Program of Study who complete the OTS Plant Molecular Biology II; BIOPL 653, Program as part of Cornell Abroad can Plant Molecular Biology I; PL BR 606, N ote: Bridging courses require linear algebra Advanced Plant Genetics. (MATH 221, Linear Algebra and Calculus; or substitute the credit earned from the biology MATH 231, Linear Algebra; or MATH 294, courses for two three-credit courses at the 400 Up to three credits for this program of Engineering Mathematics; or BTRY 421, Matrix level from list b. The OTS Program is study may be chosen from other Computation). For bridging course BIOEE 460 administered through Duke University. biological sciences courses, including Theoretical Ecology, MATH 420 Differential Applications are available at Cornell Abroad, BIO G 499, Undergraduate Research in Equations and Dynamical Systems will also 474 Uris Hall. Biology, with approval of the faculty serve as a prerequisite. 5) General Biology: The Program of Study adviser. N ote: BTRY 408 and MATH 420 can satisfy in General Biology requires a minimum 7) M icrobiology. At least 13 credits either the math requirement or a requirement of 13 credit hours in addition to including: BIOMI 290, General for an additional course. courses counted toward requirements Microbiology, Lectures; BIOMI 291, 1-10 on page 144. These 13 credits General Microbiology, Laboratory; N ote: Students who use BTRY 408 to fulfill the must include: either BIOMI 414, Bacterial Diversity or additional mathematics requirement should BIOMI 416, Bacterial Physiology, and at not use OR&IE 360, Engineering Probability a) One course from each of three different programs of study in least 2 courses from the following list: and Statistics II to fulfill the requirement for an BIOMI 391, Advanced Microbiology additional course. biology. Only those courses specifically listed as fulfilling a Laboratory; BIOMI 418, Microbial 4) Ecology and Evolutionary Biology: program of study requirement are Ecology; BIOMI 420, Microbial BIOEE 26l, Ecology and the Environ­ acceptable without permission of Genomics; or BIOMI 485, Bacterial ment, and 10 credits from the following adviser. Genetics. course lists, including at least one b) A course with a laboratory. 8) M olecu la r a n d C ell B iolog y : Chemistry course from e4ch group: 357-358 or 359-360; BIOBM 432, (a) BIOPL 241, Introductory Botany; c) And, a minimum of two upper-level Survey of Cell Biology; four credits of BIOEE 274, The Vertebrates: (300 and above) courses of two or BIOBM 430, Laboratories in Biochemis­ Structure, Function, and Evolution; more credits each. try, Molecular and Cell Biology, and at BIOEE 373, Biology of the Marine 100-level courses are not acceptable for least seven additional credits of courses Invertebrates; BIOMI 415, Bacterial meeting any of these requirements. that have a cell biological or molecular Diversity; BIOEE 471, Mammalogy; BIOPL 341 may not count as the lab course; biological orientation. The seven BIOEE 472, Herpetology; BIOEE BIO G 498 may not be used to fulfill the additional hours should include at least 475, Ornithology; BIOEE 476, requirements of this program of study. BIO G two courses from the following list: Biology of Fishes; ENTOM 212, 499 (minimum of two credits, but no more BIOAP 619, Lipids; BIOAP 658, Insect Biology. than three credits) may count as one of the Molecular Mechanisms of Hormone Action; BIOBM 407, Nature of Sensing (b) BIOEE 263, Field Ecology; BIOPL upper-level courses, and may count as the laboratory course with approval of the and Response: Signal Transduction in 447, Molecular Systematics; BIOPL Biological Systems; BIOBM 434, 448, Plant Evolution and the Fossil adviser, but it cannot count as a course representing a program of study. Applications of Molecular Biology; Record; BIOEE 452, Herbivores and BIOBM 437, Eukaryotic Cell Prolifera­ Plants: Chemical Ecology and N ote: It is possible to use a single course to tion; BIOBM 439, Molecular Basis of Coevolution; BIOEE 455, Insect fulfill more than one requirement. For Human Disease; BIOBM 631, Protein Ecology; BIOEE 456, Stream example, BIOAP 313, Histology, could count Structure and Function; BIOBM 632, Ecology; BIOEE 457 and 459, in all three areas: as a course in the Animal Membranes and Bioenergetics; BIOBM Limnology: Ecology of Lakes, Physiology program of study, as an upper 633, Biosynthesis of Macromolecules; Lectures and Laboratory; BIOEE level course, and as a course with a lab. BIOBM 635, Mechanisms of Metabolic 460, Theoretical Ecology; BIOEE Regulation and Mammalian Gene 462, Marine Ecology; BIOEE 463 6) Genetics and Development: A minimum of 13 credits, usually chosen from the Expression; BIOBM 636, Advanced Cell and 465, Plant Ecology and Biology; BIOBM 639, The Nucleus; BIO Population Biology, Lectures and following courses: BIOGD 385, Developmental Biology; BIOGD 387, G 305, Immunology; BIOGD 385, Laboratory; BIOEE 464, Macroevo­ Developmental Biology; BIOGD 483, lution; BIOEE 466 and 468, Developmental Aspects of Evolution; BIOGD 389, Embryology; BIOGD 450, Molecular Aspects of Development; Physiological Plant Ecology, BIOGD 484, Molecular Evolution; Lectures and Laboratory; BIOEE Vertebrate Development; BIOGD 480, Seminar in Developmental Biology; BIOGD 486, Advanced Eukaryotic 473, Ecology of Agricultural Genetics; BIOGD 682, Fertilization and Systems; BIOEE 478, Ecosystem BIOGD 481, Population Genetics; BIOGD 482, Human Genetics and the Early Embryo; BIOMI 290, General Biology; BIOEE 479, Paleobiology; Society; BIOGD 483, Advanced Microbiology, Lectures; BIOMI 408, BIOGD 481, Population Genetics; Viruses and Disease I; BIOMI 409, BIOGD 484, Molecular Evolution. Develpmental Biology; BIOGD 484, Molecular Evolution; BIOGD 486, Viruses and Disease II; BIOMI 485, N ote: One 400-level, four-credit course Advanced Eukaryotic Genetics; BIOMI Bacterial Genetics; BIONB 222, (including four credits from BIOSM 364) 485, Bacterial Genetics; BIONB 493, Neurobiology and Behavior II: offered at Shoals Marine Laboratory may be Developmental Neurobiology; BIOPL Introduction to Neurobiology; BIONB applied toward the 10 credits. Students are 343, Molecular Biology and Genetic 325, Neurodiseases-Molecular Aspects; encouraged to gain experience in some aspect Engineering of Plants; BIOGD 600, BIONB 425, Molecular Neurophysiol­ of field biology through course work at a Development of Sensory Systems. ogy; BIONB 495, Molecular and biological field station or work experience. Genetic Approaches to Neurosciences; BIOPL 343, Molecular Biology* and BIOLOGICAL SCIENCES - 2001-2002

Genetic Engineering of Plants; BIOPL Plant Function and Growth, Lectures (b) BIOEE 464, Macroevolution; BIOEE 347, Laboratory in Molecular Biology and Laboratory; BIOPL 247, 479, Paleobiology; ‘BIOPL 440, and Genetic Engineering of Plants; Ethnobiology; BIOPL 248, Taxonomy Phylogenetic Systematics; BIOPL BIOPL 444, Plant Cell Biology; BIOPL of Vascular Plants; BIOPL 342 and 344, 447, Molecular Systematics; ‘BIOPL 641, Laboratory in Plant Molecular Plant Physiology, Lectures and 448, Plant Evolution and the Fossil Biology; BIOPL 652, Plant Molecular Laboratory; BIOPL 343 and 347, Record; ‘BIOPL 453, Historical Biology II. Five hours of biochemistry Molecular Biology and Genetic Biogeography; BIOPL 442, Current are recommended (BIOBM 331 and Engineering of Plants, Lectures and Topics in Ethnobiology. 332, or 330 and 334). CHEM 207-208 or Laboratory; BIOPL 345, Plant Anatomy; 13) Independent Option: Students who 215-216 should be completed during BIOPL 444, Plant Cell Biology; BIOPL want to undertake a course of study the freshman year. 445, Photosynthesis; BIOPL 447, not covered by the existing programs Molecular Systematics; BIOPL 448, of study may petition the Biological 9) Neurobiology and Behavior: The two- Plant Evolution and the Fossil Record; semester introductory course sequence, Sciences Curriculum Committee. BIOEE 463 and 465, Plant Ecology and Neurobiology and Behavior I and II Information on independent option Population Biology, Lectures and and Curriculum Committee petition (BIONB 221 and 222) with discussion Laboratory; or BIOEE 466 and 468, section (four-credits per term), and forms are available in the Office of Physiological Plant Ecology, Lectures Undergraduate Biology, 216 Stimson seven additional credits. These and Laboratory. additional credits must include a course Hall. from the neurobiology and behavior Option (b) P la n t B io tech n olo g y : offerings (this course can NOT be Students are required to take BIOPL Independent Research and Honors BIONB 420, 720, or BIOG 499). 343 and 347, Molecular Biology and Program However, BIONB 420, 720 and BIO G Genetic Engineering of Plants, Lectures Biology majors are encouraged to consider 499 MAY be used to supplement this and Laboratory. Students choose, in participating in individual research under the neurobiology and behavior course to consultation with their faculty adviser, direction of a Cornell faculty member. fulfill the seven additional credits. a minimum of 10 additional credits Students interested in participating in research Students must consult their adviser for from the following list: BIOPL 241, should contact faculty members with courses that may be applied toward the Introductory Botany; BIOPL 242 and compatible research interests. Faculty seven additional credits that are not 244, Plant Function and Growth, members may consider the students’ previous listed in the Biological Sciences course Lectures and Laboratory; BIOPL 342 academic accomplishments, interests and offerings. BIO G 498 may not be used and 344, Plant Physiology, Lectures and goals, and the availability of space and to fulfill the requirements of this Laboratory; BIOPL 444, Plant Cell equipment when agreeing to supervise a program of study. Biology; BIOPL 648, Plant Biochemis­ student in his or her laboratory. Students try; PL BR 401, Plant Cell and Tissue N ote: Students who declare the Program of Culture; or PL BR 402, Plant Tissue conducting research may enroll for credit in Study in Neurobiology and Behavior after Culture Laboratory. Biological Sciences BIO G 499, Undergraduate taking BIONB 221 or 222 for only three credits Research in Biology. Registration for this must still take the 1 credit discussion section 12) Systematics and Biotic Diversity: A course is done in 216 Stimson Hall. The in BIONB 221 and 222. To arrange this, the minimum of 13 credits from the student’s research project must have signifi­ student should consult the professors in following two groups, including at least cant biological content in order to be charge of the two courses. In the event that a seven credits from group (a), and three considered for BIO G 499 credit. Students may student cannot take the discussion sections, from group (b), and at least two not earn credit for research conducted outside he or she should contact the department laboratory courses (marked with *)• of Cornell. Up to three credits of research may Director of Undergraduate Studies to BIO G 499, Undergraduate Research in be used to complete the program of studies in determine what course(s) to use to address Biology, with approval of the adviser, general biology, genetics and development, the deficiency. can be used in fulfillment of Up to four systematics and biotic diversity, as well as four credits in group (a), and can count as credits of research in neurobiology and 10) N utrition: NS 331, Physiological and one laboratory course if it has a Biochemical Bases, of Human Nutrition (4 behavior. Information about faculty research laboratory component of two or more interests and undergraduate research credits) and at least nine credits of credits. opportunities is available in the Office of additional coursework in the biological Undergraduate Biology, 216 Stimson Hall. aspects of nutrition, such as NS 31'5, (a) ‘ BIOEE 274, The Vertebrates: Obesity and the Regulation of Body Structure, Function, and Evolution; The Honors program in biological sciences is Weight; NS 332, Methods in Nutritional ‘BIOEE 371, Human Paleontology; designed to offer advanced training in Sciences; NS 431, Mineral Nutrition and •BIOEE 373, The Invertebrates: laboratory or field research through the Chronic Disease; NS 441, Nutrition and Form, Function, and Evolution; performance of an original research project Disease; NS 475, Molecular Nutrition and ‘BIOEE 471, Mammalogy; ‘BIOEE under the direct guidance of a member of the Development; NS 602, Lipids; NS 604, The 472, Herpetology; ‘BIOEE, 475, Cornell faculty. Applications for the honors Vitamins; and NS 614, Topics in Maternal Ornithology; ‘BIOEE 476, Biology program are available in the Office of and Child Nutrition. Some courses require of Fishes; BIOMI 290, General Undergraduate Biology, 216 Stimson Hall, and NS 115 Nutrition and Health: Concepts Microbiology, Lectures; ‘BIOMI 291, must be submitted in October of the senior and Controversies, which may be used as General Microbiology, Laboratory; year to the Honors Program Committee by the part of the additional nine credits. BIOMI 415, Bacterial Diversity, announced deadline. To qualify for the Lectures; ‘BIOPL 241, Introductory program, students must have been accepted N ote: For students in the College of Agricul­ Botany; ‘BIOPL 243, Taxonomy of into the biological sciences major, have ture and Life Sciences, credits in NS courses Cultivated Plants; BIOPL 247, completed at least 30 credits at Cornell, and count towards the required 55 CALS credits. Ethnobiology; ‘BIOPL 248, For students in the College of Arts and have a cumulative grade-point average of at Taxonomy of Vascular Plants; least 3.0. In addition, students must have at Sciences, NS credits will count toward the 100 BIOPL 343, The Healing Forest; least a 3 0 cumulative grade-point average in hours required in A&S if those credits fulfill BIOPL 645, Families of Tropical major requirements. all biology, chemistry, mathematics, and Flowering Plants, Lectures; ‘BIOPL physics courses. (Grades earned in courses in 11) Plant Biology: Students choose one 646, Families of Tropical Flowering other departments that are used to fulfill area of study from the following two Plants; ‘ENTOM 212, Insect Biology; biology major requirements are included in options: ENTOM 215, Spider Biology: Life on this computation.) In addition, candidates a Silken Thread; ‘ENTOM 322, Option (a) B otan y : Students are must find a Cornell faculty member in required to take Introductory Botany Insect Morphology; ‘ENTOM 331, Biological Sciences to supervise their research. Introductory Insect Systematics; (BIOPL 241). Students should then An honors candidate usually enrolls for credit ‘ENTOM 471, Freshwater Inverte­ choose, with the aid of their faculty in BIO G 499, Undergraduate Research in adviser, a minimum of three of the brate Biology; ‘ENTOM 631, Biology, under the direction of the faculty following courses, for a total of at least Systematics of the Coleoptera; PL member acting as honors supervisor, although PA 309, Introductory Mycology; *PL 10 additional credits, to round out their the Honors Program does not require botanical training: BIOPL 242 and 244, PA 319, Field Mycology. enrollment for credit. Students accepted into INDEX OF COURSES 151

the Honors program are required to partici­ for students in the College of Agriculture and C ou rse P ag e C ou rse P ag e pate in honors research seminars during both Life Sciences and as arts and sciences credits 452 782 semesters of their senior year; submit an for students in the College of Arts and 159 164 acceptable honors thesis; complete all major Sciences. 455 159 783 164 requirements; and maintain a 3.00 Cornell 456 160 786 164 cumulative and science grade-point average Course Index through graduation. Recommendation to the 457 160 787 164 C ou rse P ag e C ou rse faculty that a candidate graduate with honors P ag e 459 160 BIOMI and at what level of honors is the responsibil­ BIO G 321 156 460 160 192 164 ity of the Honors Program Committee. The 101 152 330 156 student’s final grade point average is a factor 462 160 290 164 102 152 331 156 in determining the level of honors recom­ 463 160 291 164 mended. 103 152 332 156 464 160 292 164 Students interested in the honors program are 104 152 333 156 465 160 331 164 strongly encouraged to begin their research 105 152 334 156 projects in their junior year and to consider 466 160 391 164 106 152 407 156 spending the following summer here at 467 160 394 164 Cornell engaged in full time research on their 107 152 432 156 468 160 397 165 honors project. Students interested in staying 108 152 434 156 for the summer also are encouraged to apply 469 160 404 165 109 153 435 157 to the Cornell Hughes Scholars Program. 471 160 408 ■165 110 153 436 157 Biology majors who are considering study 472 161 409 165 abroad and graduating with honors are 170 153 437 157 473 161 414 165 encouraged to meet with their academic 200 153 438 157 adviser during their sophomore year to 474 161 416 ' 165 202 153 439 157 carefully plan their academic schedule to meet 475 161 417 165 the requirements of the Honors program. 209 153 440 157 476 161 418 165 Application forms, requirements, deadline 305 153 441 157 477 161 420 165 dates for the Honors Program and the Hughes 400 153 442 157 Scholars Program, and information pertaining 478 161 485 165 401 153 443 157 to faculty research may be obtained at the 479 161 610 165 403 450 Office of Undergraduate Biology, 216 Stimson 153 157 490 161 652 Hall. 165 408 153 631 157 660 161 690 165 410 153 632 . 157 661 161 791 166 431 153 633 157 665 162 795 166 CURRICULUM COMMITTEE 450 154 636 157 668 162 796 166 Many decisions pertaining to the curriculum 498 154 639 158 669 162 797 166 and to the programs of study are made by the 499 154 641 158 Biology Curriculum Committee. The commit­ 670 162 798 166 663 154 730 158 tee consists of faculty and elected student 673 162 799 166 members and welcomes advice and sugges­ 705 154 732 158 760 162 BIONB tions from all interested parties. 706 154 733 158 763 162 111 166 BIOAP 734 158 767 162 221 166 212 154 735 158 899 162 222 166 ADVISING 214 154 736 158 999 162 322 166 Students in need of academic advice are 154 311 737 158 BIOGD encouraged to consult their advisers or come 324 166 312 154 738 to the Office of Undergraduate Biology, 216 158 184 162 325 166 Stimson Hall. 313 154 751 158 281 162 326 166 316 154 761 158 Students interested in marine biology should 282 163 327 166 visit the Shoals Marine Laboratory Office, G14 319 155 830 158 Stimson Hall. 385 163 328 167 427 155 831 158 387 163 329 167 Students interested in the multidisciplinary 458 155 832 158 program of Biology and Society should see 389 163 392 167 475 155 833 158 “Special Programs and Interdisciplinary 394 163 394 167 Studies,” in the College of Arts and Sciences 619 155 836 158 450 396 section of this catalog. 163 167 710 155 837 158 480 163 420 167 711 155 BIOEE 481 163 421 167 712 155 154 158 482 163 422 167 INDEX OF COURSES 713 155 207 158 483 163 424 167 The following course identifiers are used to 714 155 261 158 484 denote biological sciences courses in specific 163 425 167 715 155 263 areas: General Courses, BIO G; Animal v 159 485 163 426 167 719 155 264 Physiology, BIOAP; Biochemistry, Molecular 159 486 163 427 167 and Cell Biology, BIOBM; Ecology and 757 155 267 159 Evolutionary Biology, BIOEE; Genetics and 600 163 428 167 811 155 274 Development, BIOGD; Microbiology, BIOMI; 159 682 163 429 167 812 155 275 Neurobiology and Behavior, BIONB; Plant 159 684 164 430 167 Biology, BIOPL; Shoals Marine Laboratory, BIOBM 278 159 BIOSM. 685 164 440 168 132 156 371 159 Note: Biological sciences (“BIO”) courses 687 164 441 168 232 156 373 count as agriculture and life sciences credits 159 780 164 470 168 233 156 405 159 781 164 491 168 152 BIOLOGICAL SCIENCES - 2001-2002

C ourse P ag e C ou rse P ag e 109-110 is a course sequence intended for BIO G 105-106 Introductory Biology nonmajors, and meets the prerequisite for 105, fall; 106, spring. 4 credits each term 492 168 642 171 many, but not all, upper-level courses listing (or 2 credits, with permission of instruc­ 493 168 644 171 “one year of introductory biology” as a tor). Enrollment limited to 200 students. 494 168 647 171 prerequisite. Students can earn a maximum of Prerequisite: 105 is prerequisite to 106, eight credits in introductory biology (including unless written permission is obtained from 168 648 171 495 advanced placement credits). instructor. May not be taken for credit after 496 - 168 649 172 BIO G 101-104 or 109-110. No admittance BIO C 101-102 Biological Sciences, after first week of classes. Lec, T 9:05 (first 531 168 651 172 Lectures lec of fall term, R 8/30 9:05); additional 623 168 652 172 101, fall; 102, spring. 2 credits each term. study and lab hours TBA. C. H. McFadden Prerequisite: concurrent enrollment in BIO 720 168 653 172 and staff. G 103 (fall) or 104 (spring). Passing grade Designed primarily for biology majors, 721 168 654 173 (D or better) in 101 is prerequisite to 102 preprofessionals, and other students who 723 169 656 173 unless permission is obtained from desire a challenging, broad introduction to instructor. May not be taken for credit after 724 169 740 173 fundamental concepts of biology. Cell biology, BIO G 105-106 or 109-110. S-U grades physiology, anatomy (accompanied by 725 169 741 173 optional, with permission of instructor. preserved vertebrate dissection), and 790 169 742 173 Lees, M W F 9:05 or 10:10. 2 lees each biochemistry are strongly emphasized in the week; to accommodate these, students 792 169 743 173 fall semester. Subjects in the spring semester must reserve all 3 days. Evening prelims: are genetics, development, ecology, evolution, 793 169 745 173 fall, Sept. 25 and Nov. 6; spring, Feb. 21 behavior, and the diversity of organisms 794 169 746 173 and Apr. 4. Staff. (accompanied by preserved and anesthetized Designed both for students who intend to BIOPL 749 173 invertebrate dissection). Students who plan to specialize in biological sciences and for those concentrate in anatomy and physiology 240 169 840 173 who want to obtain a thorough knowledge of should consider taking this course because of 241 169 BIOSM biology as part of their general education. The the strong emphasis on organismal biology. fall semester covers the chemical and cellular 242 169 160 174 Because some testing involves the use of basis of life, energy transformations, physiol­ predissected specimens, students who object 243 169 l6 l 174 ogy, neurobiology, and behavior. The spring to dissections should take BIO G 101-104. 244 170 204 174 semester covers genetics, development, The course uses an autotutorial format and evolution, and ecology. Each topic is offers considerable flexibility in scheduling. 245 170 303 174 considered in terms of modem evolutionary Completion of the course requires mastery of 247 170 308 174 theory, and discussions of plant and animal a group of core units. Testing on these units is 248 170 309 174 systems are integrated. For those students who primarily by oral examination. Students who object to animal dissection, alternative elect to take the course must be able to meet 340 170 329 174 materials are available for study. However, deadlines. Four formal laboratory sessions are 342 170 363 174 testing will involve identification of important offered each semester; additional laboratory structures in real organisms. 343 170 364 175 work is included in the core units. Evaluation is based on written reports on experimental 344 170 365 175 BIO G 103-104 Biological Sciences, Laboratory work, practical exams, and a comprehensive 170 366 175 345 103, fall; 104, spring. 2 credits each term. final- exam. 347 170 367 175 Prerequisite: concurrent enrollment in BIO BIO G 107-108 General Biology 348 170 368 175 G 101 (fall) or 102 (spring). 103 is Summer (8-week session; 107, weeks 1—4; prerequisite to 104 unless permission is 404 170 369 175 108, weeks 5-8). 4 credits each. Prerequi­ obtained from instructor. No admittance site: one year of college or permission of 440 170 370 175 after second week of classes. S-U grades instructor- BIO G 101, 103, 105, or 107 is a 441 170 372 175 optional, with permission of instructor. prerequisite for 108. Fee, $25 for weeks Lab, MTWorR 1:25-4:25, M or W 7:30- 442 ■ 170 374 175 1—4; $15 for weeks 5-8. Lees, M-R 9-12; 10:30 p.m., or T R or S 8-11. One 3-hour labs, M T R 1:30-4:30, F 9-12. Staff. 443 171 375 175 lab each week and a weekly lec for discs, Designed for students who plan further study 444 171 376 175 special lees, etc. J. C. Glase, P. R. Ecklund, in biology and for students who want a broad and staff. course in biology as part of their general 445 171 402 175 BIO G 103-104 is designed to provide education. BIO G 107 covers biological 447 171 413 175 laboratory experience with major biological metabolism, first at the molecular leyel and phenomena in order to support an under­ 448 171 418 .175 then progressively to the organ system level. standing of the important concepts, principles, The laboratory work involves an introduction 449 171 449 175 and theories of modern biology. A second to some major techniques, vertebrate 452 171 475 175 objective of the laboratory course is to help dissection, and a survey of plant organization. students gain expertise in the methods used 453 171 477 176 BIO G 108 seeks to integrate the topics of by biologists to construct new knowledge. genetics, developmental biology, population 454 171 490 176 Students are exposed to basic concepts, biology, and ecology in a general consider­ 456 171 495 176 research methods, including laboratory and ation of biological evolution. The laboratory data transformation techniques, and instru­ 641 171 499 176 work is a continuation of the material covered mentation in the major areas of biology. First- in BIO G 107 and involves more techniques, a semester topics include biochemistry, survey of animal organization, and the design GENERAL COURSES (BIO G) physiology, plant biology, and behavior. In and performance of a field study. BIO G 107- the second semester, laboratory experience is 108 fulfills the introductory biology require­ Three introductory biology course sequences provided in the areas of genetics, biotechnol­ are taught during the academic year: BIO G ment for majors and forms a suitable ogy, invertebrate diversity, plant and animal introductory biology course sequence for 101-104, BIO G 105-106, and BIO G 109-110; development, and ecology. During the first and one during the eight-week summer students intending to go to medical school. semester, dissection of a doubly-pithed frog is For those students who object to animal session: BIO G 107-108, BIO G 101-104, 105- included. Pithing is done by the instructor. 106, and 107-108 are intended for biological dissection, alternative materials are available Dissection of several invertebrates occurs for study. However, testing will involve sciences majors and other students needing during the second semester. For those eight credits from an introductory sequence identification of important structures in real students who object to animal dissection, organisms. for majors (for example, students in a alternative materials are available for study. premedical curriculum). Any of these However, testing will involve identification of sequences meet the prerequisite for upper- important structures in real organisms. level courses listing “one year of introductory biology for majors” as a prerequisite. BIO G GENERAL COURSES 153

BIO G 109-110 Biological Principles BIO G 200 Special Studies in Biology An introductory course that includes the 109, fall; 110, spring. 3 credits each term. Fall, spring, or summer. 1-3 credits. principle and use of the scanning electron Limited to 600 students. A passing grade in Prerequisites: written permission from the microscope. Students use biological material BIO 109, AY2000 or later, is prerequisite to Office of Undergraduate Biology. Students to explore and understand some of the fine BIO 110 unless w ritten permission is must register in 216 Stimson Hall. S-U biological architecture. In addition to obtained from the instructor. Since BIO G grades optional, with permission of preparing the specimens, students use the 109-110 together constitute an integrated instructor. Hours TBA. Staff. scanning electron microscope to study and survey, 109 cannot be used to satisfy the A registration device for students who want to obtain micrographs of features that interest College of Arts and Sciences or College of take only a portion of a regular biological them. Agriculture and Life Sciences distribution sciences course—for example, only the requirement unless it is followed by 110 or lectures or only the laboratory in a course that BIO G 403 Transmission Electron equivalent. Due to overlap in content, BIO includes both. Only students who have Microscopy for Biologists G 109 may not be taken after BIO G 102/ already had training equivalent to the portion Fall. 1, 3, or 4 credits (4 credits if student 104 or BIO G 106. BIO G 110 may not be of the regular course -that is to be omitted may takes both sections). Limited to 12 taken after BIO 101-102 or BIO G 105. register in this manner. This course may not students. Prerequisites: BIOAP 31'3, BIOPL This course sequence may be used to be substituted for 100-level courses and may 345 or 443. S-U grades optional. Two fulfill the distribution requirement in the not be used in fulfillment of college distribu­ sections: Sec 01, 1 credit, weeks 1-4; sec Colleges of Agriculture and Life Sciences, tion requirements except by permission from 02, 3 credits, weeks 5-12. Students may Arts and Sciences, and Human Ecology but the Office of Undergraduate Biology register for one or both sections. Fee may may n ot be used as an introductory course be charged. Lee, T 11:15; labs, MWorTR for the major in biological sciences or as [BIO G 202 The Diversity of Life 1:25-4:25. M. V. Parthasarathy. introductory biology for premedical Fall. 3 credits. S-U grades optional. Lees, Section 01, 1 credit, weeks 1 - i, covers the requirements. Note that this course satisfies M f F 2:30. Not offered 2001-2002. principles and use of the transmission electron the prerequisite fo r many, but not all J. I. Davis, J. J. Doyle, E. Rodriguez. microscopy (TEM), with emphasis on proper second- and third-level courses in biology. The main focus of this course is on the operation of the instrument and interpretation Letter grade only. Students do not choose diversity of living and extinct species. This of images obtained. Negatively stained lab sections during course enrollment; lab diversity is examined from an evolutionary materials are used for viewing with the assignments are made during the first day perspective, with attention to the principles transmission electron microscope. Section 02, of classes. Evening prelims: fall, Sept. 25 employed in the discovery of species and in 3 credits, weeks 5-12, covers the principles and Nov. 6; spring, Feb. 21 and Apr. 4. the analysis of relationships among them. and techniques of preparing biological Lees, M W F 9:05 or 10:10; lab meets Interactions between humans and other material for transmission electron microscopy. alternate MTWRorF 2-4:25 or T 10:10- species are examined during the latter portion Using animal, plant, and microbe materials 12:35. H. Greene, R. Turgeon, C. Eberhard of the semester.] this section studies chemical fixtures, and staff. BIO G 209 Introduction to Natural cryofixations, ultrathin sectioning, Students who do not plan to major in biology Science Illustration immunogold localization, quantitative microscopy, and metal shadowing techniques. may take this broad introductory course. The Summer (6-week session). 2 credits. Students have two additional Weeks to content is designed to appeal to anyone who Limited to 12 students. Prerequisite: free­ complete laboratory assignments at the end of seeks a comprehensive knowledge of biology hand drawing or permission of instructor. each section. as part of a general education. Broad goals of S-U grades optional. Lees and labs, T R the course encompass an understanding of the 6:30-9:30 p.m. B. S. King. BIO G 408 Presentation Skills for potential benefits and limitations of science, An introduction to the art of natural science Biologists the complexity and workings of the natural illustration for publication, and to the Spring. 1 credit. Prerequisites: previous world, and the internal machinery of life— techniques of various media including pencil, research experience. Preference given to how our bodies and those of other animals pen and ink, watercolor, colored pencil, students accepted into the Honors and plants work. Fall semester covers scratchboard, and carbon dust. Potentials and Program. L. Southard and G. Hess. biological diversity, genetics, evolution, limitations of line and half-tone reproduction, This course will cover oral and written ecology, behavior and conservation biology; copyright, and portfolio presentation are communication skills used in presenting spring semester covers human physiology, discussed. research to other scientists. Topics covered plant development, genetic engineering, will include organization of scientific papers, infectious diseases, and human health. BIO G 305 Basic Immunology Lectures presentation tips for research seminars, and Laboratory sections enable small groups of (also VETMI 315) preparation of visual aids using Power Point. Fall. 3 credits. Strongly recommended: students to meet with course staff and are All students will present a 10-minute seminar basic courses in microbiology, biochemis­ used for problem-solving experiments, on their research and will evaluate other try, and genetics. S-U grades optional, with demonstrations, and discussions. There are presentations. dissections of preserved vertebrate, inverte­ permission of instructor. Lees, T R 8:30- brate, and plant materials; for those students 9:55. J. A. Marsh. BIO G 410 Teaching Contemporary who object to dissection, alternative materials A survey of immunology, with emphasis on Biology are available for study and there is no grade the biological functions of the immune Fall. 3 credits. Prerequisite: one year penalty for omitting dissection or observation response. introductory biology; permission of of animals. Testing, for students choosing to BIO G 400 Undergraduate Seminar in instructor. L. Southard and S. Merkel. be tested, will involve identification of Biology This course provides students with the important structures in real organisms. opportunity to experience teaching high Fall or spring. Variable credit (1-3 credits school science. Students will concentrate on a BIO G 170 Evolution of the Earth and Life assigned for individual seminar offerings). topic of current public interest, then develop (also EAS 102) May be repeated for credit. S-U grades teaching plans appropriate for high school Spring. 3 credits. S-U grades optional. Lees, optional. Sem TBA. Staff. students. The first part of the course consists From time to time specialized seminars on T R 9:05 or 11:15; lab, T W or R 2:00-4:25; of lectures, discussion, and laboratory field trips during lab. J. L. Cisne. topics of interest to undergraduates are experiments, which will familiarize the Earth systems and their evolution. Earth offered by visiting faculty or faculty from the students with the scientific content of the history’s astronomical context. Plate tectonics, departments of Ecology and Systematies, course. Students will then work in teams with continental drift, and their implications for Genetics and Development, or Plant Biology. high school teachers to develop their climate and life. Coevolution of life and the For students interested in Biochemistry, presentations. The final part of the course will atmosphere. Precedents for ongoing global Physiology, or Neurobiology, please see include practice presentations and teaching at descriptions under the appropriate section. change. The course covers dinosaurs, mass, regional high schools. extinctions, and human ancestry, Laboratories BIO G 401 Introduction to Scanning include work on reconstructing geological Electron Microscopy BIO G 431 Frontiers in Biophysics history and mapping ancient geography. Fall. 1/2 credit. S-U grades only. Lee TBA. Fossil-collecting on field trips. Fall or spring, weeks 1-8. 1 credit. Limited to 8 students (fall), 12 students (spring). G. Feigenson and staff. S-U grades optional. Fee may be charged. A day of lectures on Saturday TBA giving an Lee, M 10:10; lab, T R or F 9:05-12:15 or T overview of current research in biophysics at W or R 1:25-4:25. M. V. Parthasarathy. Cornell by faculty from different departments 1 -2 0 02 154 BIOLOGICAL SCIENCES - 2

across the university. Designed for under­ BIO G 663 Nanobiotechnology (also [BIOAP 214 Biological Basis of Sex graduates who are considering a career in AEP 663) Differences (also B&SOC 214 and biophysics and for graduate students who are Spring. 3 credits. Prerequisite: permission WOMNS 214) interested in biophysics research opportunities of instructor. Letter grade. Lee, T R 1:25- Spring. 3 credits. Prerequisite: one year of at Cornell. 2:40. Nanobiotechnology faculty. introductory biology. S-U grades optional. Nanobiotechnology is the application of nano- Lees, T R 1:25-2:40. Offered alternate [BIO G 450 Light and Video Microscopy and microfabrication methods to build tools years. Not offered 2001-2002. J. E. Fortune. for Biologists for exploring the mysteries of biological The structural and functional differences Spring. 3 credits. Limited to 12 students. systems. It is a graduate-level course that will between the sexes are examined. Emphasis is Prerequisites: one year of introductory cover the basics of biology and the principles placed on mechanisms of mammalian biology and permission of instructor. Lees, and practice of microfabrication techniques reproduction; where possible, special attention T R 1:25-2:30; lab, R 2:30-4:30. Not offered with a focus on applications in biomedical is given to studies of humans. Current 2001-2002. R. O. Wayne. and biological research. One objective of the evidence on the effects of gender on course is to facilitate a means through which nonreproductive aspects of life (behavior, biologists and engineers can communicate. A mental, and physical capabilities) is discussed. Theoretical and practical aspects of light team design project that stresses interdiscipli­ The course is intended to provide students microscopy, including brightfield, darkfield, nary communication and problem solving will with a basic knowledge of reproductive phase-contrast, polarization, Hoffman- be one of the course requirements. endocrinology and with a basis for objective modulation contrast, interference, differential- evaluation of sex differences in relation to ■ BIO G 705 Advanced Immunology contemporary life.] interference contrast, and fluorescence Lectures (also VETMI 705) microscopy, as well as video- and computer- Spring. 4 credits. Prerequisite: BIO G 305 BIOAP 311 Introductory Animal based digital image enhancement, are studied. or permission of instructor. Offered Physiology, Lectures (also Students learn both qualitative and quantita­ alternate years. Next offered spring 2002. VETPH 346) tive techniques to probe noninvasively the Lees, M W F 9:05. Coordinator: Fall. 3 credits. Prerequisites: one year of structure and function of living cells.] E. J. Pearce. college biology, chemistry, and mathemat­ BIO G 498 Teaching Experience Coverage at an advanced level of molecular ics. Recommended: previous or concurrent Fall or spring. 1-4 credits. Enrollment and cellular immunology. course in physics. S-U grades optional, limited. Prerequisites: previous enrollment with permission of instructor. Evening BIO G 706 Immunology of Infectious prelims. Lees, M W F 11:15. E. R. Loew. in the course to be taught or equivalent. Diseases (also VETMI 719) Arts students may not count this course A general course in animal physiology Spring. 2 credits. Prerequisite: BIO G 305 toward graduation. They may, however, emphasizing principles of operation, or permission of instructor. S-U grades upon petition to their class dean, carry regulation, and integration common to a optional, with permission of instructor. few er than 12 other credits and remain in broad range of living systems from the cellular Lee, R 10:10-12:05. Offered alternate years. good standing. This would affect D ean’s to the organismal level. Structure/function Coordinator: E. Denkers. List eligibility, but not eligibility fo r relationships are stressed along with underly­ Coverage at an advanced level of the graduating with distinction. S-U grades ing physico-chemical mechanisms. immunology of diseases caused by selected optional, with permission of instructor. bacterial, viral, protozoan, and helminthic BIOAP 312 Farm Animal Behavior (also Hours TBA. Staff. parasites. ANSC 305) Designed to give qualified undergraduate Spring. 2 credits. Prerequisites: introduc­ students teaching experience through actual Related Courses in Other Departments tory animal physiology (AN SC 100 and involvement in planning and assisting in The Sea: An Introduction to Oceanography 150 or equivalent). Recommended: at least biology courses. This experience may include (Biological Sciences [BIOEE] 154) Medicine one animal production course or equiva­ supervised participation in a discussion group, and Civilization (Biology and Society 322) lent experience. S-U grades optional. Lees, assisting in a biology laboratory, assisting in T R 11:15. E. A. Oltenacu, K. A. Houpt. field biology, or tutoring. Biological sciences Pathogenic Bacteriology and Mycology (Biological Sciences [BIOMI] 404 and Veterinary The behavior of production species (avian and courses currently offering such experience Microbiology 318) mammalian) influences the success of any include BIO G 105-106; BIOAP 311, 313, 319; management program. Students study BIOBM 330, 331; BIOEE 274, 475; BIOGD Viruses and Disease (Biological Sciences [BIOMI] behaviors relating to communication, learning, 281; and BIOMI 291, 292. 408 and Veterinary Microbiology 408) social interactions, reproduction, and feeding BIO G 499 Undergraduate Research in of domestic animals, and their physiological Biology basis. Management systems for commercial livestock production and their implications for Fall or spring. Variable credit. Students in ANIMAL PHYSIOLOGY (BIOAP) the College o f Arts and Sciences may not animal behavior and welfare are stressed. register fo r more than 6 credits per term BIOAP 212 Human Physiology for Non- BIOAP 313 Histology: The Biology of the with one supervisor or 8 credits per term Biology Majors Tissues with more than one supervisor. Prerequi­ Spring. 3 credits. May not be taken for Spring. 4 credits. Prerequisite: one year of site: written permission of staff member credit after BIOAP 311. Limited to 130 introductory biology. Recommended: who supervises the work and assigns the students. This course may be used toward BIOBM 330 or 331, or their equivalents; grade. Students must register in the Office the science distribution requirement of the and previous or concurrent enrollment in of Undergraduate Biology in 216 Stimson College of Arts and Sciences and the BIOAP 311- S-U grades optional, with Hall. Each student must submit an Group B distribution requirement of the permission of instructor. Evening prelims. independent study statement describing College of Agriculture and Life Sciences. Lees, M W 12:20; labs, M W 1:25-4:25. the proposed research project during This course may not be used to fulfill the Staff. course registration. Applications are requirements of any program of study in Provides students with a basis for understand­ available in the college offices and in 216 the biological sciences major. Lees, M W F ing the microscopic, fine-structural, and Stimson Hall. Any faculty member in 1:25; disc, M W or F 2:15. M. D. Baustian. functional organization of vertebrates, as well Biological Sciences may act as a supervi­ Introduction to the principles of physiology as methods of analytic morphology at the cell sor. Supervisors outside of Cornell are not governing the function of the human body. and tissue levels. Dynamic interrelations of acceptable. S-U grades optional. Hours Emphasis will be placed on reproduction, structure, composition, and function in cells TBA. Staff. pregnancy and development, and immunol­ and tissues are emphasized. The course may Practice in planning, conducting, and ogy and the defense of the organism against include work with vertebrate animals. reporting independent laboratory and library disease. Major organ systems will be surveyed research programs. Up to three credits of to illustrate how physiologists study the BIOAP 316 Cellular Physiology research may be used to complete the function of living systems, and how this Spring. 4 credits. Limited to 72 students, Programs of Study in general biology, genetics knowledge has shaped the management of with preference given to students studying and development, and systematics and biotic health and disease. The contribution of in animal physiology. Each lab limited to diversity, and four credits of research in information based sciences of genetics, 36 students. Prerequisite: concurrent or neurobiology and behavior. molecular biology, and the emerging previous enrollment in BIOBM 330 or 331 biotechnologies to the study of human and 332 or 333. Evening prelims. Lees, M physiology will be covered. ANIMAL PHYSIOLOGY 155

W F 11:15; lab, MorT 1:25-4:25. membrane and epithelial transport; electro­ [BIOAP 715 Stress Physiology: To Be A. Quaroni and staff. physiology; cardiovascular physiology; Discussed as Part of Animal Welfare Lectures introduce students to the most gastrointestinal physiology; renal physiology; Fall. 1 credit. Prerequisite: BIOAP 311 or current information on the way cells function and acid/base physiology. The lectures equivalent required. Offered alternate apd regulate themselves and neighboring incorporate clinical correlations whenever years. Next offered fall 2002. K. A. Houpt. c)blls, and on what molecules are involved in appropriate. Occasional guest lecturers talk The emphasis will be on physiological t|iese regulatory processes. Laboratories about work and careers in basic research and/ assessment of stress.] fjrovide an introduction to cell and organ or clinical medicine. Recommended for culture and to immunological techniques used BIOAP 719 Graduate Research in Animal biological sciences majors, pre-med and pre- Physiology (also VETPH 628) t(j> study cell structure and function in vivo and vet students, and beginning graduate students ik vitro. Experiments performed in the in physiology, nutrition, and animal science. Fall or spring. Variable credit. Prerequi­ laboratory are closely related to, and provide sites: written permission of the section practical experience with, subjects covered in BIOAP 475 Mechanisms Underlying chair and of the staff member who the lectures. Vertebrate animals are used in Mammalian Developmental Defects supervises the work and assigns the grade. (also NS 475) this course. No experimentation is performed S-U grades optional. Hours TBA. Staff. on live animals. Spring. 3 credits. Prerequisites: BIOBM Similar to BIO G 499 but intended for 330, 331-332, or 333 (may be taken graduate students who are working with BIOAP 319 Animal Physiology concurrently). M W 9:10; lab, R 2:00. faculty members on an individual basis. Experimentation D. Noden and P. Stover. Fall. 4 credits. Designed for upper-level For description, see NS 475. BIOAP 757 Current Concepts in undergraduate and graduate students Reproductive Biology studying in physiology, and other students BIOAP 619 Lipids (also NS 602) Fall. 3 credits. Limited to 20 students. interested in biomedically related Fall. 2 credits. Lees, T R 11:15. Prerequisites: undergraduate degree in professions. Graduate students in the field A. Bensadoun. biology and a strong interest in reproduc­ of Physiology and related fields without Advanced course on biochemical, metabolic, tive biology. S-U grades optional. Lec/disc, equivalent background are strongly and nutritional aspects of lipids. Emphasis is T R 10:10-12:05. Offered alternate years. encouraged to enroll. Each of 2 afternoon placed on critical analysis of current topics in J. E. Fortune, W. R. Butler, and staff. laboratory sections is limited to 40 lipid methodology; lipid absorption; lipopro­ A team-taught survey course in reproductive students. Prerequisite: concurrent or tein secretion, molecular structure, and physiology/endocrinology. Lectures by a previous enrollment in BIOAP 311 or catabolism; molecular biology, function and number of reproductive biologists on various permission of instructor. Lee, R 12:20; lab, regulation of lipoprotein receptors; mecha­ aspects of male reproductive function M or W 12:20-5:00 (includes disc section). nism of hormonal regulation of lipolysis and (endocrine regulation, testis function, E. R. Loew and staff. fatty acid synthesis; and cholesterol metabo­ spermatogenesis, and sperm physiology/ A series of student-conducted in vitro and in lism and atherosclerosis. function); female reproductive function vivo experiments designed to illustrate basic (endocrinology, ovarian development and BIOAP 710-718 Special Topics in functions, oocyte physiology/function); physiological processes in animals, with Physiology emphasis on relevance to humans, and to fertilization and early embryo development; Fall or spring. 1 or 2 credits for each topic. pregnancy; parturition; puberty; and reproduc­ introduce students to physiology research May be repeated for credit. Enrollment in techniques, instrumentation, experimental tive technology. Student participation in the each topic may be limited. S-U grades form of discussions and/or presentations. design, and interpretation of results. Tech­ optional, with permission of instructor. niques include anesthesia, surgical proce­ Lectures, laboratories, discussions, and BIOAP 811 Advanced Physiological dures, vivisection under anesthesia, and real­ seminars on specialized topics. Methods I time computer recording and analysis of data. Fall. 2 credits. Enrollment limited. Experiments with living tissues and live [BIOAP 711 Readings in Applied Animal Behavior Prerequisites: graduate student status or animals examine properties of blood, muscle, permission of course coordinator. S-U and nerves; cardiovascular, respiratory, and Fall. 1 credit. Prerequisite: BIOAP 311 or grades only. Lab TBA. Coordinator: J. Ray. renal function and their control; and endo­ equivalent. Offered alternate years. Next This is a course for graduate students in crine regulation of renal, cardiovascular, and offered in 2002. Lee, 1 hour each week physiology and related disciplines. Experi­ reproductive tissue activity. Experimental TBA. K. A. Houpt.) ments are carried out in the laboratories of resources include live animals, frogs, rats, BIOAP 712 Thermoregulation and faculty members in the Field of Physiology to rabbits, and sheep which are not always Exercise acquaint graduate students with the latest euthanized after the laboratory exercises. Fall. 1 credit. Offered alternate years. techniques/methods in biomedical/physiologi- Written reports of laboratory activities are D. Robertshaw. cal research. Three modules are offered each required. Grading is based on evaluation of An examination of the competing demands on semester by arrangement with the course these reports, take-home case studies, the body of exercise and heat exposure with coordinator. laboratory performance, and weekly quizzes. particular emphasis on the cardiopulmonary BIOAP 812 Advanced Physiological BIOAP 427 Fundamentals of system and integration of thermoregulatory Methods II reflexes. Endocrinology (also AN SC 427) Spring. 2 credits. Enrollment limited. Fall. 3 credits. Prerequisite: animal or [BIOAP 713 The Physiology of Ingestive Prerequisites: graduate student status or human physiology or permission of Behavior: Food and Water Intake permission of course coordinator. S-U instructor. Lees, M W F 9:05. P. A. Johnson. Fall. 1 credit. Prerequisite: introductory grades only. Lab TBA. Coordinator: J. Ray. For description, see AN SC 427. biology. Next offered in fall 2002. This is a course for graduate students in BIOAP 458 Mammalian Physiology T. R. Houpt. physiology and related disciplines. Experi­ Spring. 3 credits. Enrollment limited. A series of f to 1.5 hour weekly sessions in ments are carried out in the laboratories of Graduate student auditors allowed. which the physiological mechanisms of faculty members in the Field of Physiology to Prerequisite: BIOAP 311 or equivalent. hunger and thirst, primarily in mammals, will acquaint graduate students with the latest Students not meeting this prerequisite must be considered by the group. There will be a techniques/methods in biomedical/physiologi- ■ obtain written permission of instructor in mix of background readings, individual cal research. Three modules are offered each T8 014 Vet Research Tower before the first reports, and general discussion.) semester by arrangement with the course coordinator. class. Evening prelims. Lees, M W F 10:10. [BIOAP 714 Cardiac Electrophysiology K. W. Beyenbach and staff. Fall. 1 credit. Offered alternate years. Next The course offers an in-depth treatment of Related Courses in Other Departments offered fall 2002. R. Gilmour. selected topics in mammalian and human Adaptations of Marine Organisms (Biological Survey of cardiac potentials, passive mem­ physiology. Emphasis is on concepts and a Sciences [BIOSM] 413) brane properties, ion channels, and cardiac working knowledge of physiology. Selected arrhythmias. Emphasis on nonlinear dynamical Advanced Work in Animal Parasitology (Veterinary topics include: basic functional elements of aspects of cardiac electrophysiology and Microbiology 737) biological systems; recurrent themes in cardiac arrhythmias.] physiology; design of multicellular animals; Animal Development (Veterinary Anatomy 507) mammalian fluid compartments; homeostasis; Animal Reproduction and Development (Animal Science 300) BIOLOGICAL SCIENCES - 2001-2002

Developmental Biology (Biological Sciences some familiarity with iteration, arrays, and BIOBM 330, 331, or 332. Lees, M W F [BIOGD] 385) procedures. 10:00-12:00. S. Ely or H. T. Nivison. Embryology (Biological Sciences [BIOGD] 389) For course description see COM S 321. The content of this course is similar to that of BIOBM 330, however, it is presented in Fundamentals of Endocrinology (Animal BIOBM 330-332 Principles of lecture format rather than as individualized Science 427) Biochemistry instruction. The topics include the structure Introductory biochemistry is offered in three Insect Morphology (Entomology 322) and function of proteins, enzyme catalysis, formats: individualized instruction (330) and metabolism, and the replication and expres­ Integration and Coordination of Energy Metabolism lectures (331 and 332) during the academic sion of genes. (Biological Sciences [BIOBM] 637 and year, and lectures (333) during the summer. Nutritional Sciences 636) Individualized instruction is offered to a BIOBM 334 Computer Graphics and Neuroanatomy (Veterinary Anatomy 504) maximum o f250 students each semester. Molecular Biology Lectures are given fa ll semester (33 V, spring Fall or spring. 1 credit. Prerequisite: Sensory Function (Biological Sciences semester (332), and summer (333). concurrent registration in BIOBM 330. If [BIONB] 492) space permits, students who have Teaching Experience (Biological Sciences BIOBM 330 Principles of Biochemistry, completed BIOBM 331 or 333 and have Individualized Instruction [BIO G] 498) either taken or are concurrently taking, Fall or spring. 4 credits. Prerequisites: one BIOBM 332 will be permitted to register Undergraduate Research in Biology (Biological year of introductory biology for majors and Sciences [BIO G] 499) during the first week of classes. one year of general chemistry and CHEM J. E. Blankenship, P. C. Hinkle, and staff. 257 or 357-358 (CHEM 358 may be taken Visualization of complex biomolecules using concurrently) or equivalent, or permission Silicon Graphics computers. Group presenta­ of instructor. Concurrent registration in tions on current topics in molecular biology. BIOCHEMISTRY, MOLECULAR AND BIOBM 334 is encouraged. May not be CELL BIOLOGY (BIOBM) taken for credit after BIOBM 331, 332, or BIOBM 407 Nature of Sensing and 333- S-U grade optional for graduate Response: Signal Transduction in BIOBM 132 Orientation Lectures in students only. Evening prelims: fall, Oct. Biological Systems (also PLPA 407) Biochemistry 11 and Nov. 8; spring, Feb. 26 and Apr. 4. Spring. 3 credits. Prerequisite: BIOBM 330 Spring, weeks 1-3. No credit. Primarily for Hours'TBA. J. E. Blankenship, P. C. Hinkle, or 333 or 331 and previous or concurrent freshmen, sophomores, and transfer and staff. registration in 332. Recommended: BIOGD students. S-U grades only (registered Fourteen units that cover protein structure and 281. S-U grades optional. Lees T R 10:10- students receive an unsatisfactory grade for function, enzymes, basic metabolic pathways, 11:25. T. P. Delaney. nonattendance). Lee, S 10:10-11:00, for DNA, RNA, protein synthesis, and an The responses of organisms and cells to their first three S of semester. Section chair and introduction to gene cloning. No formal surroundings are examined to illustrate how staff. lectures, autotutorial format. biological systems sense their biotic and Discussions by six professors about their abiotic environment and Communicate sensing BIOBM 331 Principles of Biochemistry: into appropriate responses. A wide variety of research and promising areas for research in Proteins and Metabolism the future. response systems will be explored to identify Fall. 3 credits. Prerequisites: one year of their unique features and to illustrate how BIOBM 232 Milestones In Molecular introductory biology for majors, one year similar processes are utilized by widely Biology and Genetics of general chemistry, and CHEM 257 or divergent organisms. Examples are drawn Spring. 3 credits. Prerequisites: Advanced 357-358 (CHEM 257 or 357 should not be from prokaryote, plant, and animal systems Placement Biology. Letter grade. Lees, M taken concurrently) or equivalent, or for environmental sensing, control of W 9:05. Disc, F 9:05. J. M. Lopez. permission of instructor. May not be taken development, and responses during disease. Lectures cover seminal genetic, biochemical, for credit after BIOBM 330 or 333. S-U Discussion will also examine the role of and cell and molecular biological experiments grades with permission of instructor. genetics and biochemistry in understanding that were recognized by Nobel prizes. Evening prelim: Oct. 25. Lees, M f F signal transduction pathways, as well as the Discussion will focus on current scientific 10:10. G. W. Feigenson. way these systems are perturbed by mutation consequences of these seminal findings, The chemical reactions important to biology, and disease. together with their societal and/or economic and the enzymes that catalyze these reactions, ramifications. are discussed in an integrated format. Topics BIOBM 432 Survey of Cell Biology include protein folding, enzyme catalysis, Spring. 3 credits. Prerequisite: BIOBM 330, [BIOBM 233 Introduction to bioenergetics, and key reactions of synthesis 333, or 331, and previous or concurrent Biomolecular Structure (also and catabolism. registration in 332, or equivalent. CHEM 233) Recommended: BIOGD 281. S-U grades Fall. 2 credits. Limited to 30 students. BIOBM 332 Principles of Biochemistry: optional for graduate students only. Lees, Prerequisites: CHEM 207-208 or equiva­ Molecular Biology M W 8:40-9:55. W. J. Brown, V. M. Vogt. lents. Lees, T R 2:30-3:20. Next offered fall Spring. 2 credits. Prerequisites: one year of A survey of a wide array of topics focusing on 2002. S. E. Ealick. introductory biology for majors and the general properties of eucaryotic cells. The This course is intended for students with a previous or concurrent registration in topics include methods used for studying basic understanding of chemistry who are organic chemistry, or permission of cells, the structure and function of the major considering a program of study in biochemis­ instructor. May not be taken for credit after cellular organelles, and analyses of cellular try. The interrelationship between the BIOBM 330 or 333- S-U grades optional, processes such as mitosis, endocytosis, cell structure and function of biologically with permission of instructor. Lees, T R motility, secretion, cell-to-cell communication, important molecules is explored. Emphasis is 12:20. B. K. Tye. gene expression, and oncogenesis. Some of placed on understanding the way in which the A comprehensive course in molecular biology the material is covered in greater depth in three-dimensional arrangements of atoms that covers the structure and properties of BIOBM 437; BIOGD 483; and BIOBM 632, determine the biological properties of both DNA, DNA replication and recombination, 636, and 639. small molecules and macromolecules such as synthesis and processing of RNA and proteins, proteins and enzymes. The study of molecular the regulation of gene expression, and the BIOBM 434 Applications of Molecular structure is aided by interactive computer principles and uses of recombinant DNA Biology to Medicine, Agriculture, and graphics for visualizing three-dimensional technologies. Industry structures of molecules.] Fall. 3 credits. Enrollment limited to 36 BIOBM 333 Principles of Biochemistry: students. Prerequisites: BIOBM 330 or 333 BIOBM 321 Numerical Methods in Proteins, Metabolism, and Molecular or 331 and 332. Recommended: BIOGD Computational Molecular Biology Biology 281. S-U grades optional. Lees, M W F (see COM S 321) Summer (8-week session). 4 credits. 11:15. J. M. Calvo, S. Ely. Fall. 3 credits. Prerequisites: at least 1 Prerequisites: one year of introductory Lecture topics include large scale sequencing course in calculus, such as MATH 106, 111, biology for majors, one year general of genomes, drug discovery based upon or 191 and a course in linear algebra, such chemistry, and CHEM 257, or 357-358, or genomics, pharmaco-genomics, mapping and as MATH 221 or 294 or BTRY 417. No equivalents, or permission of the instruc­ cloning human disease genes, DNA and edible particular course in programming is tor. May not be taken for credit after vaccines, transgenic animals, engineering required, but the student should have BIOCHEMISTRY, MOLECULAR AND CELL BIOLOGY 157 plants resistant to insects, and gene therapy. insensitivity syndromes, inborn errors of components, immunofluorescence and Discussions will relate to problem sets and to metabolism, gene fusions resulting in hybrid electron microscopy, and in vitro assays. some implications of molecular biology to proteins, gene amplification, gene inactivation, society. disruption of signaling pathways, disruption of [BIOBM 443 Experimental Molecular metabolic pathways, and the molecular Neurobiology (also BIONB 430) BIOBM [435J-436 Undergraduate Spring. Next offered spring 2003. Biochemistry Seminar actions of environmental poisons and toxins. Examples of diseases will be selected to D. Deitcher. 435, fall; 436, spring. 1 credit each term. emphasize various aspects of cell biology, Experiments include PCR, cloning of DNA May be repeated for credit. Limited to physiology, and immunology that have been fragments, RNA purification, restriction digests, upperclass students. Prerequistites: BIOBM presented in other courses. In addition, the bacterial transformation, and DNA sequencing. 330, 333, or 331 and 332 or written methods used to identify the underlying Experiments emphasize how molecular permission of instructor. S-U grades only. biochemical and genetic basis of the diseases, techniques can be applied to studying Seminar time TBA. Organizational meeting as well as possible pharmaceutical and genetic neurobiological problems.] first W of each semester at 4 p.m. Next therapies for treating the diseases, will be BIOBM 450 Lab Projects in Molecular offered fall 2002. Fall: G. P. Hess; spring: presented. staff. Biology and Biochemistry Selected papers from the literature on a given BIOBM 440-443 Laboratories in Spring. 3 credits. Enrollment limited to 10 topic are evaluated critically during six or Molecular Biology, Biochemistry, students. Prerequisites: 4 credits from any seven two-hour meetings. Cell Biology and Molecular of the following lab courses; BIOBM 440, Neurobiology 441, 442, or 443 plus permission of BIOBM 437 Eukaryotic Cell Proliferation Laboratory instruction in these areas is instructor. Letter grade only. Lab, R 12:20- (also TOX 437) organized as a set of 2 credit modules, 4:25 plus at least 3 additional lab hours by Fall. Variable credits. Students may take each module is taught for half a semester individual arrangement with the instructor. lectures for 2 credits, or take both lectures during the academic year and some Disc, F 2:30. S. Ely. and discussions for 3 credits. Enrollment modules are taught during the summer. Students work individually on projects of their for discussion section is limited to 20 Students may sign up for two courses for a choice pertaining to a common theme. Skills students, with preference given to total of 4 credits; limited space is available developed include literature and database graduate students. Prerequisite: BIOG 101- for students taking only one course for 2 searching, project and experimental design, 102 or BIOG 105-106 and BIOBM 330 or credits. Which courses are offered in each keeping a legal notebook, weekly communi­ BIOBM 331-332. Recommended: semester depends on scheduling con­ cation of results, and an oral presentation. BIOGD281 and BIOGM 432. S-U grades straints and student preferences. Enroll­ Experimental techniques may include site- optional. Lees, T R 12:20-1:10. Disc, TBA. ment limited, with preference given to directed mutagenesis, affinity purification, R.-H. Chen undergraduates having Biochemistry or whole-genome expression systems, molecular The course covers a wide spectrum of issues Molecular and Cell Biology Programs of evolution of a compound, and exploration of related to cell proliferation in eukaryotes. Study and to graduate students with a the human genome. Lectures include various aspects of the minor in the Field of Biochemistry, regulation of cell division cycle and signal BIOBM 631 Protein Structure and Molecular and Cell Biology. Prerequisites: Function transduction pathways, with additional topics BIOBM 330, or 333, or 331 and previous on oncogenesis, cell aging, and cell death. Fall. 3 credits. Prerequisites: BIOBM 330 or or concurrent enrollment in 332, or 332 333 or 331 and 332 and organic chemistry. The facts as well as concepts and logics and previous or concurrent enrollment in behind findings are presented in the lectures. Recommended: physical chemistry. S-U 331, and permission of instructor. Form to grades optional. Lees, M W F 9:05. Research articles are analyzed and discussed apply for admission to this course is in depth during discussion section. L. Nicholson. found on the web [http:// Presentations on the principles of protein BIOBM 438 Seminar in Applied Molecular 132.236.135.25/signup.html]; for Fall structure and the nature of enzymatic Biology semester courses, apply from February catalysis. Specific topics include protein Fall. 2 credits. Enrollment limited to 20 15 until one week before CoursEnroll folding, stability, dynamics and evolution, students. Prerequisites: BIOBM 330 or 333 begins; for Spring semester courses, folded conformations and structure prediction, or 331, and 332. Recommended: BIOGD apply from September 15 until one ligand binding energetics, and the structural 281. S-U grades optional. Sem. R 12:20- week before CoursEnroll begins basis of catalysis. 2:15. J. Calvo, S. Ely. Confirmation of enrollment status will be A seminar course focusing on topics such as: sent by e-mail during the week preceding BIOBM 632 Membranes and drugs that extend the life of AIDS patients; CoursEnroll. Fall and Spring: Labs, M W Bioenergetics applications of human embryonic stem cell 12:20-4:25 (disc, F 1:25) or T 9:05-4:25 Spring. 2 credits. Prerequisite: BIOBM 330 technology; genetically engineering plants for (disc, R 11:15 or 1:25) or R 9:05-4:25 (disc, or 333 or 331 and 332 or equivalent. resistance to insects; bringing erythropoeitin to T 1:25). Summer (3-week session): M-F Lees, T R 11:15. Offered alternate years. market; mining genetic variation within 10-5:30. P. C. Hinkle. isolated human populations; and benefits from Structure and dynamics of biological BIOBM 440 Experimental Molecular membranes, physical methods, model stimulating and blocking angiogenesis. Groups Biology of students will explore the underlying science membranes, ionophores, ion-transport 2 credits. S. Ely or H. T. Nivison. ATPases, mitochondrial and chloroplast and will also consider some business and/or Experiments include cloning of DNA social implications for these topics. electron transfer chains, and examples of fragments, restriction mapping, DNA transport from plants, animals, and bacteria. BIOBM 439 Molecular Basis of Human sequencing, Southern blotting, and PCR. The Emphasis given to structure of membrane Disease experiments emphasize quantitative aspects as proteins. Fall. 2 credits. Prerequisites: BIOBM 330 or well as experimental design. BIOBM 331-332 and genetics Ce.g., BIOGD BIOBM 633 Biosynthesis of BIOBM 441 Experimental Proteins and Macromolecules 281) or permission from instructor. Enzymology Recommended: cell biology (e.g., BIOBM Fall. 2 credits. Prerequisite: BIOBM 330 or 2 credits. S. Ely or H. T. Nivison. 333 or 331 and 332. Recommended: 432 or BIOAP 316) and physiology (e.g., • Experiments include purification of enzymes BIOAP 311 or BIOAP 458). S-U grades BIOGD 281. Lees, T R 9:05. J. W. Roberts, by salt fractionation, ion exchange chromatog­ D. B. Wilson. optional. Lees, T R 11:15. W. K. Krause. raphy, affinity chromatography, determination This course will examine how changes in the Synthesis of DNA, RNA, and proteins, and of kinetic parameters for an enzyme, analysis regulation of gene expression. normal expression, structure, and activity of of proteins by rate zonal sedimentation, SDS- gene products caused by genetic mutations polyacrylamide gel electrophoresis, and BIOBM 636 Advanced Cell Biology and environmental agents lead to human immunoblotting. Spring. 2 credits. Prerequisites: BIOBM diseases. The material will focus on how 330 or 333 or 331 and 332, and 432, or BIOBM 442 Experimental Cell Biology proteins with modified structures and their equivalents. Lee, T R 9:05-9:55. biochemical activities cause alterations in Spring only. 2 credits. T. Huffaker. A. P. Bretscher. normal cellular processes, as well as the Experiments include culture of animal cells, With the availability of whole genome physiological consequences of these changes. purification and analysis of subcellular sequences, new families of genes are being Topics will be selected from hormone identified. It is the goal of functional genomics 158 BIOLOGICAL SCIENCES - 2001-2002

to elucidate the role of the gene products in BIOBM 751 Ethical Issues and majoring in the Field of Biochemistry, the functional organization of cells. This Professional Responsibilities Molecular and Cell Biology. S-U grades course provides an integrated view of how Spring. 1 credit. Limited to graduate only. Sem and disc TBA. Fall: G. P. Hess; this can be achieved employing molecular, students beyond first year. S-U grades spring: J. Roberts. genetic, and cell biological approaches. The only. Organizational meeting will be held A seminar course with critical discussion by discussion will center around a detailed on the first W of the semester. Sem, W students of original research papers. A variety discussion of topics such as the cytoskeleton, 3:35—4:25 Additional sections may be of topics in biochemistry, molecular and cell secretion, endocytosis, cell polarity, and offered. P. Hinkle. biology are covered. related topics. Together with BIOBM 437, 632, Ethical issues in research and the professional and 639 this course provides broad coverage responsibilities of scientists are discussed in a Related Courses in Other Departments of the cell biology subject area. case-study format. Topics to be discussed Lipids (Biological Sciences [BIOAP] 619 and include: regulations; data selection, manipula­ Nutritional Sciences 602) BIOBM 639 The Nucleus tion, and representation; fraud, misconduct, Spring. 2 credits. Prerequisite: BIOBM 330 or and whistle-blowing; conflicts of interest and Molecular Aspects of Development (Biological 333 or 331 and 332, and 434, or their commitment; authorship, ownership, and Sciences [BIOGD] 483) equivalent. Recommended: BIOGD 281. Lee, intellectual properties; peer review and Molecular Biology Techniques for Animal Biologists T R 10:10. J. T. Lis. confidentiality; scientific response to external (Animal Science 650) Lectures on topics of eucaryotic gene pressure; legal liabilities; and professional Molecular Mechanisms of Hormone Action organization, regulation of gene expression, codes of ethics. RNA processing, chromatin structure, the (Biological Sciences [BIOAP] 658 and Veterinary Medicine 758) structure and movement of chromosomes, and BIOBM 761 Topics in Cancer Cell Biology the architecture of the nucleus. This course (also VETMM 761) Teaching Experience (Biological Sciences covers the structure and function of the Spring. Series of 1 credit graduate sections [BIO G] 498) that reflect the “cancer expertise” of the nucleus at the molecular and cell biological Undergraduate Research in Biology (Biological levels, and together with BIOBM 437, 632 and Cornell faculty. (Course Director: Sciences [BIO G] 499) 636, provides broad coverage of the cell Dr. B. U. Pauli). biology subject area. For description, see VETMM 7 6 l. BIOBM 641 Laboratory in Plant BIOBM 830 Biochemistry Seminar Molecular Biology (also BIOPL 641) Fall or spring. No credit. Sem, F 4:00. Staff. ECOLOGY AND EVOLUTIONARY Fall. 4 credits. Prerequisites: BIOGD 281 or Lectures on current research in biochemistry, BIOLOGY (BIOEE) equivalent, BIOBM 330 or 331 or presented by distinguished visitors and staff equivalent, and permission of instructor. members. Lectures are open to everyone, but BIOEE 154 The Sea: An Introduction to S-U grades with permission of instructor. registration is limited to graduate students in Oceanography (also EAS 104) Lab, T 9:05-4:30. J. B. Nasrallah, Biochemistry, Molecular and Cell Biology. Spring. 3 or 4 credits (4-credit option M. R. Hanson. BIOBM 831 Advanced Biochemical includes one 2 1/2 hour laboratory each Selected experiments on gene expression, Methods I week). S-U grades optional. Lees, T R gene transfer, and assay of reporter genes in 11:40-12:55; lab, M or W 2:00-4:25, or M Fall. 6 credits. Required of, and limited to, plants. The course emphasizes the application 7:30-9:55 P M. C. H. Greene, W. M. White. first year graduate students in the Field of of molecular biology methodology to plant A survey of the physics, chemistry, geology, Biochemistry, Molecular and Cell Biology. systems. Additional lab time is required to and biology of the oceans for both science S-U grades only. Labs and discussions 12 complete assignments. and non-science majors. Topics include: sea­ hours each week TBA. Organizational floor spreading and plate tectonics, marine meeting first R of semester, 10:10. BIOBM 730 Protein NMR Spectroscopy sedimentation, chemistry of seawater, ocean (also VETPR 730) V. M. Vogt and staff. currents and circulation, the oceans and The first half of this course comprises an Spring. 2 credits. Prerequisites: CHEM 389 climate, ocean ecology, coastal processes, intensive laboratory covering fundamental and 390 or CHEM 287 and 288 or marine pollution, and marine resources. permission of instructor. S-U grades aspects of modern molecular biology and cell optional. Lee TBA. L. K. Nicholson, biology. The second half of the course BIOEE 207 Evolution (also HIST 287 and R. E. Oswald. compromises research in the laboratory of a S&TS 287) The student acquires the tools necessary for professor chosen by the student (See BIOBM Fall or summer. 3 credits. Intended for in-depth understanding of multidimensional, 832). Students must enroll separately for each students with no background in college multinuclear NMR experiments. Schemes for half. biology. May not be taken for credit after magnetization transfer, selective excitation, BIOBM 832 Advanced Biochemical BIOEE 278. Does not meet the evolution­ water suppression, decoupling, and others are Methods II ary biology requirement for the biological presented. The application of these techniques sciences major. S-U grades optional. Fall: Spring. 6 credits. Required of, and limited to proteins for resonance assignment, structure Lees, T R 10:10; disc, 1 hour each week to, first year graduate students in the Field determination, and dynamics’ characterization TBA. Summer (6-week session): Lees and of Biochemistry, Molecular and Cell discs, M W 6:00-9:00 p.m. W. B. Provine. is studied. Biology. S-U grades only. Lab TBA. Staff. Evolution is the central concept in biology. Research in the laboratories of two different BIOBM 732-737 Current Topics in This course examines evolution in historical Biochemistry professors chosen by the student. Arrange­ and cultural contexts. This course aims to ments are made jointly between the Director Fall or spring. 1/2 or 1 credit for each understand the major issues in the history and of Graduate Studies and the research adviser. topic. May be repeated for credit. current status of evolutionary biology, and Prerequisite: BIOBM 330 or 333 or 331 and BIOBM 833 Research Seminar in explore the implications of evolution for 332 or equivalent. S-U grades only. Biochemistry culture. Issues range from controversies over Lectures and seminars on specialized topics. Fall or spring. 1 credit each term. May be mechanisms of evolution in natural popula­ Topics for fall and spring to be announced in repeated for credit. Required of, and tions to the conflict between creationists and the division’s course supplement published at limited to, second-, third-, and fourth-year evolutionists. the beginning of each semester. graduate students majoring in biochemis­ BIOEE 261 Ecology and the Environment try. S-U grades only. Sem, M 12:20-1:30. BIOBM 738 Macromolecular Fall or summer. 4 credits. Prerequisite: one Crystallography (also CHEM 788) T. C. Huffaker. Each student presents one seminar per year year of introductory biology. S-U grades Spring. 3 credits. S-U grades optional. optional. Lees, M W F 11:15; disc, W or R Prerequisite: permission of instructor. Lees, on his or her thesis research and then meets with instructors and thesis committee 1:25, 2:30, or 3:35- N. G. Hairston and staff. M W F 10:10. Offered alternate years. An introduction to the science of ecology, the S. E. Ealick. members for evaluation. study of interactions between organisms and Lectures briefly cover the fundamentals of BIOBM 836-837 Methods and Logic in their environments. Major topics include crystallography and focus on methods for Biochemistry, Molecular and Cell demography, succession, biodiversity, determining the three-dimensional structures Biology biogeochemistry and ecosystems, and the of macromolecules. 836, spring; 837, fall. 1 credit each term. evolution of adaptations. The influences of Limited to first-year graduate students enemies, competitors, and mutualists on ECOLOGY AND EVOLUTIONARY BIOLOGY 159

populations and communities are discussed. Laboratories include dissections of preserved introductory biology for majors. Permission The effects of climate and human activities on vertebrate animals and noninvasive live of faculty required because it is off ecological processes are also considered. animal demonstrations. campus. Students in BIOEE 373 are Ecological principles are used to explain the strongly encouraged to take BIOEE 477. issues associated with several environmental [BIOEE 275 Human Biology and Evolution (also ANTHR 275 and NS Three week, full time course. Daily and problems. 275) evening lectures, laboratories, and field work. Course is taken during the summer; BIOEE 263 Field Ecology Fall. 3 credits. S-U grades optional, with enroll for credit during the subsequent fall permission of either instructor. Lees, M W Fall. 3 credits. Limited to 25 students. semester. Total cost for room, board, and F 10:10; disc, M 10:10 or TBA. Lees every Prerequisite: concurrent or previous overhead at SML: $1,150. Offered alternate W and F; occasional lectures on M. enrollment in BIOEE 261. Lee, R 1:25; lab, years. C. D. Harvell, J. G. Morin. Offered alternate years. Not offered 2001- F 12:20-4:25; 1 weekend field trip to the An introduction to the biology and evolution 2002. K. A. R. Kennedy, J. D. Haas. Hudson Valley. P. L. Marks. of the major invertebrate phyia, concentrating An introduction to the biology of H om o Field exercises designed to give students on marine representatives. In addition to the sa p ien s through an examination of human direct experience with field work, with evolution of form and function, lectures cover evolution, biological diversity, and modes of emphasis on developing observational skills, aspects of ecology, behavior, physiology, adaptation to past and present environments. journal keeping, and a landscape perspective. chemical ecology, and natural history of Evolutionary theory is reviewed in relation to Topics include plant succession, niche invertebrates. The Shoals Marine Laboratory the current evidence from the fossil record relationships of insects, influence of herbi­ exposes students to a wealth of marine and and studies of the evolution of human vores and competition on plant performance, terrestrial invertebrates in their natural behavior. A survey of human adaptation decomposition of soil litteq foraging behavior, habitats. Regular field excursions allow an covers a complex of biological and behavioral census methods, and use of scientific excellent opportunity to study freshly responses to environmental stress. Human collections. collected and in situ representatives of most diversity is examined as the product of long­ of the major phyla. BIOEE 264 Birds in Biology term evolutionary forces and short-term Fall. 3 credits. Limited to 25 students. May adaptive responses. Topics such as creation­ BIOEE 405 Biology of the Neotropics not be taken for credit after BIOEE 475. ism, the Piltdown fraud, the sociobiology Fall. 2 credits. Prerequisite: introductory Intended primarily for biology nonmajors. debate, genetic engineering, race and IQ, and biology (majors, non-majors, or equiva­ S-U grades optional. Lees and discs, T R racism are presented as examples of current lent), or permission of instructor. S-U 8:40-9:55; 2 field trips TBA. Offered issues in human biology.] grades optional. Lee and disc, W 7:30-9:30 alternate years. A. A. Dhondt. BIOEE 278 Evolutionary Biology p.m. P. H. Wrege, A. S. Flecker. This course explores exciting new insights in This course is an introductory survey of the Fall or spring. 3 or 4 credits. (4-credit biology using detailed examples drawn from biology of the New World tropics, with option involves writing component and bird studies. Subject matter is suitable for non­ primary focus on moist lowland forests. The two discussion sections per week; limited majors, but of interest to majors as well. objectives are to learn basic characteristics and to 20 students per section each semester. Topics will be drawn from a variety of phenomena important to understanding Students may not preregister for the biological disciplines. These include behav­ neotropical biology, to gain firsthand 4-credit option; interested students ioral ecology (mating systems, territorial knowledge of the resources in tropical biology complete an application form on the first behavior, song), population ecology (migra­ available at Cornell, and to learn how to day of class.) Limited to 300 students. tion, population limitation, micro-evolution, organize and execute a meaningful seminar Prerequisite: 1 year of introductory biology competition), evolutionary biology (trade-offs presentation. in life histories, optimal clutch size), and or permission of instructor. S-U grades conservation biology (habitat fragmentation, optional. Evening prelims: fall, Sept. 27 [BIOEE 452 Herbivores and Plants: inbreeding, acid rain). Lectures will be and Nov. 1; spring, Feb. 28 and Apr. 2. Chemical Ecology and Coevolution interspersed with discussion of selected Lees, T R 9:05; disc, 1 hour each week (also ENTOM 452) papers. TBA. Fall, M. Geber; spring, M. Shulman. Spring. 3 credits. Prerequisites: one year of The course considers explanations for patterns introductory biology, BIOEE 261, CHEM BIOEE 267 Introduction to Conservation of diversity and for the apparent “good fit” of 257 or 357/358 and 251 or 301, or Biology organisms to the environment. Topics covered permission of instructor. S-U grades Fall. 3 credits. May not be taken for credit include the genetic and developmental basis optional. Field trips, additional lectures, or after NTRES 450. Intended for both science of evolutionary change, processes at the laboratory demonstrations may be held in and non-science majors. Completion of population level, the theory of evolution by place of F lecture. Lees, M W F 11:15. BIOEE 267 is not required for NTRES 450. natural selection, levels of selection, concepts Offered alternate years. Not offered 2001- S-U grades optional. Lees, M W 9:05; disc, of fitness and adaptation, modes of speciation, 2002. P. P. Feeny. F 9:05 or R 2:30; 1 Saturday field trip. long-term trends in evolution, rates of Topics include: significance of plant chemistry A. S. Flecker, J. W. Fitzpatrick. evolution, and extinction. Students taking the in mediating interactions between plants and An exploration of biological concepts related four-credit option read additional materials herbivorous animals; mechanisms and to conserving the earth’s biodiversity, from the primary literature and write a series strategies of plant finding and exploitation by introducing ecological and evolutionary of essays in place of the regular prelims. animals, especially insects, and of defense and principles important for understanding major escape by plants; evolutionary hypotheses for conservation problems. Topics include [BIOEE 371 Human Paleontology (also ANTHR 371) ecological patterns of resistance and attack; patterns of species and ecosystem diversity, and implications for human food and Fall. 4 credits. Prerequisite: one year of causes of extinction, genetic risks of small agriculture.] populations, design of nature preserves, introductory biology or ANTHR 101 or strategies for protecting endangered species, permission of instructor. Lees, M W F 2:30; BIOEE 455 Insect Ecology (also ecosystem restoration, and the value of lab, 1 hour each week TBA; occasional ENTOM 455) biodiversity. field trips. Offered alternate years. Not Fall. 3 credits. Prerequisites: BIOEE 26l or offered 2001-2002. K. A. R. Kennedy. equivalent and ENTOM 212 or knowledge BIOEE 274 The Vertebrates: Structure, A broad survey of the fossil evidence for of another taxon. S-U grades optional. Function, and Evolution human evolution with special attention to Lees, M W F 11:15. Offered alternate years. Spring. 4 credits. Prerequisite: one year of skeletal and dental anatomy, geological R. B. Root. introductory biology. Fee, $25. S-U grades contexts, paleoecology, dating methods, Topics include the nature and consequences optional. Lees, M W F 9:05; lab, M T W or archaeological associations, and current of biotic diversity, biogeography, coevolution, R 1:25-4:25. K. R. Zamudio. theories of human origins and physical adaptive syndromes exhibited by various An introductory course in vertebrate diversity. guilds, population regulation, impact of organismal biology which explores the insects on ecosystems, comparative and structure and function of vertebrates with an BIOEE 373 Biology of the Marine Invertebrates functional analysis of communities, and emphasis on trends in vertebrate evolution. differences in the organization of natural and Fall (but course must be taken in the Lectures will cover topics such as the origin managed systems. Ecological and evolutionary previous summer at the Shoals Marine and evolution of various vertebrate groups, principles are integrated by thorough study of sensory systems, thermoregulation, life history, Laboratory [SML]). 5 credits. Limited to 24 exemplars. locomotion, feeding, size, and scaling. students. Prerequisite: one year of BIOLOGICAL SCIENCES - 2001-2002

[BIOEE 456 Stream Ecology (also An introduction to the models used to Field and laboratory exercises designed to ENTOM 456 and NTRES 456) construct ecological theory and analyze data give firsthand experience with the ecology Spring. 4 credits. Limited to 60 students. on ecological dynamics, and to the mathemati­ and population biology of plants. Emphasis is Recommended: BIOEE 26l. S-U grades cal and computer methods used to analyze on making observations and measurements of optional. Field project with lab papers. these models. Applications from individual plants in the field and greenhouse, and on Lees, T R 9:05; lab, TWorR 1:25-4:25. decision-making through multispecies and data analysis.] Offered alternate years. Not offered 2001- spatial dynamics introduce the main themes in [BIOEE 466 Physiological Plant Ecology, 2002. B. L. Peckarsky. theoretical ecology: optimization, dynamics, Lectures Lecture: addresses the patterns and processes and the links between process and pattern. Spring. 3 credits. Limited to 45 students. occurring in stream ecosystems, including The lab includes instruction in computer Prerequisite: BIOEE 261 or introductory channel formation; water chemistry; watershed programming and review of mathematical plant physiology. S-U grades optional, with influences; plant, invertebrate, and fish methods.] permission of instructor. Lees, M W 8:40- community structure; nutrient.cycling; trophic dynamics, colonization, and succession; [BIOEE 462 Marine Ecology (also 9:55; optional disc TBA. Offered alternate years. Not offered 2001-2002. Staff. community dynamics; conservation; and the EAS 462) impacts of disturbances. Lab: field projects Spring. 3 credits. Limited to 75 students. A detailed survey of the physiological approaches used to understand the relation­ include descriptive and experimental Prerequisite: BIOEE 261. Lees and discs, ships between plants and their environment. techniques, hypotheses testing, and writing of M W F 10:10. Offered alternate years. Not Lectures explore: physiological adaptation; scientific papers related to environmental offered 2001-2002. C. D. Harvell. limiting factors; resource acquisition and assessment.] Lectures and discussion focus on current research in broad areas of marine ecology allocation; photosynthesis^ carbon, and energy [BIOEE 457 Limnology: Ecology of Lakes, with an emphasis on processes unique to balance; water use and water relations; Lectures marine systems. A synthetic treatment of nutrient relations; linking physiology, Fall. 3 credits. Prerequisite: BIOEE 26l or multiple levels of organization in marine development, and morphology; stress written permission of instructor. Recom­ systems including organismal, population, physiology; life history and physiology; the mended: introductory chemistry. Letter community, ecosystems, and evolutionary evolution of physiological performance; and grade, S-U by permission only. Lees, M W biology. Examples are drawn from all types of physiology at the population, community, and F 11:15. Offered alternate years. Not marine habitats, including polar seas, ecosystem levels. Readings draw from the offered 2001-2002. N. G. Hairston, Jr. temperate coastal waters, and tropical coral primary literature and textbooks.] Limnology is the study of inland fresh waters reefs.] and other, nonmarine, environments. This BIOEE 467 Seminar in the History of [BIOEE 463 Plant Ecology and Biology (also HIST 415, B&SOC 447, course focuses on lakes and ponds, which are and S&TS 447) discussed as distinct aquatic environments Population Biology, Lectures Summer (6-week session). 4 credits. with clear terrestrial boundaries, and within Fall. 3 credits. Prerequisite: BIOEE 261 or Limited to 18 students. S-U grades which ecological interactions are especially 278 or equivalents, or permission of optional: W. B. Provine. evident. In lakes, interactions between instructor. Recommended: some taxonomic Specific topic changes each year. organisms are often strong and adaptations familiarity with vascular plants and easily recognized. Physical and chemical concurrent enrollment in BIOEE 465. Lees, [BIOEE 468 Physiological Plant Ecology, properties of the environment impact M W F 11:15. Offered alternate years. Not Laboratory offered 2001-2002. M. A. Geber, organisms in important ways and organisms, Spring. 2 credits. Limited to 15 students. likewise, influence physics and chemistry. As P. L. Marks. Prerequisite: previous or concurrent a result, lakes provide excellent systems for This course examines the biological and enrollment in BIOEE 466. Lab, W 1:25— historical factors affecting the structure of understanding the links between physical 4:25, plus additional lab hours TBA. (thermal and mixing), chemical (dissolved plant communities, and the distribution, Offered alternate years. Not offered 2001- abundance, and population dynamics of elements and compounds), and organismal 2002. Staff. individual species. The influence of the dynamics. Lakes are exciting environments for A detailed survey of the physiological environment, disturbance history, competition, study in their own right, and for gaining approaches used in understanding the and herbivory on the organization of plant perspective on ecological and evolutionary relationships between plants and their processes in general.] communities are considered. Plant popula­ environment. Laboratories apply physiological tions are also studied through an analysis of techniques to specific ecological problems and [BIOEE 459 Limnology: Ecology of Lakes, plant life histories and plant-plant and plant- cover aspects of experimental design and Laboratory animal interactions. Throughout the course an computer-aided data analysis. Most laborato­ Fall. 2 credits. Prerequisite: concurrent or attempt is made to blend empirical patterns, ries run past the three-hour period, with previous enrollment in BIOEE 457. Letter experimental results, and theory. Readings are students spending an average of three hours/ grade, S-,U by permission only. Lab, drawn from the primary literature.] week in additional lab time for this course.] TWorR 1:25-4:25; 1 weekend field trip. Fee, $12 (for food on field trip). Offered BIOEE 464 Macroevolution [BIOEE 469 Food, Agriculture, and alternate years. Not offered 2001-2002. Spring. 4 credits. Limited to 25 students. Society (also B&SOC 469 and N. G. Hairston, Jr. and staff. Prerequisite: BIOEE 278 or permission of S&TS 469) Laboratories and field trips devoted to studies instructor. S-U grades optional, with Spring. 3 credits. Limited to 20 students. of the biological, chemical, and physical permission of instructor. Lees, T R 10:10- Prerequisite: an introductory ecology properties of lakes and other freshwater 11:25; disc, 1 hour each week TBA. course or permission of instructor. S-U environments. Exercises focus on understand­ Offered alternate years. A. R. McCune. grades optional, Lees, T R 1:25-2:40. Not ing the freshwater environment, on experi­ An advanced course in evolutionary biology offered 2001-2002. A. G. Power. mentation, and on understanding ecological centered on large-scale features of evolution. A multidisciplinary course dealing with the processes within lakes. Optional vertebrate Areas of emphasis include patterns and social and environmental impact of food dissection (fish) during one laboratory processes of speciation, phylogeny reconstruc­ production in the United States and develop­ exercise and during a portion of the weekend tion, the origin of variation, causes of major ing countries. Agroecosystems of various field trip.] evolutionary transitions, and patterns of kinds are analyzed from biological, economic, diversification and extinction in the fossil and social perspectives. The impacts of [BIOEE 460 Theoretical Ecology record. Discussion of these problems involve traditional, conventional, and alternative Spring. 4 credits. Enrollment limited. data and approaches from genetics, morphol­ agricultural technologies are critically Prerequisites: completion of the Biological ogy, systematics, paleobiology, development, examined in the context of developed and Sciences mathematics requirement or and ecology. developing economies. Specific topics include equivalent, and either one additional pest management, soil conservation, plant semester of mathematics, statistics or [BIOEE 465 Plant Ecology and Population Biology, Laboratory genetic resources, biotechnology, and modeling (e.g., ABEN 250, ABEN 367, sustainable development.] NTRES 305, NTRES 340, NTRES 410, Fall. 1 credit. Prerequisite: concurrent BIONB 422) or permission of instructor. enrollment in BIOEE 463. Lab, F 12:05- [BIOEE 471 Mammalogy S-U grades optional. Lees, T R 1:25-2:40; 5:00. Offered alternate years. Not offered Fall. 4 credits. Recommended: BIOEE 274. lab, M 2:00-4:25. Offered alternate years. 2001-2002. M. A. Geber, P. L. Marks. S-U grades optional, with permission of Not offered 2001-2002. S. P. Ellner. instructor. Carpooling to the Vertebrate ECOLOGY AND EVOLUTIONARY BIOLOGY 161

Collections at Cornell Business and BIOEE 475 Ornithology A survey of the major groups of organisms Technology Park is necessary several times Spring. 4 credits. Limited to 30 students. and their evolutionary histories. Intended to during the semester. Fee, $15. Lees, M T F Prerequisite: permission of instructor by fill out the biological backgrounds of earth 12:20; lab, MTorW 1:25-4:25; 1 weekend preregistering in E235 Corson. Recom­ and atmospheric science students concerning field trip required. Offered alternate years. mended: BIOEE 274. S-U grades optional, the nature and significance of the fossil record Not offered 2001-2002. Staff. with permission of instructor. Carpooling for their respective studies. Lectures on the evolution, classification, to the Vertebrate Collections at Research BIOEE 490 Topics in Marine Biology distribution, and adaptations of mammals. Park is necessary once a week. Fee, $15. Laboratory and fieldwork on systematics, Lees and labs, T R 12:20-4:25; occasional Spring. 2 credits. May be repeated for ecology, and natural history of mammals of field trips and special projects. Offered credit. Primarily for undergraduates. the world, with primary emphasis on the alternate years. D. W. Winkler. Limited to 15 students. Prerequisite: North American fauna. Systematics laborato­ Lectures cover various aspects of the biology permission of instaictor. S-U grades ries held in the museum at Research Park. of birds, including anatomy, physiology, optional. Lee, F 1:25-3:20. J. Morin and Live animals are studied in the field and are systematics, evolution, behavior, ecology, and M. Shulman. sometimes used in the laboratory for biogeography. Laboratory includes dissection Seminar courses on selected topics in marine nondestructive demonstrations or experi­ of dead material, studies of skeletons and biology; may include laboratory or field trips. ments. The systematics laboratory exercises plumages, and specimen identification of Topics and time of organizational meeting are are based on museum specimens.] avian families of the world and species of shown in departmental course offerings listed New York. Independent projects emphasize on the web site. [BIOEE472 Herpetology research skills. BIOEE 660 Field Studies in Ecology and Spring. 4 credits. Limited to 35 students. Systematics Recommended: BIOEE 274. S-U grades [BIOEE 476 Biology of Fishes Fall or spring. Variable credit. Prerequi­ optional, with permission of instructor. Fall. 4 credits. Limited to 24 students. sites: BIOEE 261, a taxon-oriented course, Fee, $30. Lees, T R 12:20; labs, T R 1:25— Recommended: BIOEE 274 or equivalent and permission of instructor. S-U grades 4:25; occasional field trips and special experience in vertebrate zoology. S-U optional, with permission of instructor. projects. Offered alternate years. Not grades optional, with permission of Lees and field trips TBA. Estimated costs: offered 2001-2002. H. W. Greene. instructor. A small lab fee may be required. TBA. Staff. Lectures cover various aspects of the biology of Lees, M W F 10:10; lab, M 1:25-4:25; with This course provides students with opportuni­ amphibians and reptiles, including evolution, additional lab time TBA; 2 field trips. ties to learn field techniques and new biotas zoogeography, ecology, behavior, and Offered alternate years. Not offered 2001- by participating in an intensive series of field physiology. laboratory topics include systemat­ 2002. A. R. McCune. exercises. Extended field trips may be ics, morphology, and behavior. Live animals are An introduction to the study of fishes: their scheduled during fall break, intersession, or studied in the field and are used in the structure, evolution, distribution, ecology, spring break. The regions visited, trip laboratory for nondestructive demonstrations physiology, behavior, classification, and objectives, and other details are announced by and experiments. The systematics laboratory identification, with emphasis on local species. the various instructors at an organizational exercises are based on museum specimens and Two field trips, including one full day meeting held at the beginning of the semester. dissection of preserved materials.] weekend trip required. Live animals are studied in the field and are sometimes used in Meetings on campus are devoted to orienta­ [BIOEE 473 Ecology of Agricultural the laboratory for nondestructive demonstra­ tion and reports on completed projects. Systems (also CSS 473) tions or experiments. The systematics and [Section 01: Life Histories of Marine and Fall. 3 credits. Limited to 45 students. dissection laboratories use preserved Freshwater Invertebrates Prerequisite: BIOEE 261 or permission of specimens.] Fall. 2 credits. Prerequisite: undergraduates instructor. S-U grades optional. During the must have previous experience or course BIOEE 477 Marine Invertebrates Seminar first 6 weeks of class, the Thursday work with marine or freshwater inverte­ meetings may run to 5:00 because of field Fall. 1 credit. Prerequisite: BIOEE 373 or brates. Two extended weekend field trips permission of instructor. Sem, 1 hour each trips. Lees and discs, T R 2:30-3:45. in either early September and October or Offered alternate years. Not offered 2001- week TBA. Offered alternate years. over winter break. Fee, TBA (to help cover 2002. A. G. Power, E. C. M. Fernandes. C. D. Harvell, J. G. Morin. transportation and housing). Offered Analysis of the ecological processes operating Discussions and directed readings center alternate years. Not offered 2001-2002. in agricultural systems, with an emphasis on arouhd current research themes in Inverte­ C. D. Harvell, N. G. Hairston, Jr. the interactions between organisms. Topics brate Biology. Designed as an on-campus Field trips to either the Shoals Marine include nutrient dynamics in agroecosystems, companion course to the field-based BIOEE Laboratory and Shackelton Point Field Station plant competition and facilitation, intercrop­ 373, Biology of the Marine Invertebrates. or to the Coast of Mexico. Students employ ping, the ecology of species invasions, Students will write individual research essays experimental approaches to study the mutualism in agroecosystems, plant-herbivore based on projects done in the field. evolution of invertebrate life histories.] relations, plant-pathogen interactions, biological pest control, and evolutionary [BIOEE 478 Ecosystem Biology Section 02: Graduate Field Course in processes in agriculture. Case studies from Spring. 4 credits. Prerequisite: BIOEE 261 Ecology both the tropics and the temperate zone are or equivalent. S-U grades optional. Lees Spring. 3 credits. Restricted to graduate used to illustrate important concepts.] and discs, T R 10:10-12:05. Offered students. A fee will be required to help alternate years. Not offered 2001-2002. cover food and lodging for trip to Florida. [BIOEE 474 Laboratory and Field L. O. Hedin. Offered alternate years. P. L. Marks, Methods in Human Biology (also Analysis of ecosystems in terms of energy R. B. Root. AHTHR 474) flow and nutrient cycles, emphasizing an The course is designed to give graduate Spring. 5 credits. Limited to 16 students. experimental approach and comparative students experience in defining questions and Prerequisite: one year of introductory aspects of terrestrial, freshwater, and marine designing field investigations. The class is biology or ANTHR 101 or permission of ecosystems. Consideration of anthropogenic based at the Archbold Biological Station in instructor. Recommended: permission of effects on ecosystems, such as from acid central Florida over spring break and during instructor by preregistering in E231 precipitation and offshore oil pollution. the following week. The class visits several Corson. Independent research project Analysis of climate change and regional ecosystems including sand pine scrub, cattle required. Lees and labs, T R 10:10-12:05; environmental change from an ecosystem ranches, cypress swamps, everglades, and additional hours TBA. Offered alternate perspective.] coral reefs. years. Not offered 2001-2002. K. A. R. Kennedy. BIOEE 479 Paleobiology (also EAS 479) BIOEE 661 Environmental Policy (also Practical exercises and demonstrations of Fall. 4 credits. Prerequisites: one year of ALS 661 and B&SOC 461) modern approaches to the methodology of introductory biology for majors and either Fall and spring. 3 credits each term. biological anthropology. Emphasis on BIOEE 274, EAS 375, BIOEE 373, or (Students must register for 6 credits each comparative human anatomy, osteology, permission of instructor. S-U grades term, since an “R” grade is given at the description of skeletal subjects, optional. Lees, M W F 12:20; lab W 2:00- end of the fall term.) Limited to 12 paleopathology, skeletal maturation, and 4:25. W. Allmon. students. Prerequisite: pennission of relevant field techniques for the archaeologist instructor. Sem, R 2:30-4:30. D. Pimentel. and forensic anthropologist.] 162 BIOLOGICAL SCIENCES - 2001-2002

This course focuses on complex environmen­ A survey of the historical background of Entomology (ENTOM 201-215, 331,453, 470-471) tal issues. Ten to twelve students, representing present-day concepts of human evolutionary Natural Resources (NTR ES 201-279-271,301- several disciplines, investigate significant variations and adaptations in space and time. 302, 305, 418-419) environmental problems. The research team The formation of biological anthropology as spends two semesters preparing a scientific an area of scientific inquiry within the social Plant Pathology (PLPA 309, 319) report for publication in S cien ce or and biological sciences is reviewed. Students BioScience. Thus far, every study has been select their own topics within a broad range Shoals Marine Laboratory (BIOSM 303-499) published. of readings in the history of Western concepts Statistics (BTRY 101-102, 261,301,451; ILRST of human origins, diversity, and place in 210-411) [BIOEE 665 Limnology Seminar nature.] Spring. 1 credit. May be repeated for credit. Primarily for graduate students; BIOEE 760 Special Topics in Evolution written permission of instructor required and Ecology for undergraduates. S-U grades optional. Fall or spring. 1-3 credits. May be repeated GENETICS AND DEVELOPMENT Sem TBA. Offered alternate years. Not for credit. Enrollment limited. S-U grades (BIOGD) offered 2001-2002. N. G. Hairston, Jr. optional, with permission of instructor. A seminar course on advanced topics in Hours TBA. Staff. [BIOGD 184 Understanding Genetics freshwater ecology.] Independent or group intensive study of Spring. 3 credits. May not be taken for special topics of current interest. Content credit after BIOGD 281 or 282. This course BIOEE 668 Principles of varies and is arranged between student and Biogeochemistry may be used toward the science distribu­ staff member. tion requirement of the College of Arts and Spring. 4 credits. Limited to 20 students. Sciences and the Group B distribution Prerequisite: solid background in ecology, BIOEE 763 Workshop in Biogeochemistry requirement of the College of Agriculture environmental chemistry, or related Fall or spring. 1 credit. May be repeated and Life Sciences. This course may not be environmental science. Permission of for credit. Limited to 15 students. used td fulfill the requirements for any instructor required for undergraduates. S-U Prerequisite: BIOEE 668. S-U grades only. program of study in the biological sciences grades optional. Lees and discs, T R 10:10— Workshop and disc, TBA. Staff. major. S-U grades optional. Lees, M W F 12:05. Offered alternate years. L. O. Hedin, Provides a workshop-forum in which graduate 2:30. Offered alternate years. Next offered staff. students interact with invited world-leaders in 2003. T. D. Fox. Lectures cover the biotic controls on the biogeochemistry. Workshop topics will An introduction to genetics for students chemistry of the environment and the change each semester. A one-week workshop majoring in fields other than biology. Genetics chemical control of ecosystem function. will be preceded by seven, one-hour is a rapidly developing science that is Emphasis is on cycles of major elements and preparatory discussions of readings. providing insight into all aspects of biology minor elements globally and in selected BIOEE 767 Current Topics in Ecology and practical tools which increasingly affect ecosystems, stressing the coupling of element and Evolutionary Biology our lives. This course shows how major cycles. A comparative approach is used to Fall. 4 credits. Prerequisite: permission of conclusions about inheritance have been illustrate similarities and differences in element instructor required for undergraduates. S-U derived from the experimental evidence, cycling among ecosystems. Analysis of both grades only. Lees and discs, T R 10:10- drawing on examples from the biology of theoretical and applied issues, including humans, other animals, plants, fungi, and global atmospheric changes and factors 12:05. Staff. Critical evaluation and discussion of theory bacteria. It also illustrates current and future controlling the acidification of lakes and soils. and research in ecology and evolutionary applications of genetic discoveries. For [BIOEE 669 Plant Ecology Seminar biology. Lectures by faculty and student-led example, the basic principles of inheritance, in Spring. 1 credit. May be repeated for discussions of topics in areas of current conjunction with methods for the isolation credit. Suggested for students majoring or importance. and detection of specific gene fragments, is used to understand the detection of genetic minoring in plant ecology. S-U grades BIOEE 899 M.S. Thesis Research optional. Sem TBA. Not offered 2001-2002. diseases and the identification of individuals Fall or spring. 1-15 credits. Prerequisite: Staff. (DNA fingerprinting). Other topics to be admission to the Field of Ecology and Includes review of current literature, student covered include the origin of mutations, use Evolutionary Biology. S-U grades optional. research, and selected topics of interest to of genetic methods to alter the properties of Hours TBA. E&EB field faculty. participants.] organisms and the influence of inheritance on Thesis research conducted by an M.S. student behavior.] BIOEE 670 Graduate Seminar in in the Field of Ecology and Evolutionary Vertebrate Biology Biology with advice and consultation of a BIOGD 281 Genetics Fall or spring. 1 credit. May be repeated major professor who is a member of the Field. Fall, spring, or summer (8-week session). 5 credits. Not open to freshmen in fall for credit. Primarily for graduate students; BIOEE 999 Ph.D. Dissertation Research written permission of instructor required semester. Enrollment may be limited to for undergraduates. S-U grades only. Sem Fall or spring. 1-15 credits. Prerequisite: 200 students. Prerequisite: one year of admission to the Field of Ecology and TBA. Staff. introductory biology or equivalent. No Seminar presentations and discussions by Evolutionary Biology as a Ph.D. student. admittance after first week of classes. students on areas of current research in S-U grades optional. Hours TBA. E&EB Students do not choose lab sections during vertebrate biology. Topics vary from semester field faculty. course enrollment; lab assignments are Dissertation research conducted by a Ph.D. to semester. made during first day of classes. Lees, T R student in the Field of Ecology and Evolution­ 10:10-12:05; lab, T or W or R or F 2:30- Section 01: Mating Systems, Parentage, ary Biology with advice and consultation of a 4:25. Problem-solving sessions strongly and Sexual Selection in Plants and major professor who is a member of the Field. recommended, T or W 8:30-9:45 (addi­ Animals tional session by arrangement). T. D. Fox, Fall. 1 credit. Primarily for graduate Related Courses in Other Departments M. L. Goldberg, R. J. MacIntyre. students; written permission of instructor A general study of the fundamental principles required for undergraduates. S-U grades Agricultural and Biological Engineering (ABEN 371, 456) of genetics in eukaryotes and prokaryotes. only. Sem TBA. K. R. Zamudio, Discussions of gene transmission, gene action M. A. Geber. Anthropology (ANTHR 375) and interaction, gene linkage and recombina­ tion, gene structure, gene and chromosome [BIOEE 673 Human Evolution: Concepts, Biological Sciences (BIO G 400, 498, 499; BIOGD History, and Theory (also ANTHR 387, 481,484; BIOMI 290, 414, 418, BIONB, mutations, genes in populations, and 673) 221,422, 427; BIOPL 241,248, 447, 448) extrachromosomal inheritance. Aspects of Fall. 3 credits. Prerequisite: one year of recombinant DNA technology are discussed. introductory biology or ANTHR 101 or Biology and Society (S&TS 206, 427) In the laboratory, students perform experi­ permission of instructor. Lee, M 2:30; sem ments with microorganisms and conduct an Crop and Soil Sciences (CSS 260, 366, p.m. independent study of inheritance in D roso­ and disc, W 7:30-9:30 Offered 415-416, 420) alternate years. Not offered 2001-2002. p h ila . K. A. R. Kennedy. Earth and Atmospheric Sciences (EAS 101-102, 204, 302, 321) GENETICS AND DEVELOPMENT 163

BIOGD 282 Human Genetics BIOGD 481 Population Genetics BIOGD 484 Molecular Evolution Spring. 2 or 3 credits (2 credits if taken Fall. 4 credits. Prerequisite: BIOGD 281, Spring. 3 credits. Prerequisites: BIOGD 281 after BIOGD 281). Each discussion limited BIOEE 278, or equivalents. Lees, M W F and organic chemistry. Lees, M W 8:40- to 25 students. Prerequisite: 1 year of 10:10; disc, M 2:30 or T 1:25. C. F. Aquadro. 9:55. Offered alternate years. R. J. MacIntyre. introductory biology or equivalent; Population genetics is the study of the An analysis of evolutionary changes in genes permission of instructor required for transmission of genetic variation through time and their protein products. Theories on the students who have taken BIOGD 281. S-U and space. The class explores how to quantify evolution of the genetic code, the construction grades optional. Lees, M W 10:10 (Lees, this variation, what the distribution of of phylogenetic trees from biochemical data also F 10:10 first 3 weeks only); disc, R variation tells us about the structure of natural and the role of gene duplications in evolution 10:10 or F 10:10 or 11:15. Staff. populations, and about the processes that lead are discussed. The second half of the course A course designed for nonmajors. Lecturers to evolution. Topics include the diversity and concerns the evolution and the organization provide the technical background needed to measurement of genetic variation, mating and of genomes from viruses to higher eukaryotes, understand controversial personal, social, and reproductive systems, selection and fitness, including the evolution of satellite DNA legal implications of modem genetics that are genetic drift, migration and population sequences and transposable elements. discussed in section meetings. structure, mutation, multilocus models, the genetics of speciation, quantitative traits, and • [BIOGD 485 Bacterial Genetics (also BIOMI 485) BIOGD 385 Developmental Biology the maintenance of molecular variation. Fall. 2 credits. Graduate students, see Fall. 3 credits. Prerequisite: BIOGD 281. Emphasis is placed on DNA sequence BIOMI 685. Prerequisite: BIOGD 281. Lees, M W F 11:15. K .J. Kemphues. variation, and the interplay between theory Recommended: BIOMI 290 and BIOBM An introduction to the morphogenetic, and the data from experiments and natural cellular, and genetic aspects of the develop­ populations. Specific case studies include the 330 or 331 and 332 or 333. Lees, W 7:30- 9:25 p.m. Next offered fall 2002. Staff. mental biology of animals. population genetic issues involved in DNA For course description, see BIOMI 485 ] BIOGD 387 Developmental Aspects of fingerprinting, the genetic structure and Evolution evolution of human populations, and the [BIOGD 486 Advanced Eukaryotic Fall. 2 credits. Prerequisite: BIOGD 281. study of adaptation at the molecular level. Genetics S-U grades optional. Lees, T R TBA. Examples are drawn from studies of animals, Spring. 4 credits. Enrollment may be A. W. Blackler. plants, and microbes. limited to 50 students. Prerequisites: An examination of the developmental BIOGD 482 Human Genetics and Society BIOGD 281, BIOBM 330 or 333 or 331 and 332. S-U grades optional. Lees, T 12:20- mechanisms that underlie evolutionary change Fall. 4 credits. Enrollment limited to 24 2:15 and R 12:20-1:10; disc R 1:25-2:15 or and organismal diversity and of the develop­ senior biological sciences majors, with F 11:15-12:05. Next offered spring 2003. mental constraints that contribute to evolution­ preference given to students studying E. E. Alani. ary conservatism. molecular biology and genetics. Prerequi­ The course develops fundamental skills in sites: BIOGD 281 and BIOBM 330 or 333 BIOGD 389 Embryology eukaryotic genetic analysis through lectures or 331 and 332, and permission of Spring. 3 credits. Preference given to and by reading, analyzing, and presenting instructor. S-U grades optional. Disc, T R seniors. Prerequisites: 1 year of introduc­ research articles. Concepts are presented 2:30-4:25. R. A. Calvo. tory biology and a knowledge of mamma­ within the context of a well-studied field, such Presentation of some of the science and lian adult anatomy. Lees, T R 10:10; labs, as chromosome segregation. The basic tools technology of human genetics, plus discussion T or R 2—4:25. A. W. Blackler. that have been developed to study this field of the ethical, social, and legal implications of A course in the embryonic development of are used to analyze other topics such as recent advances in the field. Among the topics vertebrate animals, with emphasis on the vegetative and meiotic cell cycle control, considered are assisted reproductive strategies, comparative aspects of morphogenesis and embryonic development, pathogen resistance eugenics, genetic counseling, genetic function at the tissue and organ levels. The in plants, and human genetics.] laboratory has a strong morphogenetic bias, screening (pre-implantation, prenatal, emphasizing the comparative aspects of neonatal, pre-symptomatic, carrier, and [BIOGD 600 Development of Sensory developmental anatomy and preparation for workplace), wrongful life and wrongful birth, Systems medical studies. genetic effects of abused substances, genetics Spring. 2 credits. Prerequisites:'introduc­ and behavior, human cloning, and therapy for tory biology, genetics, development, and BIOGD 394 Circadian Rhythms (also genetic diseases. Students lead many neurobiology, or permission of instructor. ENTOM 394 and BIONB 394) discussions. There is a major writing compo­ S-U or letter grades. Lee, M 7:00-8:40 p.m. Fall. 2 credits. Prerequisite: ENTOM 212, or nent to the course. Offered alternate years. Next offered BIOGD 281, or BIONB 221 or 222, or spring 2003. K. Whitlock. permission of instructor. S-U grades [BIOGD 483 Advanced Developmental Biology This course will explore the unique and optional. Lee, W 7:30-9:10 p.m. Offered shared mechanisms used in sensory system Spring. 3 credits. Prerequisites: BIOGD 1 alternate years. J. Ewer. development of both vertebrates and 281; BIOBM 332 or 330 or 333; and For description, see ENTOM 394. invertebrates. The first class of the course will BIOGD 385 or permission of instructor. provide a general introduction to the BIOGD 450 Vertebrate Development Lees, T R 2:30-4:00. Offered alternate development of sensory systems in vertebrates Spring. 3 credits. Prerequisite: introductory years. Next offered spring 2003. and invertebrates. Following classes will biology. S-U and letter grades. Lees, T R M. F. Wolfner. involve the reading of current and classic 11:40-12:55. S. Ely. An advanced course in developmental papers in sensory system development. This course is designed to examine the biology, with emphasis on the molecular Students will choose a topic and articles from development of characteristics that make events underlying developmental processes. a list provided by the instructor. Students will vertebrates unique. The course will start with Simultaneously, a molecular biology course be responsible for an oral presentation and an introduction to recent evolutionary and that focuses on how development modulates short paper.] molecular approaches to understanding the and uses transcriptional, post-transcriptional, rise of vertebrate structures. The development translational, and post-translational regulation BIOGD 682 Fertilization and the Early of vertebrate structures, such as neural crest, of gene expression and cellular events such as Embryo specialized sense organs, and limbs, will be signal transduction and cell-cell communica­ Spring. 2 credits. Prerequisites: BIOGD examined in detail with emphasis on the tion. Numerous developmental systems are 281; BIOBM 332, 330 or 333; and BIOGD cellular and molecular events controlling their discussed, and analyzed in microorganisms, 385 or permission of instructor. Lee, R development. plants and, especially, animals including fruit 2:30-4:25. Offered alternate years. BIOGD 480 Seminar in Developmental flies, nematode worms, and vertebrates such M. F. Wolfner. Biology as mice, frogs, and humans. Course readings This course treats the earliest events in the include original research articles. Discussion Spring. 1 credit. May be repeated for formation of a new organism. The methods emphasizes specific experiments and credit. Limited to juniors and seniors. and findings of genetic, developmental, and approaches, and results and their interpreta­ Prerequisite: BIOGD 281. S-U grades only. molecular analyses are discussed. Readings in tion.] Seminar TBA. Staff. the recent literature and discussions focus on pre-gastrulation embryos from several animal species. Topics include fertilization (sperm/ egg binding, sperm entry into egg), pro- 164 BIOLOGICAL SCIENCES - 2G£3 1-2 m

nuclear fusion, egg activation, initiation and Each graduate student presents one seminar BIOMI 290 General Microbiology terminating the cleavage, division period, per year based on his or her thesis research. Lectures cytoplasmic determinants, and changes in The student then meets with the thesis Fall, spring, or summer (6-week session). nuclear and cytoplasmic architecture. committee members for an evaluation of the 2 or 3 credits (2 credits if taken after presentation. BIOMI 192). Prerequisites: 1 year of [BIOGD 684 Advanced Topics in introductory biology for majors and 1 year Population Genetics BIOGD 787 Seminar in Genetics and of college chemistry, or equivalent. Spring. 2 credits. Limited to 20 students. Development Recommended: concurrent registration in Prerequisites: BIOGD 481 or equivalent Fall and spring. 1 credit. Limited to BIOMI 291. Lees, M W F 11:15. Staff. and written permission of instructor. S-U graduate students in Genetics and A comprehensive overview of the biology of grades optional. Lee, T 2:30-4:25. Offered Development. S-U grades only. Sem, M microorganisms, with emphasis on bacteria. alternate years. Not offered 2001-2002. 4-5:00. Staff. Topics include microbial cell structure and Next offered spring 2003. C. F. Aquadro. Seminars in current research in genetics and function, physiology, metabolism, genetics, An in-depth exploration of current areas of developmental biology conducted by diversity, and ecology. Applied aspects of research in population genetics. Readings distinguished visitors and staff. microbiology are also covered such as primarily from recent books and the current biotechnology, the role of microorganisms in literature. Specific topics are announced the Related Courses in Other Departments environmental processes, and medical previous fall and in the division’s catalog Advanced Plant Genetics (Plant Breeding 606) microbiology. supplement. Format includes lectures, discussion, and presentations by students.] Animal Development (Veterinary Anatomy 507) BIOMI 291 General Microbiology Biosynthesis of Macromolecules (Biological Laboratory [BIOGD 685 Advanced Bacterial Sciences [BIOBM] 633) Fall or spring, 2 credits. Summer (6-week Genetics (BIOMI 485) session), 2 credits. Prerequisite: concurrent Fall. 2 credits. Limited to graduate students Current Topics in Biochemistry (Biological Sciences or previous enrollment in BIOMI 290. Lee, [BIOBM] 731-736) in Biological Sciences; see BIOMI 485. F 12:20; labs, M W 12:20-2:15 or 2:30-4:25, Prerequisites: BIOGD 281 or equivalent, Evolutionary Biology (Biological Sciences or T R 10:10-12:05, 12:20-2:15, or 2:30- BIOBM 330 or 331 and 332 or equivalent, [BIOEE] 278) 4:25. C. M. Rehkugler. and permission of instructor. Recom­ Laboratory in Molecular Biology and Genetic A study of the basic principles and techniques mended: BIOMI 290 or equivalent. Lee, W Engineering of Plants (Biological Sciences of laboratory practice in microbiology, and 7:30-9:25; disc, R 10:10-11:00. Not-offered [BIOPL] 347) fundamentals necessary for further work in 2001-2002. Next offered fall 2002. Staff. the subject. For course description, see BIOMI 685.1 Laboratory in Plant Molecular Biology (Biological Sciences [BIOPL] 641) BIOMI 292 General Microbiology [BIOGD 687 Developmental Genetics Molecular Biology and Genetic Engineering of Discussion Fall. 2 credits. Limited to 20 students. Plants (Biological Sciences [BIOPL] 343) Spring. 1 credit. Prerequisite: concurrent or Prerequisites: BIOGD 281 and 385 or their previous enrollment in BIOMI 290. S-U equivalents. S-U grades optional. Lee TBA. Plant Cytogenetics (Plant Breeding 446) grades only. Disc TBA. C. M. Rehkugler. Offered alternate years. Not offered 2001- Plant Genome Organization (PLBR 653-03) A series of discussion groups in specialized 2002. K. J. Kemphues. areas of microbiology to complement BIOMI Plant Growth and Development (Biological Sciences Selected topics focus on the use of genetic 290. analysis in understanding mechanisms of [BIOPL] 644) development. Topics are drawn primarily from Plant Molecular Biology I (Biological Sciences BIOMI 331 General Parasitology (also studies in fruitflies, nematodes, mice and fish. [BIOPL] 653) VETMI 331) Possible topics include pattern formation, cell Spring. 2 credits. Prerequisites: one year of Plant Molecular Biology II (Biological Sciences introductory biology. Lees, T R 3:35-4:25. lineage, neural development, maternal [BIOPL] 652) information in development, germ cell D. Bowman. development, sex determination, and Protein-Nucleic Acid Interactions (Biological An introduction to the basic animal parasites, intercellular communication. Students read Sciences [BIOMI] 692) stressing systematics, taxonomy, general current literature and are encouraged to The Nucleus (Biological Sciences [BIOBM] 639) biology, ecological interactions, and behavior discuss each topic in class.] of non-medically important groups. The Undergraduate Research in Biology (Biological course will introduce the major animal BIOGD 780 Current Topics in Genetics Sciences [BIO G] 499) parasites, protozoan, nematode, platyhelm- Fall or spring. 2 credits. May be repeated Molecular Neurobiology BIONB 420/720 (also inth, acanthocephalan, annelid, and arthro­ for credit. Primarily for graduate students, BIOBM 435/735) pod. with preference given to majors in the Field of Genetics, written permission of BIOMI 391 Advanced Microbiology instructor required for undergraduates. Laboratory Fall. 3 credits. Prerequisites: BIOMI 290, Limited to 20 students. No auditors. S-U MICROBIOLOGY (BIOMI) grades optional, with permission of 291, and BIOBM 330 or 331 or 333. instructor. Seminar TBA. Staff. BIOMI 192 Microorganisms on the Planet Preference given to biological sciences Earth students in the microbiology program of BIOGD 781 Problems in Genetics and study. Lab, M W 1:25-4:25; disc, F 1:25. Development Summer. 3 credits. May not be taken for credit after BIOMI 290. S-U grades J. B. Russell, W. C. Ghiorse, Fall. 2 credits. Limited to first-year graduate optional. Lees, M-F 10:00-11:15. J. P. Shapleigh, S. H. Zinder! students in the Field of Genetics and R. P. Mortlock. A laboratory course that illustrates basic Development. Disc TBA. Staff. A course in microbiology designed to principles of experimental microbiology. The An introduction to the research literature in introduce students, who have a limited course is organized into four modules which selected areas through weekly problem sets background in science, to the microorganisms last three weeks each: (1) ecology, (2) and discussions. that populate our planet earth. Among the physiology, (3) genetics, and (4) structure and BIOGD 782-783 Current Genetics/ microorganisms studied are the bacteria, the function. Students are encouraged to take this Development Topics archaebacteria, some of the single-celled course during their third year of study. Spring. 1/2 or 1 credit for each topic. May plants and animals, and the viruses. Topics BIOMI 394 Applied and Food be repeated for credit. S-U grades only. covered are the basic nature of microorgan­ Microbiology (also FOOD 394) Lectures and seminars on specialized isms, their evolution on earth, their composi­ Fall. 2-3 credits. Prerequisites: BIOMI 290- topics to be announced. Staff. tion and growth, their role in the ecology of 291. M W F 12:20-1:10. C. A. Batt. this planet, their role in human history and Microorganisms play a central role in a variety BIOGD 786 Research Seminar in disease, and their use in bioengineering. This Genetics and Development of food, agricultural, and environmental course is not a prerequisite for advanced processes. This course will present a Fall and spring. 1 credit. Limited to and courses in microbiology. required of second-, third-, and fourth-year comprehensive survey of the roles that microorganisms play in industrial/biotechno- graduate students in Genetics and logical processes as well as their importance Development. S-U grades only. Sem, W in the safety and production of foods. Issues 12:20-1:30. Staff. MICROBIOLOGY 165

related to the biochemistry, genetics, and [BIOMI 414 Bacterial Diversity mutants; genetic exchange, recombination and physiology of microorganisms important in Spring. 3 credits. Prerequisites: BIOMI 290, mapping; complementation, epistasis and these processes will be reviewed. A tworcredit and 291, BIOBM 330 or 331 or 333 suppression; transposons; gene expression core section on food microbiology is recommended. Lees, M W F 11:15. Offered and regulation; and genetics of bacterial complemented by a one-credit section on alternate odd years. Next offered spring pathogenesis.] industrial/biotechnology applications. 2003. S. H. Zinder. A consideration of the evolutionary biology, BIOMI 610 Introduction to Chemical and BIOMI 397 Environmental Microbiology physiology, ecology, genetics, and practical Environmental Toxicology (also TOX (also CSS 398) potential of important groups of bacteria. 610) Fall. 3 credits. Prerequisites: BIOES 261 or Topics include molecular methods for Fall. 3 credits. Prerequisite: graduate BIOMI 290 or CSS (SCAS) 260 or permis­ determining bacterial phytogeny, the evolution standing in the field or consent of the sion of instructor. Lees, M W F 10:10. of diverse mechanisms of energy conserva­ instructor. Letter grades. Lee, M W F 11:15— W. C. Ghiorse, E. L. Madsen. tion, fixation of carbon and nitrogen, and 12:05. A. Hay. The biological properties, evolution, and adaptation to extreme environments.] Introduction to the general principles of behavior of microorganisms in natural systems toxicology including the sources, mechanisms, are discussed in relation to past and present BIOMI 416 Bacterial Physiology and targets of toxic agents. Special attention environmental conditions on Earth and other Spring. 3 credits. Prerequisites: BIOMI 290, will be given to the interaction between toxic living planets. The functional role of microor­ 291, and BIOBM 330 or 331 or 333, or agents and biological systems at both the ganisms in ecologically and environmentally their equivalents. Lees, M W F 11:00. organismal and ecological level. The effects of significant processes is also considered through Offered alternate even years. both anthropogenic and natural toxins will be discussion of specific topics such as nutrient J. P. Shapleigh. examined with respect to genetic and and toxic elemental cycles, transformation of The focus of the course is on physiological developmental toxicity as well as carcinogen­ pollutant chemicals, wastewater treatment, and metabolic functions of bacteria. Consider­ esis and specific organ toxicity. environmental biotechnology, and astrobiology. ation is given to chemical structure, regulation, BIOMI 652 (Section 02) Molecular Plant- growth, and energy metabolism. Special Microbe Interactions (BIOPL 652, [BIOMI 404 Pathogenic Bacteriology and attention is given to those aspects of bacterial Mycology (also VETMI 404) Sec 02) metabolism not normally studied in biochem­ Spring. 1 credit. Prerequisites: BIOGD 281, Spring. 2 or 3 credits (3 credits with lecture istry courses. and seminar with permission of instructor BIOBM 330 or 331 or 333, and BIOPL 653 for undergraduates). Prerequisites: BIOMI BIOMI 417 Medical Parasitology (also (section 01) or their equivalents. S-U 290 and 291 ■ Strongly recommended: BIO VETMI 431) grades optional. Lees, M W F 12:20 (12 G 305. Lees, M W 10:10; sem, F 10:10. Fall. 2 credits. Prerequisites: courses lees) Jan. 21-Feb. 15. S. C. Winans. Offered alternate odd years. Not offered pertaining to zoology and biology. Lees, For course description, see BIOPL 652, Sec 02. 2001-2002. D. Debbie. T R 3:35-4:25. D. Bowman. BIOMI 690 Prokaryotic Biology This is a course in medical microbiology, A systematic study of anthropod, protozoan, Fall and spring. 4 weeks/8 lectures. presenting the major groups of bacterial and and helminth parasites of public health 1 credit/section to be offered. T R. Time mycotic pathogens important to human and importance with emphasis on epidemiologic, TBA. veterinary medicine. The emphasis of this clinical, and zoonotic aspects of these course is infection and disease pathogenesis. parasitisms. Section 1—Microbial Structure and Topics include disease causality; interactions Function of host, pathogen and environment, including BIOMI 418 Microbial Ecology Fall. J. P. Shapleigh. immunity to bacteria and fungi; and principles Spring. 3 credits. Prerequisites: BIOMI 290 Discussion of those macromolecules and of antimicrobial therapy and drug resistance. and 291, or BIOMI 398 and instructor’s assemblages of macromolecules that together A companion seminar addresses the current permission, and BIOBM 330 or 331 and define the structure of the prokaryotic cell. and classic literature related to microbial 332. Lees, M W F 10:10-11. E. R. Angert. This will include external structures, such as pathophysiology on the cellular and molecular Understanding the role of microorganisms in cell wall, flagella, pili, and peptidoglycan and level.] natural environments is one of the greatest internal structures such as specialized vesicles challenges facing microbiologists. This course and other large complexes. BIOMI 408 Viruses and Disease I (also will introduce current biochemical and VETMI 408) macrom'olecule sequence-based methods to Section 2—Microbial Genetics Spring. 2 credits. Prerequisites: BIOMI 290, assess community diversity and microbial Fall. J. D. Helmann. 291; BIO G 305; and permission of activity in a variety of ecosystems. Other Reviews the fundamental concepts of instructor. Recommended: BIOGD 281. topics discussed include bacterial growth and microbial genetics including mutations and Lees, M W 7:30 p.m. Offered alternate even survival, population biology, and microbial their analysis, plasmids, conjugation, years. J. Casey. interactions. transformation, transduction, transposition, The course covers basic concepts in virology recombination, repair, and mutagenesis. [BIOMI 420 Microbial Genomics with emphasis on virus-host interactions, Section 3—Microbial Physiology/Diversity strategies for gene regulation, and mecha­ Spring. 2 credits. Prerequisites: BIOMI 290, Fall. S. H. Zinder. nisms of pathogenicity. Selected viral BIO G 281, BIOBM 330, or equivalent. The major energy conserving modes of infections that result in immune dysfunction Lees, T R 10:10-11:00. Offered alternate metabolism and their phylogenetic distribu­ and neoplasis are highlighted in the context of odd years. Not offered 2001-2002. tions among both bacteria and archaea are approaches to prevent or reduce the severity J. P. Shapleigh and J. D. Helmann. reviewed. Topics include fermentation, of diseases. Genomic information is revolutionizing biology. We will discuss the impact of respiration, photosynthesis, and pathways of [BIOMI 409 Viruses and Disease II (also genomic information on the study of microbial carbon and nitrogen fixation. VETMI 409) physiology, evolution, and biotechnology. Section 4—Microbial Pathogenesis Fall. 2 credits. Prerequisites: BIOMI 290 Topics will include both techniques (auto­ Spring. S. C. Winans. and 291. Recommended prerequisites: mated DNA sequencing, assembly, annotation, An introduction to the fundamental concepts BIOMI 408, BIOBM 330-332, BIOBM 432. DNA chips) and applications (genome-wide of bacterial pathogenesis including the normal Lees, T R 2:30-3:20 p.m. Offered alternate analysis of transcription, functional flora, pathogen efltry and colonization, the even years. G. Whittaker. genomics).] production and regulation of toxins, horizontal This course will be complementary to BIOMI [BIOMI 485 Bacterial Genetics transfer of pathogenesis determinants, and the 408, Viruses and Disease I, and will empha­ roles of both specific and nonspecific host Fall. 2 credits. Graduate students, see size RNA viruses. The course will be complete defenses. Examples will include bacterial BIOMI 685. Prerequisite: BIOGD 281. in its own right. As such, completion of pathogens of both animals and plants. BIOMI 408 is not a requirement, but is Recommended: BIOMI 290 and BIOBM encouraged. The structure and classification of 330 or 331 and 332 or 333. Lees, W 7:30- Section 5—Environmental Microbiology viruses, virus entry, genome replication and 9:25 p.m. Not offered 2001-2002. Staff. Spring. E. L. Madsen. assembly will be studied with particular Concepts and principles of formal genetic A core course of concepts, methods, and emphasis on virus-host cell interactions. analysis as applied to prokaryotes, with current literature that reveals the Vaccination, chemotherapy and evolution of emphasis on enterobacteria and their viruses. multidisciplinary nature of environmental viruses will also be discussed.] Topics include mutagenesis and isolation of microbiology and its relationship to prokary- 166 BIOLOGICAL SCIENCES - 2001-2002

otic biology. The crucial roles that microor­ Introduction to Scanning Electron Microscopy cells, synaptic mechanisms, neurochemistry, ganisms play in catalyzing biogeochemical (Biological Sciences [BIO G] 401) motor systems, sensory systems, learning, and reactions throughout the biosphere will be Introductory Mycology (Plant Pathology 309) memory. Some discussion sections include discussed. dissections of preserved brains. Light and Video Microscopy for Biologists BIOMI 791 Advanced Topics in (Biological Sciences [BIO G] 450) [BIONB 322 Hormones and Behavior (also PSYCH 322) Microbiology Limnology: Ecology of Lakes, Lectures (Biological Fall or spring. 1 credit. May be repeated Sciences [BIOEE] 457) Fall. 3 credits; 2 lectures plus a section in for credit. Prerequisite: graduate standing which students will read and discuss in microbiology. S-U grades only. Disc Magical Mushrooms, Mischievous Molds (Plant original papers in the field, give an oral TBA. S. C. Winans, E. R. Angert. Pathology 201) presentation, and write a term paper. Reading and presentation by graduate Microbiology for Environmental Engineering (Civil Limited to juniors and seniors. Prerequisite students of current literature in selected areas and Environmental Engineering 451) (1 of the following): PSYCH 223, or of modern microbiology. BIONB 221, or BIONB 222, or one year of Plant Virology (Plant Pathology 645) introductory biology plus a course in BIOMI 795-796 Current Topics in Principles of Biogeochemistry (Biological Sciences psychology. Letter grades only. Lee M W F Microbiology [BIOEE] 668) 11:15. Not offered 2001-2002. E. Adkins Fall, 795; spring, 796. 1/2 or 1 credit for The Soil Ecosystem (Crop and Soil Science 366) Regan. each topic. May be repeated for credit. See PSYCH 322 for description.] Designed primarily for graduate students in microbiology. Prerequisite: upper-level BIONB 324 Biopsychology Laboratory courses in microbiology. S-U grades only. (also PSYCH 324) . Lees TBA. Staff. NEUR0BI0L0GY AND BEHAVIOR Fall. 4 credits. Limited to 20 upperclass Lectures and seminars on special topics in (BI0NB) students. Prerequisites: laboratory microbiology. experience in biology or psychology, BIONB 111 Brain Mind and Behavior BIONB 221 and 222 or PSYCH 123 and BIOMI 797 Scientific Communication (also PSYCH 111 and COGST 111) 222; and permission of instructor. Labs, T Skills Spring. 3 credits. Intended for freshmen R 1:25-4:25. T. DeVoogd. Fall and spring. 1 credit each semester. and sophomores in the humanities and See PSYCH 324 for description. F 2:30-3:20. social sciences; junior and senior non­ The ability to communicate effectively is BIONB 325 Neurodiseases— Molecular biology majors by permission only. Letter essential for success as a scientist. The primary Aspects grades only. Lees, M W F 9:05- E. Adkins goal of this course is to provide students with Fall. 3 credits. Prerequisites: two courses Regan and R. R. Hoy. an opportunity to develop self-confidence and from BIONB 222, BIOGD 281, BIOBM 330, See COGST 111 for description. refine their formal oral presentation skills. or 331; co-registration in one of the two is Students will be asked to present topical BIONB 221 Neurobiology and Behavior I: acceptable. S-U grades optional. Lees, T R seminars that will be critically evaluated by Introduction to Behavior 9:05; disc, T 1:25, 2:30, or 3:35. the instructor. Feedback for improving the Fall. 3, 4, or 5 credits (4 credits with one T. R. Podleski. presentation and peer evaluations will be discussion per week; 5 credits with two The intent of this course is to teach students emphasized. Taken by students in the discussions per week and participation in how to use recombinant DNA techniques for Graduate Field of Microbiology during their the Writing in the Majors program). 4- or the study of neurodiseases. How are genes first two semesters, a third semester is 5-credit option required of students in the responsible for diseases identified and how optional. neurobiology and behavior program of are the functions of these genes studied? study. Each 4-credit discussion section is Attention is focused on those neural diseases BIOMI 798 Graduate Research Seminar limited to 20 students, with preference in which significant advances have been made in Microbiology given to students studying neurobiology using these techniques, for example, Fall and spring. 1 credit each semester. and behavior. Enrollment in the 5-credit Alzheimer’s, Huntington’s, Prion diseases, Required of all graduate students in the option is limited to 12 students. Students schizophrenia, depression, disorders affecting Graduate Field of Microbiology. S-U grades may not preregister for the 5-credit option; ion channels, and muscular dystrophies. only. F 1:25-2:15. Staff. interested students complete an applica­ Emphasis is placed on how these studies All graduate students in the Field of Microbiol­ tion form on the first day of class. Not provide a useful approach to studying the ogy are required to attend BIOMI 798 and are open to freshmen. Prerequisite: 1 year of mammalian nervous system by exposing the required to present a seminar concerning their introductory biology for majors. May be functions of genes that would be difficult to research at least once each year. taken independently of BIONB 222. S-U identify in other ways. BIOMI 799 Microbiology Seminar grades optional. Lees, M W F 12:20; disc BIONB 326 The Visual System TBA. P. W. Sherman and staff. Fall and spring. Required of all graduate Spring. 4 credits. Prerequisite: BIONB 222 A general introduction to the field of behavior. students in the Graduate Field of Microbi­ or BIOAP 3H, or permission of instructor. ology and open to all who are interested. Topics include evolution and behavior, behavioral ecology, sociobiology, chemical S-U grades optional. Lees, M W F 10:10; Sem TBA. Staff. disc, 1 hour each week TBA. Offered ecology, communication, rhythmicity, alternate years. H. C. Howland. orientation and navigation, and hormonal Related Courses in Other Departments The visual systems of vertebrates are mechanisms of behavior. Advanced Food Microbiology (Food Science 607) discussed in breadth and depth as well as BIONB 222 Neurobiology and Behavior II: some aspects of invertebrate vision. Topics Advanced Immunology Lectures (Biological Introduction to Neurobiology covered include the optics and anatomy of Sciences [BIO G] 705 and Veterinary Spring. 3 or 4 credits (4 credits with eyes, retinal neurophysiology, structure and Microbiology 705) discussion and written projects). 4-credit function of higher visual centers, ocular Advanced Work in Bacteriology, Virology, or option required of students studying motility and ocular and visual system Immunology (Veterinary Microbiology 707) neurobiology and behavior. Each development. discussion limited to 20 students, with Bacterial Plant Diseases (Plant Pathology 647) BIONB 327 Evolutionary Perspectives on preference given to students studying Human Behavior Basic Immunology, Lectures (Biological Sciences neurobiology and behavior. Not open to [BIO G] 305 and Veterinary Microbiology 315) Fall. 3 credits. Prerequisites: BIONB 221 freshmen. Prerequisites: 1 year of and permission of instructor required. Ecology of Soil-Borne Pathogens (Plant introductory biology for majors and 1 year Letter grades only. T R 2:55-4:10. Pathology 644) of chemistry. May be taken independently of BIONB 221. S-U grades optional. Lees, S. T. Emlen. Food Microbiology, Laboratory (Food Science 395) A Socratically taught, discussion-based course M W F 12:20; disc TBA. A. H. Bass arid dealing with evolutionary perspectives on Food Microbiology, Lectures (Food Science 394) staff. human behavior. Each fall a different topic is A general introduction to the field of cellular Immunology of Infectious Diseases and Tumors selected for study. All class members read and (Biological Sciences [BIO G] 706 and and integrative neurobiology. Topics include discuss primary papers and recent books. Veterinary Microbiology 719) neural systems, neuroanatomy, developmental Each student is responsible for leading neurobiology, electrical properties of nerve NEUROBIOLOGY AND BEHAVIOR 167 multiple discussions, for writing an original [BIONB 421 Effects of Aging on Sensory function in nerve cells and other excitable paper, and for peer-reviewing papers of other and Perceptual Systems (also cells (e.g., muscle, heart, glands). The latest class members. PSYCH 431 and 631) electrophysiological and molecular genetic Fall. 3 or 4 credits. The 4-credit option experiments will be reviewed. Diversity of BIONB 328 Biopsychology of Learning involves a term paper or creation of a electrophysiology deriving from channel and Memory (also PSYCH 332) relevant web site. Limited to 25 students. structure and expression patterns will be Spring. 3 credits. Prerequisites: 1 year of Prerequisites: an introductory course in considered in the contexts of behavior and biology and either a course in biopsychol­ biology or psychology, plus a second behavioral plasticity (learning), neural ogy or BIONB 222. S-U grades optional. course in perception or neurobiology or development, and channel evolution. Course Lees, M W F 11:15. T. DeVoogd. cognitive science or biopsychology. T R format includes written and oral presentations, See PSYCH 332 for description. 10:10-11:25. Not offered 2001-2002. reviewing scientific literature in selected areas, BIONB 329 Ecology of Animal Behavior B. P. Halpern. and proposing new experiments.] For description see PSYCH 431.] (also BIOSM 329) BIONB 426 Animal Communication Summer. 4 credits. Prerequisite: 1 year of BIONB 422 Modeling Behavioral Spring. 4 credits. Prerequisites: BIONB 221. introductory college biology. Recom­ Evolution Letter grade only. T R 2:55-4:10; disc, mended: course work in ecology, Spring. 4 credits. Limited to 25 students. 1 hour each week TBA. J. Bradbury, psychology, or behavior. S-U grades Prerequisites: BIONB 221, 1 year of S. Vehrencamp. optional. A special 2-week course offered calculus, 1 course in probability or An integrated approach to animal communica­ at Cornell’s Shoals Marine Laboratory statistics, and permission o f instructor tion, organized into three parts: 1) the physics (SML) on an island off Portsmouth, N.H. (Office: W309 Mudd Hall; phone: and physiology of producing, transmitting, For more details and an application, 254-4352). This course is open to and receiving signals; 2) optimal strategies for consult the SML office, G14 Stimson Hall. advanced undergraduates and graduate encoding information, using information to Daily lees, labs, and fieldwork for 2 weeks. students. S-U grades optional. Lees, T R make decisions, and designing signals; and 3) SML faculty. 2:30-4:00; computer lab, 1 class period per the behavioral ecology of signal evolution. See BIOSM 329 for description. week TBA. Offered alternate years. BIONB 427 Animal Social Behavior [BIONB 392 Drugs and the Brain H. K. Reeve. This is an intensive lecture and computer lab Spring. 4 credits. Limited to 30 students. Spring. 4 credits. Prerequisites: BIONB 222 course on modeling strategies and techniques Prerequisites: BIONB 221 and BIOEE 26l or equivalent course in neurobiology with or 278, and advance permission of permission of instructor. A knowledge of in the study of behavioral evolution. Popula­ tion-genetic (including quantitative-genetic), instructor. S-U grades optional, with biochemistry is useful but not required. static optimization, dynamic programming, permission of instructor. Lees and discs, S-U grades optional. T R 10:10—11:25; disc and game-theoretic methods are emphasized. T R 2:30-4:25. Offered alternate years. TBA. Offered alternate years. Not offered These approaches are illustrated by applica­ T. D. Seeley. 2001-2002. R. Harris-Warrick. tion to problems in optimal foraging, sexual An intensive course for upper-division An introduction to neuropharmacology, with selection, sex ratio evolution, animal students interested in behavorial ecology and an emphasis on the neural mechanisms of communication, and the evolution of sociobiology. Lectures, discussions, and psychoactive drugs. Topics include a brief cooperation and conflict within animal social student presentations examine topics introduction to neuropharmacology and a groups. Students learn to assess critically including adaptation, communication, mating discussion of the major neurotransmitter recent evolutionary theories of animal systems, sexual selection, sex ratios, inbreed­ families. The rest of the course will cover the behavior, as well as to develop their own ing and outbreeding, altruism, kin recognition, major psychoactive drugs, including cocaine, testable models for biological systems of and conflict and cooperation in animal heroin, psychedelics, marijuana and alcohol, interest or to extend pre-existing models in societies. as well as pharmaceuticals for the treatment of novel directions. The M athematica software anxiety, schizophrenia and depression. The [BIONB 428 Topics in Behavior program is used as a modeling tool in the course will include a term paper in the form Fall or spring. 2-4 credits. (Credits based accompanying computer lab (no prior of a grant proposal to study a current problem on number of lectures and/or field experience with computers required). in neuropharmacology.] exercises as outlined in the division’s catalog course supplement and subject to BIONB 394 Circadian Rhythms (also BIONB 424 Neuroethology (also PSYCH 424) approval through the associate director’s BIOGD 394 and ENTOM 394) Spring. 4 credits. Prerequisites: BIONB 221 office.) May be repeated for credit. Fall. 2 credits. Prerequisite: ENTOM 212, or or 222, or 1 year of introductory biology Primarily for undergraduates. S-U grades BIOGD 281, or BIONB 221 or 222, or for majors and permission of instructor. optional. Not offered 2001-2002. Staff. permission of instructor. S-U grades S-U grades optional. M W F 11:15; disc, Courses on selected topics in behavior; can optional. Lee, W 7:30-9:10 p .m. Offered 1 hour each week TBA. C. D. Hopkins. include lecture and seminar courses; may alternate years. J. Ewer. Neuroethologists take a comparative and include laboratory. Past topics have included For description, see ENTOM 394. evolutionary approach to study the nervous animal orientation, insect behavior, bio­ BIONB 396 Introduction to Sensory system. They ask, how do brains of animals rhythms, and communication. Topics, Systems (also PSYCH 396 and 696) compare and how did they come about instructors, and time of organizational meeting Spring. 4 credits. Limited to 25 students. through the process of evolution? How are are listed in the division’s catalog supplement Prerequisites: an introductory course in neural circuits adapted to species-typical issued at the beginning of each semester.] behavior? What is the hope and interest in the biology or biopsychology, plus a second [BIONB 429 Olfaction and Taste: course in behavior, biopsychology, study of a large diversity of animals, com­ Structure and Function (also cognitive science, neuroscience, or pared to a specialized look at just a few PSYCH 429) mammalian species? Can we hope to perception. Students are expected to have Spring. 3 or 4 credits (4 credits with term understand how animals with specialized knowledge of elementary physics, paper or research project, which can, but chemistry, and behavior. S-U grades behaviors have specialized nervous systems? need not, study nonhuman vertebrates). What is the sensory world of a real animal optional. Lees, M W F 10:10. Offered Preference given to junior and senior and how does it vary from species to species? alternate years. B. P. Halpern. psychology and biology majors and These and other questions will derive this See PSYCH 396 for description. graduate students. Graduate students, see introductory survey of neuroethology PSYCH 629. Prerequisite: a 300-level BIONB 420/720 Topics in Neurobiology including: exotic senses; amazing motor course in biopsychology or equivalent. and Behavior programs; surprising integration. Fall or spring. Variable credit. May be Lees, T R 9:05. Offered alternate years. Not repeated for credit. Primarily for under­ [BIONB 425 Molecular Neurophysiology offered 2001-2002. B. P. Halpern. graduates. S-U grades optional. TBA. Staff. Spring. 3 credits. Prerequisite: BIONB 222 See PSYCH 429 for description.] Courses on selected topics in neurobiology or permission of instructor. S-U grades [BIONB 430 Experimental Molecular and behavior; can include lecture and seminar optional. Lees, T R 2:55-4:10. Offered Neurobiology (also BIOBM 443) courses. Topics, instructors, and time of alternate years. Not offered 2001-2002. Spring. 2 credits variable. Limited to 12 organizational meetings are listed in the D. McCobb. students. Prerequisites: co-meeting with catalog supplement issued at the beginning of Course focuses on ion channels, the primary BIOBM 430 lab. Mandatory registration via the semester. proteins generating cellular electrical signals 168 BIOLOGICAL SCIENCES - 2001-2002

web page: www.mbg.comell.edu/ [BIONB 492 Sensory Function (also properties of sound, physiological mecha­ courses.html. Letter grade only. Disc, 1 PSYCH 492 and 692) nisms for sound production and hearing, and hour each week on day other than lab Spring. 4 credits. Limited to 25 students. the behavioral contexts in which sounds are day; Lab T or R all day, or M and W Prerequisite: a 300-level course in used. Acoustic techniques are provided in the afternoons, to be coordinated with other biopsychology, or BIONB 222, or BIOAP laboratory where students learn how to BIOBM 430 sections. Offered alternate 311, or equivalent. Students are expected record, synthesize, and analyze sounds with years. Not offered 2001-2002. to have knowledge of elementary physics, the aid of recorders and Mac and/or PC D. L. Deitcher. chemistry, and behavior. S-U grades computers running customized software. Labs See BIOBM 443 for description.] optional. Lees, M W F 10:10. Offered are designed around the lecture material and alternate years. Not offered 2001-2002. provide practical “real-world” exercises BIONB 440 Electronics in Neurobiology FI. C. Howland, B. P. Halpem. designed to stimulate discovery of fundamen­ Fall. 4 credits. Limited to juniors, seniors, See PSYCH 492 for description.] tal principles described in lectures. Class and graduate students. Prerequisites: a [BIONB 493 Developmental Neurobiology research projects on a selected topic in calculus course. S-U grades optional. Lees, bioacoustics are required. Engineering Fall. 3 credits. Prerequisite: BIONB 222 or T R 8:40-9:55. Lab, W 1:25-4:25. Offered students with interests in music, audio permission of instructor. S-U grades alternate years. B. R. Land. analysis, and digital signal processing are optional, with permission of instructor. The course will emphasize understanding of encouraged. the electrical functioning of the nervous Lees, T R 2:55-4:10. Offered alternate system and will enable students to build years; not offered 2001-2002. R. Booker. BIONB 531 From Signals to Syntax (also instrumentation to study the nervous system. Lectures covering the development of the COGST 531, LING 531, PSYCH 531, It will be taught by mathematical analysis, nervous system, taking examples from both COM S 531) simulation, and construction of circuit vertebrates and invertebrates. Emphasis is on Spring. 4 credits. Prerequisites: COGST examples drawn from practical neurobiologi- cellular and molecular issues, that is, how do 101. S-U grades optional. M 4:15-6:15 p.m. cal instrumentation problems and the nerve cells differentiate both morphologically S. Edelman. electronic basis of neurons. and biochemically? The role of cues such as See COGST 531 for description. hormones and developmental genes in neural [BIONB 441 Computers in Neurobiology development is discussed. Readings are taken [BIONB 623 Chemical Communication (also CHEM 622) Fall. 4 credits. Limited to juniors, seniors, from original journal articles.] and graduate students. Prerequisites: a Fall. 3 credits. Primarily for research- calculus course. S-U grades optional. Lees, [BIONB 494 Comparative Vertebrate oriented students. Limited to 30 students. T R 8:40-9:55. Lab, W 1:25-4:25. Offered Neuroanatomy Prerequisites: 1 year of introductory alternate years. Not offered 2001-2002. Spring. 3 credits. Intended for juniors, biology for majors or equivalent, course B. R. Land. seniors, and graduate students. Prerequi­ work in biochemistry, and CHEM 358 or This course is an introduction to computer site: BIONB 222 or equivalent. S-U grades equivalent. S-U grades optional. Lees, M W instrumentation techniques and data reduc­ optional. Lees, T R 10:10-11:30. Offered 10:10; disc, F 10:10. Offered alternate tion. It will give a basic understanding of the alternate years. Not offered 2001-2002. years. Not offered 2001-2002. T. Eisner, techniques used for coupling a biological A. H. Bass. J. Meinwald, W. L. Roelofs, and guest experiment to a computer. It will include Organization and evolution of neuroanatomi- lecturers. techniques to convert raw data to scientific cal pathways as substrates for species-typical The production, transmission, and reception visualization. Some computer modeling vertebrate behaviors. The course is divided of chemical signals in communicative examples drawn from practical neurobiologi- into three major sections: development, interactions of animals, plants, and microor­ cal problems will be done.] general principles of brain organization, and ganisms. Studies of insects are emphasized. co-evolution of vertebrate brain and behavior.] Specific topics are treated with varying BIONB 470 Biophysical Methods (also emphasis on chemical, biochemical, ecologi­ A&EP 470 and VETPR 470) [BIONB 495 Molecular and Genetic cal, behavioral, and evolutionary principles.] Spring. 3 credits. Prerequisites: basic Approaches to Neuroscience knowledge of and interest in physics and Fall. 3 credits. Limited to juniors, seniors, BIONB 720 Seminar in Advanced Topics mathematics is expected, but strong efforts and graduate students. Prerequisites: in Neurobiology and Behavior are made to give an intuitive understand­ BIONB 222 and BIOBM 330 or 332. Letter Fall or spring. Variable credit. May be ing of the mathematics and physics grade only. Lees, T R 2:55-4:10. Offered repeated for credit. Primarily for graduate involved. Some knowledge of physical alternate years. Not offered 2001-2002. students; written permission of instructor chemistry, molecular and cell biology, or D. Deitcher. required for undergraduates. S-U grades neurobiology will be helpful. Depending Focus of the course is on how different optional. Sem TBA. Staff and students. on individual background, all students will molecular and genetic approaches have led to Designed to provide several study groups find certain aspects easy and other aspects major advances in neuroscience. Lectures, each semester on specialized topics. A group demanding. Letter grades only. Lees, T R student presentations, and discussions may meet for whatever period is judged 8:40-9:55. M. Lindau. examine original research articles. Topics adequate to enable coverage of the selected See A&EP 470 for description. include ligand-gated channels, potassium topics. Ordinarily, topics are selected and channels, seven membrane spanning circulated during the preceding semester. BIONB 491 Principles of receptors, development of the neuromuscular Discussion of current literature is encouraged. Neurophysiology junction, neurotransmitter release, second Suggestions for topics should be submitted by Spring. 4 credits. Limited to 20 students. messengers, and learning and memory.] faculty or students to the chair of the Prerequisite: BIONB 222 or written Department of Neurobiology and Behavior. permission of instructor. S-U grades for BIONB 496 Bioacoustic Signals in graduate students with permission of Animals and Man BIONB 721 Introductory Graduate instructor. Lees, M W 10:10; lab, M or T Fall. 3 credits. Limited to 12 junior, senior, Survey in Neurobiology and Behavior 12:20-4:25; additional hours TBA. and graduate students. Prerequisites: 1 Fall. 2 credits. Required of graduate B. R. Johnson. year of introductory biology, PHYS 101- students majoring in neurobiology and A laboratory-oriented course designed to 102 or 207-208, and permission of behavior. Concurrent registration in teach the theory and techniques of modem instructor. S-U grades optional. Lees, M W BIONB 221 and 222 not required. S-U cellular neurophysiology. Lecture time is used 9:05; lab TBA. Offered alternate years. grades only. Lees and discs, TBA. to introduce laboratory exercises and discuss C. W. Clark. C. D. Hopkins and staff. results, to supplement laboratory topics, and Humans and animals live in a world of sound. Lectures, readings, and discussion introduce for discussion of primary research papers. Mechanisms for sound production and first-year graduate students to the research Extracellular and intracellular recording and perception are extremely varied. Acoustic activities of the faculty in the Graduate Field voltage clamp techniques are used to analyze signals mediate social interactions, and are of Neurobiology and Behavior. Class meets motor neuron and sensory receptor firing used to scan the environment for food and to weekly for two hours. Students also prepare a properties, and examine the cellular basis for aid in navigation. For many species acoustic research proposal on a potential topic for their resting and action potentials and synaptic sensing plays a critical role in predator thesis research (in the format of an NSF or transmission. Invertebrate preparations are detection and avoidance. This course teaches NIH grant). This proposal will be prepared in used as model systems. Computer acquisition students about animal acoustic signaling by consultation with one or more relevant faculty and analysis of laboratory results are introducing them to various animal acoustic members. emphasized. systems. The course presents the physical PLANT BIOLOGY 169

[BIONB 723 Advanced Topics in Animal [BIONB 792 Advanced Laboratory in O TS Undergraduate Semester Abroad Programs Behavior Cellular and Molecular Neurobiology Shoals Marine Laboratory Program Fall or spring. Variable credit. May be Fall or spring. 2 credits. May be repeated repeated for credit. Primarily for graduate for credit. Primarily for graduate students. students in behavior. Prerequisite: Prerequisites: BIOBM 330 or 331 or permission of instructor. S-U grades equivalent, BIONB 491 or equivalent, and optional. Sem TBA. Not offered 2001-2002. written permission of instructor. S-U grades PLANT BIOLOGY (BIOPL) Staff. optional. Lab TBA. Not offered 2001-2002. BIOPL 240 Green World/Blue Planet A seminar on a specific topic in animal Staff. Spring. 3 credits. S-U grades optional. Lees, behavior. The instructor presents lectures A two-week intensive laboratory course T R 1:25-2:40. K. J. Niklas, E. R. Turgeon, during the first few course meetings; the designed to provide experience with a specific T. G. Owens. remainder of the course is devoted to student technique currently used in cellular and This course focuses on helping individuals presentations. Topic and instructor are listed molecular neurobiology. The technique under understand how scientific information relates in the division’s catalog supplement issued at study and instructor in charge vary from to the issues they face as citizens, in manage­ the beginning of the semester.] semester to semester and are listed in the ment decision making, and in public policy. division’s catalog supplement issued at the [BIONB 724 Field Methods in Animal To what extent should genetic engineering of beginning of the semester.] Behavior crop plants be permitted? Should we place Fall or spring. Variable credit. May be [BIONB 793 Advanced Topics in limits on fossil fuel consumption as a means repeated for credit. Primarily for graduate Integrative Neurobiology of limiting global warming and global climate students in behavior. Prerequisite: Fall or spring. Variable credit. May be change? Must human endeavors be restricted permission of instructor. S-U grades repeated for credit. Primarily for graduate in certain areas to maintain diversity? The optional. Sem and fieldwork TBA. Not students; written permission of instructor format of this course is interactive, with offered 2001-2002. Staff. required for undergraduates. S-U grades lectures and discussions about how we as a A seminar-field experience course designed optional. Lees and discs TBA. Not offered society deal with controversial issues. for first-year graduate students in animal 2001-2002. Staff. BIOPL 241 Introductory Botany behavior. Weekly seminars discussing field A course designed to provide in-depth Fall. 3 credits. Lees, T R 9:05; lab, M T V methodology, data collection, and hypothesis knowledge of current research in anatomical or R 1:25-4:25, or M W 7:30-10:30 p.m. testing are followed by an intensive period (10 and physiological bases of vertebrate and K. J. Niklas. days to two weeks) in the field. Specific topics invertebrate behavior. Readings are primarily Introductory botany for those interested in the and field sites vary from semester to semester. from specialty books and selected journal plant sciences. Emphasizes structure, Topic and instructor are listed in the division’s articles. Topic and instructor are listed in the reproduction, and classification of an- catalog supplement issued at the beginning of division’s catalog supplement issued at the giosperms and the history of life on earth. the semester.] beginning of the semester.] Laboratory emphasizes development of skills [BIONB 725 Behavioral Ecology [BIONB 794 Advanced Laboratory in handling plant materials, including Workshop Techniques in Integrative identification. First and second weeks of Fall. 2 credits. May be repeated for credit. Neurobiology laboratory are field trips, starting with the first Primarily for graduate students; written Fall or spring. Variable credit. May be day of classes. Those who register fo r an permission of instructor required for repeated for credit. Prerequisite: permis­ evening laboratory are still required to attend undergraduates. S-U grades only. Seminar sion of instructor based upon a personal the afternoon field trips. TBA. Offered alternate years. Not offered interview. Lab TBA. Not offered 2001- BIOPL 242 Plant Function and Growth 2001-2002. J. W. Bradbury, 2002. Staff. S. L. Vehrencamp. A laboratory in the integrative, or neuroethol- Spring. 3 credits. S-U grades optional. A hands-on workshop designed to familiarize ogical, approach to studies of animal Primarily for undergraduates in agricultural graduate students with quantitative techniques behavior. Designed to provide practical sciences, but also for any biological as applied to behavioral ecology. Only one working knowledge of research methods in sciences students wanting to know about technique area will be considered each year. anatomical, physiological, and behavioral plant function. Suitable as a second-level Possible areas include spatial statistics, approaches to studies of vertebrate and course for nonmajors to satisfy the biology multivariate statistics, bioacoustical analysis, invertebrate behavior. Laboratory technique to distribution requirement. Prerequisites: 1 and event analysis. The material covered in be covered and instructor are listed in the year of introductory biology and/or BIOPL the first weeks will attempt to bring everyone, division’s catalog supplement issued at the 241. Recommended: 1 year of introductory regardless of background, up to a common beginning of the semester.] chemistry. Concurrent enrollment in BIOPL starting point. All participants will do weekly 244 required of plant science undergradu­ homework, prepare an introduction to one ates and highly recommended for other Related Courses in Other Departments science majors. May not be taken for credit sub-area, and create homework problems for Animal Behavior (Psychology 535) that sub-area. The topic for a given semester . after BIOPL 342 except by written is listed in the catalog supplement issued at Biochemistry and Human Behavior (Psychology 361 permission of instructor. Evening prelims the beginning of the semester.] and Nutritional Sciences 361) Feb. 21 and March 28. Lees, M W F 10:10. Brain and Behavior (Psychology 425) P. J. Davies. [BIONB 790 Advanced Topics in Cellular How plants function and grow. Examples deal and Molecular Neurobiology Developmental Biopsychology (Psychology 422) with crop plants or higher plants where Fall or spring. Variable credit. May be Evolution and Development (BIOGD 480/780; possible, though not exclusively. Topics repeated for credit. Limited to graduate BIOEE 760; BIO G 400) include cell structure and function; plant students and advanced undergraduates metabolism, including photosynthesis; light studying neurobiology and behavior. Evolution of Human Behavior (Psychology 326) relations in crops; plant-water relations; water Prerequisite: BIONB 222. S-U grades Human Behavior: A Sociobiological Perspective uptake, transport, and transpiration; irrigation optional. Lees and sem TBA. Not offered (Anthropology 476) of crops; sugar transport; mineral nutrition; 2001-2002. Staff. Insect Behavior Seminar (Entomology 662) growth and development—hormones, A lecture-seminar course on selected topics in responses to light, flowering, fruiting, cellular and molecular neurobiology. Students Neurobiology of Animal Behavior (Biological dormancy, and abscission; stress; tissue read original papers in the scientific literature Sciences [BIOSM] 327) culture'; and genetic engineering of plants. and lead discussions of these articles. Primates and Evolution (Anthropology 490) Suggestions for topics may be submitted by [BIOPL 243 Taxonomy of Cultivated faculty or students to the chair of the Section Primate Behavior and Ecology (Anthropology 390) Plants (also HORT 243) of Neurobiology and Behavior. Topic and Teaching Experience (Biological Sciences Fall. 4 credits. Prerequisite: 1 year of instructor are listed in the division’s catalog [BIO G] 498) introductory biology or written permission of instructor. May not be taken for credit supplement issued at the beginning of the The Brain and Sleep (Psychology 440/640) semester.] after BIOPL 248. Lees, M W F 10:10; labs, Undergraduate Research in Biology (Biological M or W 2—4:25. Offered alternate years. Sciences [BIO G] 499) Not offered 2001-2Q02. M. A. Luckow. BIOLOGICAL SCIENCES - 2001-2002

A study of ferns and seed plants, their chemistry of natural products and biological A companion to BIOPL 343 with laboratory relationships, and their classification into assays in the discovery of chemicals and their activities that focus on the practice of plant families and genera, emphasizing cultivated role in nature will be described. Classical biotechnology. Students will transfer genes to plants. Particular emphasis is placed on examples of drug development, from quinine plants by a variety of methods, and will gaining proficiency in identifying and to taxol, in the course of chemical prospecting analyze their expression in the host genome distinguishing families and in preparing and will be discussed. An overall medicinal by use of reporter gene assays, and by the using analytic keys. Attention is also given to purpose in chemoprospecting will be preparation and analysis of nucleic acids. emphasized, with mention of specific the economic importance of taxa, to the basic BIOPL 348 The Healing Forest worldwide spread of diseases pressing for taxonomic literature, and to the elements of Spring. 2 credits. Prerequisites: introduc­ new drugs.] nomenclature.] tory biology or plant biology, or permis­ BIOPL 244 Plant Function and Growth, BIOPL 342 Plant Physiology, Lectures sion of instructor. Lec/disc, R 2:30-4:25. Laboratory Spring. 3 credits. Prerequisites: 1 year of Offered alternate years. D. M. Bates, Spring. 2 credits. Prerequisite: concurrent introductory biology and either concurrent E. Rodriguez. enrollment in BIOPL 242. May not be enrollment in BIOPL 344 or written An ethnobotanical consideration of the role of taken for credit after BIOPL 344. Disc and permission of instructor. May not be taken plants in traditional and western medicine. lab, MTorW 12:20-4:25. T. Silva. for credit after BIOPL 242 unless written Studies of indigenous and lay societies Experiments exemplify concepts covered in permission is obtained from instructor. illustrate the ecological, systematic, biochemi­ BIOPL 242 and offer experience in a variety of Lees, T R 10:10-11:25. T. G. Owens. cal, and cultural aspects of herbal medicines biological and biochemical techniques, from An integrated and interdisciplinary study of and are placed in the broader context of such the cellular to whole plant level. the processes that contribute to the growth, interdependent themes as the conservation of competition, and reproduction of plants. biological and cultural diversity, human BIOPL 245 Plant Biology Topics include, but are not limited to, plant health, bioprospecting, compensation for Summer (6-week session). 3 credits. water relations, membrane properties and indigenous knowledge, and sustainable Limited to 24 students. Lees, M-F 11:30- processes, photosynthesis, plant respiration, development. 12:45; labs, M W 2-5:00. T. Silva. mineral and organic nutrition, stress physiol­ BIOPL 404 Crop Evolution, Introductory botany, including plant identifica­ ogy, control of growth and development, and tion. Emphasizes structure, reproduction, and Domestication and Diversity (also responses to the environment. Emphasis is on PL BR 404) classification of flowering plants. Much of the the relationship between structure and Spring. 2 credits. S-U letter. Prerequisites: laboratory work is conducted outdoors taking function from the molecular to the whole- Genetics 281 or Plant Breeding 225 or advantage of several outstanding natural areas plant level. which are available for study. Those who lack permission of the instructor. Lees, T R college-level biology are expected to work BIOPL 343 Molecular Biology and 9:05. S. Kresovich. closely with the instructor on supplemental Genetic Engineering of Plants See PL BR 404, for description. instructional materials. Spring. 2 credits. Prerequisite: 1 year [BIOPL 440 Phylogenetic Systematics general biology or permission of instructor. Spring. 4 credits. Limited to 24 students. BIOPL 247 Ethnobiology S-U grades optional. Lees, T R 11:15. Prerequisite: introductory biology or Fa]l. 3 credits. S-U grades optional. Lees, T M. E. Nasrallah. permission of instructor. Lees, T R 10:10; R 11:15; disc, R 12:20 or 1:25, or F 12:20. An introduction to current studies involving labs, T R 2:00-4:25. Offered alternate D. M. Bates. recombinant DNA technology and its years. Not offered 2001-2002. K. C. Nixon. A consideration of the principles, methods, application to the improvement of plants. The Basic and advanced theory and methods of and issues of ethnobiology. Emphasis is on course emphasizes genetic transformation phylogenetic analysis. Students are introduced the past and present ecological, evolutionary, methodology, gene expression systems, and to cladistic analysis using parsimony and gain economic, and cultural interrelationships of strategies for increasing productivity. The experience with computer-aided analysis of humans in traditional and lay societies with course is directed towards undergraduates taxonomic data, including both morphological their plants and animals, as a means of who wish to become familiar with the theory and molecular data sources. Topics discussed understanding the place and future of humans and practice of plant biotechnology. in the biosphere. Traditional medicines, include applications of phylogenetic methods underutilized organisms, resource manage­ BIOPL 344 Plant Physiology, Laboratory to biogeography and evolutionary studies.] ment, and ownership of nature, and method­ Spring. 2 credits. Prerequisite: concurrent [BIOPL 441 Systematics and Evolution of ology are among the topics covered. enrollment in BIOPL 342. May not be Crops taken for credit after BIOPL 244. Similar to Fall. 2 credits. Prerequisite: an advanced- BIOPL 248 Taxonomy of Vascular Plants BIOPL 244 but at a more advanced level. Spring. 4 credits. Prerequisite: 1 year of level course in the plant sciences with Lab, R 1:25-4:25; disc, R 12:20. T. Silva. taxonomic content or permission of introductory biology. May not be taken for Experiments exemplify concepts covered in instructor. Lees, R 12:20-2:15. Offered credit after BIOPL 243. S-U grades BIOPL 342 and offer experience in a variety of alternate years. Not offered 2001-2002. optional. Lees, M W F 9:05; lab, W or R biological and biochemical techniques, from Staff. 1:25-4:25. J. I. Davis. the cellular to whole plant level, with An integrated study of the systematics and An introduction to the classification of emphasis on experimental design. vascular plants, with attention to the goals of evolution of agronomic and horticultural taxonomy, the processes of plant evolution, [BIOPL 345 Plant Anatomy species. Processes of domestication, the and the means of analyzing evolutionary Fall. 4 credits. Limited to 15 students. evolutionary history of selected cultigens, the relationships among plants. The laboratory Prerequisite: 1 year of introductory biology nature of weeds and land races, classification concentrates on methods of plant identifica­ or a semester of botany. Lees, M W 9:05; and nomenclature as applied to cultivated tion and presents an overview of vascular labs, M W 2-4:25. Offered alternate years. plants, and underexploited plant resources are plant diversity, with particular attention to the Not offered 2001-2002. Staff. among the topics considered.] flowering plants. A descriptive course with equal emphasis on [BIOPL 442 Current Topics in development and mature structure. Lecture, Ethnobiology [BIOPL 340 Methods in Chemical laboratory, and reading are integrated in a Prospecting Fall. 2 or 4 credits (4 credits with an study guide. The laboratory offers the independent research component and Spring. 2 credits. Prerequisites: Intro opportunity to develop the practical skills Biology (BIOG 101-104, 105-106, or 107- term paper). Prerequisites: BIOPL 247, 348, required to make anatomical diagnoses and to or permission of instructor. Lec/disc, T 108) required. Completion or concurrent write anatomical descriptions.] enrollment in organic chemistry, recom­ 2:30-4:25. Offered alternate years. Not mended. TBA. Offered alternate years. Not BIOPL 347 Laboratory in Molecular offered 2001-2002. E. Rodriguez, offered 2001-2002, E. Rodriguez and Biology and Genetic Engineering of D. M. Bates. M. Aregullin. Plants Explorations of the interrelationships of plants Student participants will learn theory and Spring. 2 credits. Limited to 24 students. and animals with humans from a wide range methodologies in ethnobotany, chemical Prerequisite: BIOPL 343 or permission of of perspectives. Topics considered are ecology and zoopharmacognosy as they apply instructor. Concurrent enrollment is BIOPL contemporary issues, theory, and methodol­ in a multidisciplinary fashion to chemical 343 is encouraged. S-U grades optional. ogy of ethnobotany and ethnobiology, and prospecting. The use of techniques in the Lab, W 12:25-4:25. M. E..Nasrallah. the role of plants and animals in human lives, in subsistence and exchange, and in thought.] PLANT BIOLOGY 171

BIOPL 443 Topics and Research BIOPL 449 Green Signals and Triggers— Selected experiments on gene expression, Methods in Systematics The Plant Hormones (also HORT gene transfer, and assay of reporter genes in Fall or spring. 1-2 credits (1 credit per 449) plants. The course emphasizes the application section). Prerequisite: written permission Fall. 1 credit. S-U grades optional. of molecular biology methodology to plant of instructor. S-U grades optional. Staff. Prerequisite: 1 year of introductory biology systems. Additional lab time is required to A series of one-credit modules on specialized and permission of instructor. F 1:25-2:15. complete assignments. topics in systematics. Topics and instructors Offered alternate years. P. J. Davies. vary each semester. May not be taught every A study of plant hormones and how they [BIOPL 642 Plant Mineral Nutrition (also semester. Topics and instructors are listed in regulate plant growth and development. CSS [SCAS] 642) the division’s catalog supplement issued at the Topics covered include the discovery, role in Spring. 3 credits. Prerequisite: BIOPL 342 beginning of the semester. growth and development, mode of action, and or equivalent. Lees, M W F 10:10. Offered practical uses of the plant hormones auxin, alternate years. Not offered 2001-2002. BIOPL 444 Plant Cell Biology gibberellins, cytokinins, abscisic acid, L. V. Kochian, R. M. Welch. Fall. 4 credits. Limited to 24 students. ethylene, and brassinosteroids. A detailed study of the processes by which Prerequisites: 1 year of introductory plants acquire and use mineral nutrients from biology or permission of instructor. Lees, [BIOPL 452 Systematics of Tropical the soil. Topics include: the uptake, transloca­ M W F 9:05; lab, M or W l:25^f:25. Plants tion, and compartmentation of mineral R. O. Wayne. Spring. 3 credits. Prerequisite: BIOPL 243 elements; root-soil interactions; the metabo­ Evidence from microscopy, physiology, or BIOPL 248. Letter grades only. Lee, M W lism of mineral elements; the involvement of biochemistry, and molecular biology is used to 10:10; lab, T 1:25-4:25. Offered every three mineral nutrients in various physiological try to unravel the mystery of the living cell. years. Not offered 2001-2002. K. C. Nixon. processes; and the nutrition of plants adapted The dynamics of protoplasm, membranes, and The families of plants encountered solely or to extreme environmental stresses (e.g., acid the various organelles are studied. The chiefly in tropical regions are considered in a soils). Specific mineral elements are empha­ mechanisms of cell growth and division, the phylogenetic context in lectures, discussions, sized to illustrate these topics.] relationship of the cytoskeleton to cell shape and laboratory, with the aim of providing and motility, the interaction of the cell with its basic points of recognition for, and an BIOPL 644 Regulatory Factors in Plant environment, and the processes that give rise understanding of, diversity and relationships Growth and Development to multicellular differentiated plants are in these families.] Fall. 1-2 credits (1 credit per section). investigated. Prerequisites: BIOPL 242 or 342 or BIOPL 453 Principles and Practice of equivalent, or permission of instructor. [BIOPL 445 Photosynthesis Historical Biogeography (also Two modules, which can be taken Fall. 3 credits. Prerequisites: 1 year of ENTOM 453) together or in isolation. These are college chemistry and mathematics. Fall. 3 credits. Prerequisite: a course in compatible with BIOPL 652-03 and BIOPL Recommended: 1 year of college , physics systematics or permission of instructors. 653—04 (Molecular Aspects of Plant and plant physiology. Lees, M W F 10:10. S-U grades optional. Lees, T R 10:10; lab Development II and I respectively). S-U Offered alternate years. Not offered 2001- T 1:25-4:25. Offered alternate years. grades optional. Offered alternate years. 2002. T. G. Owens. J. K. Liebherr and M. A. Luckow. An introduction to the processes of photosyn­ A survey of techniques in historical biogeogra­ Section 01 Plant Hormones thesis in plants, algae and bacteria. An phy, and the development of modern Oct. 5. P. J. Davies. interdisciplinary approach is emphasized biogeographic theory in the context of Plant Hormones: their role in plant growth incorporating biochemical, biophysical, classical, ecological, and phylogenetic and development, analysis, biosynthesis and physiological and molecular perspectives. The analytical methods. Geological and paleonto­ mode of action, including signal transduction, course covers the range of processes involved logical aspects of biogeography will be examined from a physiological, biochemical in photosynthesis beginning with light presented, and large-scale biogeographic and molecular point of view. The course harvesting and primary photochemistry patterns discussed. Laboratories will focus on covers auxin, gibberellin, cytokinin, ethylene, through electron transport and inorganic computer applications and discussion of abscisic acid, brassinosteroids, and other carbon fixation. Emphasis is placed on the controversial issues. compounds as appropriate. regulation of photosynthesis from the cellular [BIOPL 454 Systematics of Tropical Sec 02 Phytochrome and Photomorpho­ to the whole-plant level.] Plants: Field Laboratory genesis BIOPL 447 Molecular Systematics Spring break. 1 credit. Limited to 15 1 credit. Lees M W F 9:05 (12 lees). Nov. 5- Dec. 7. P. J. Davies. Fall. 3 credits. Prerequisites: BIOEE 278 or students. Prerequisite: concurrent A study of the regulation of plant growth and BIOGD 281 or BIOBM 330, or BIOBM 332, enrollment in BIOPL 452 or permission of development by light as perceived through or written permission of instructor. Lees, T instructor. Letter grades only. For more the pigments phytochrome and cryptochrome. R 8:30-9:55. Offered alternate years. details and application, contact the This will include the biochemistry and J. J. Doyle. L. H. Bailey Hortorium, 467 Mann Library. molecular biology of phytochrome and the The theory and practice of using molecular Offered every three years. Not offered way in which phytochrome modulates plant evidence, particularly DNA sequence data, for 2001-2002. K. C. Nixon. growth, including molecular and genetic addressing diverse systematic and evolution­ An intensive orientation to families of tropical analysis of its effects, and the mechanisms by ary questions. Emphasis is on phylogeny flowering plants represented in forests of the which it acts. The role of phytochrome in reconstruction, particularly in eukaryotic American Tropics. Emphasis on field detecting and modulating growth in natural systems. The organization and evolution of identification combined with laboratory and agricultural environments will also be nuclear and organellar genomes is described analysis of available materials in a “whole- covered. from the standpoint of their suitability for biology” context.] systematic and evolutionary studies. BIOPL 456 Biomechanics of Plants (also BIOPL 647 Seminar in Systematic Botany [BIOPL 448 Plant Evolution and the ABEN 456) Fossil Record Fall. 3 credits. Prerequisites: upper division Fall or spring. 1 credit. May be repeated for credit. Prerequisite: written permission Spring. 3 credits. Prerequisite: BIOPL 241 undergraduate or graduate status, of course coordinator required for or equivalent, or permission of instructor. completion of introductory sequence in undergraduates. S-U grades optional. Sem, Lees, T R 9:05; lab, R 12:20-2:15. Offered biology and one year of calculus, or T 12:20. Bailey Hortorium staff. alternate years. Not offered 2001-2002. permission of instructor. S-U or letter grade Lectures and discussions led by staff, visitors, K. J. Niklas, W. L. Crepet. optional. Lees, T R 11:15-12:05; disc, W and students on topics of current importance An introduction to evolution, surveying major 2:30-3:20. J. R. Cooke and K. J. Niklas. to systematic botany. changes in plants from the origin of life to the See ABEN 456, for description. present. Emphasis is placed on plant form and BIOPL 641 Laboratory in Plant Molecular [BIOPL 648 Plant Biochemistry function, adaptations to particular ecologic Biology (also BIOBM 641) Spring. 3 credits. Prerequisites: organic settings, and evolutionary theory as it relates Fall. 4 credits. Prerequisites: BIOGD 281 or chemistry, biochemistry, and a course in to plants.] equivalent, BIOBM 330 or 331 or plant physiology. Lees, M W F 9:05. equivalent, and permission of instructor. Offered alternate years. Not offered 2001- S-U grades with permission of instructor. 2002. T. G. Owens and staff. Lab, T 9:05-4:30. J. B. Nasrallah, M. R. Hanson. BIOLOGICAL SCIENCES - 2001-2002

Selected areas of plant biochemistry are Section 01 Molecular Plant-Pathogen series. The course is divided into two sections: reviewed in the context of the plant life cycle Interactions (also PLPA 662) 1) Gene discovery; this covers genetic, and responses to the environment. Topics 1 credit. Lees, M W F 10:10 (12 lees) Jan. molecular, and genomics approaches to the include metabolism of lipids, carbohydrates, 21-Feb. 15. T. P. Delaney, A. R. Collmer, isolation of plant genes; and 2) Gene organic acids, phenolic compounds', and S. G. Lazarowitz. characterization; this covers DNA sequencing, proteins; nitrogen and sulfur assimilation; An examination of the molecular properties gene database analysis, assessment of gene respiration; photosynthesis; development and that control the development of host-parasite expression, production of transgenic plants, replication of chloroplasts; and cell-wall interactions in both microorganisms (bacteria, and functional genomics approaches. composition and properties. Attention is paid viruses, and fungi) and higher plants. Emphasis is on understanding the appropriate to operation of control mechanisms.] Contemporary theories describing the genetic approach that is needed for different and molecular mechanisms of microbial experiments. BIOPL 649 Physiology of Ion and Water pathogenesis and plant resistance are Section 02 Plant Biotechnology (also Transport in Plants discussed. Fall. 1-3 credits (1 credit per section). PLBR 653.2 and PLPA 663) Prerequisite: BIOPL 342 or equivalent, or Section 02 Molecular Plant-Microbe 1 credit. Lees, M W F 1:25 (12 lees) Oct. 1- permission of the instructor. Three Interactions (BIOMI 652) Oct. 29- M. Zaitlin, E. D. Earle. modules that may be taken independently, 1 credit. S-U grades optional. Lees, M T F This course deals with production and uses of although section 01 is recommended 12:20 (12 lees) Jan. 21-Feb. 15. Offered transgenic plants for agricultural and industrial before taking section 02. Offered alternate alternate years. S. C. Winans. purposes. Topics include procedures for gene years. R. M. Spanswick. Course focuses on the interactions of introduction and control of gene expression, Agrobacteria and R h iz o b ia with plants. Topics as well as strategies for obtaining transgenic Section 01 Basic Principles of Ion on Agrobacterium -plant interactions include plants that are resistant to insects, diseases, Transport and Electrophysiology plant-microbe recognition mechanisms, and herbicides, produce useful products, or 1 credit. Lees T R 10:10-11:30 (9 lees). T-DNA transfer process, oncogenesis, and use have improved nutritional and food process­ Aug. 30-Sept. 27. of Agrobacterium to produce transgenic ing characteristics. Regulatory and social The topic of this section is the biophysical plants. Topics on R h izo b iu m -plant interactions issues relating to plant biotechnology are basis of ion transport across cell membranes, include regulation of nitrogenase activity and discussed. including membrane structure, ion fluxes and expression, organization and function of the their measurement, the thermodynamic [Section 03 Plant Genome Organization sym plasmid, nodule development, and plant (also PLBR 653.3) criterion for active transport, and the genetics involved in plant-microbe interaction. relationship between ion transport and the 1 credit. S-U grades optional. Lees, M W F electrical properties of cell membranes. [Section 03 Molecular Aspects of Plant 10:10 (12 lees). Offered alternate years. Development II Not offered 2001-2002. S. D. Tanksley. Section 02 Ion Transport in Plants 1 credit. S-U grades optional. Lees, M f F This section covers the structure and variation 1 credit. Lees T R 10:10-11:30 (9 lees). Oct. 10:10 (12 lees) Not offered 2001-2002. of plant nuclear genomes, including changes 2-Oct. 30. Staff. in genome size, centromere/telomere Topics include: transport of the major ions in The molecular genetics of plant development. structure, DNA packaging, transposable plant cells and whole plants; properties of This module focuses on vegetative develop­ elements, genetic and physical mapping, proton ATPases and their relationship to the ment and includes topics such as the positional gene cloning, genomic sequencing transport of ions, sugars, and amino acids at development of the shoot, root, and vascula­ and comparitive genomics.] the plasma membrane and tonoplast; ion ture, and the operation of the vegetative shoot channels in plant cell membranes; intercellular Section 04 Molecular Aspects of Plant apical meristem. The module is a companion Development I ion transport via plasmodesmata; and long to BIOPL 653, Sec 04 (Plant Development I).] distance ion transport in higher plants. 1 credit. Lees, M W F 10:10 (12 lees) Oct. [Section 04 Plant Gene Evolution and 31-Nov. 30. J. B. Nasrallah. Section 03 Water Transport in Plants Phylogeny This module focuses on the molecular 1 credit. Lees T R 10:10-11:30 (11 lees). 1 credit. Lees, M W F 1:25 (12 lees). genetics of plant development. Current Nov. 1-Dec. 6. Offered alternate years. Not offered 2001- approaches to the elucidation of the molecular Topics include: water relations of plant cells 2002. J. J. Doyle. signals and pathways that lead to the and tissues using water potential terminology; Practical applications of molecular systemat- establishment of the differentiated state of permeability of plant cells to water and the ics/evolution for plant molecular biologists floral cells and organs are discussed. Topics role of aquaporins; and transport of water and other non-systematists. The course include cell-cell signalling in the establishment through whole plants, including transpiration, focuses on two basic issues: methods and of pattern and differentiation of specialized stomatal physiology and the effects of water principles for inferring relationships among cell types, and the control of developmental stress. genes and the use of data to hypothesize pathways by endogenous and external cues. [BIOPL 651 Quantitative Whole-Plant relationships among plants. Evolutionary The module is a companion to BIOPL 652, Physiology patterns and processes of genes and gene Sec 02 (Molecular Aspects of Plant Develop­ Fall. 3 credits. Prerequisites: introductory families are discussed, as well as rates of ment II). physics, calculus, and plant physiology. sequence evolution, paralogy and orthology, Section 05 Molecular Biology of Plant S-U grades only. Lees, T R 10:10-11:30. the effects of recombination and concerted Organelles Offered alternate years. Not offered 2001- evolution of gene phytogenies, and the 1 credit. S-U grades with permission. Lees, 2002. R. M. Spanswick. implications of using gene or allele phytog­ M W F 1:25 (12 lees) Oct. 31-Nov. 30. An exploration of the extent to which enies to infer organismal evolutionary M. R. Hanson, D. B. Stem. physiological processes and their interactions patterns.] An in-depth examination of the molecular can be formulated in a quantitative manner BIOPL 653 Plant Molecular Biology I biology of plant mitochondria and plastids. and integrated to describe various aspects of Fall. 1-5 credits (1 credit per section). Topics include the organization, evolution, plant behavior, including growth and yield. Prerequisites: BIOGD 281 and BIOBM 330 and expression of organelle genomes, RNA Consideration is given to characterization of or 332, or their equivalents. Recom­ editing, and the expression of nuclear genes the plant environment, energy balance, gas mended: BIOBM 331. S-U grades optional. encoding structural or regulatory organelle exchange, water relations, photosynthesis, Coordinator: J. B. Nasrallah. proteins. Special topics include respiration, translocation, nutrient supply, and A series of four-week modules on specialized mitochondrially-encoded cytoplasmic male the timing of developmental events.] topics. sterility, transformation and expression of BIOPL 652 Plant Molecular Biology II foreign genes in chloroplasts, and the use of Section 01 Concepts and Techniques in genetics to investigate nucleus-organellar Spring. 1-2 credits (1 credit per section). Plant Molecular Biology (also PLPA 663.01) interactions. Prerequisites: BIOGD 281 and BIOBM 330 1 credit. Lees, M W F 10:10 (12 lees) Sept. or 332, or their equivalents. Recom­ 3-Sept. 28. T. P. Delaney, G. B. Martin. Section 06 Molecular Breeding and mended: BIOBM 331. S-U grades optional. This is an introductory module that provides a Genetic Diversity (also PLBR 653.6) A series of four-week modules on specialized broad overview of molecular biology concepts 1 credit. Lees M W F 10.T0 (12 lees). topics. Coordinator: J. B. Nasrallah. relevant to the plant sciences. This section Oct. 1-Oct. 29. Offered alternate years. serves as a prerequisite to other modules in S. D. Tanksley. the BIOPL 653 (fall) and BIOPL 652 (spring) COURSES IN MARINE SCIENCE 173

Application of DNA markers to the identifica­ BIOPL 746 Research Seminar in the ecology and evolutionary biology program tion, manipulation and isolation of genes Systematic Botany of study requirements, students in marine important to plant and animal productivity Spring. 1 credit. Limited to graduate biology are encouraged to enroll in the using molecular genetic techniques. Students students, except by permission of following courses: learn how to design and execute experiments instructor. Disc, T 12:20. Bailey Hortorium to identify quantitative trait loci (QTLs), as staff. 1) BIOEE. 154, The Sea: An Introduction to well as how to apply molecular markers to A student-led seminar presentation based on Oceanography, plant and animal breeding programs. his or her thesis research or a related topic. 2) BIOSM 364, Field Marine Science, BIOSM [BIOPL 654 Botanical Nomenclature BIOPL 749 Graduate Research in Botany 375 Field Marine Biology and Ecology, or a 400-level BIOSM field course at the Fall or spring. Variable credit. May be Fall. 1 credit. Prerequisite: written Shoals Marine Laboratory, permission of instructor. S-U grades only. repeated for credit. S-U grades optional. Lee and disc TBA. Offered alternate years. Hours TBA. Staff. 3) BIOEE 462, Marine Ecology. Not offered 2001-2002. Staff. Similar to BIO G 499 but intended for Undergraduate Specialization In Ocean An analysis of the International Code of graduate students who are working with Sciences Botanical Nomenclature and its application to faculty members on an individual basis. various plant groups.] Science of Earth Systems majors have the [BIOPL 840 Current Topics in Plant option of specializing their program of study BIOPL 656 Topics in Plant Evolution Physiology in the area of ocean sciences. This interdisci­ Spring. 1 credit. Prerequisite: BIOPL 448 or Fall or spring. 2 credits. May be repeated plinary specialization is intended for students equivalent background in evolution, or for credit. S-U grades only. Sem TBA. Not with interests in understanding the interaction written permission of instructor. Lab and offered 2001-2002. Staff. of biological, chemical, geological, and disc TBA. Offered alternate years. Molecular aspects of plant hormone biosyn­ physical processes in ocean systems. In K. J. Niklas. thesis, signal transduction and action. Seminar addition to fulfilling the Science of Earth A series of selected topics to provide a reports by graduate students on current Systems general requirements (see the SES background in plant evolution, paleobotanical literature in experimental plant physiology or program description in Interdisciplinary literature, and evolutionary theory. Among the related areas.] Centers, Programs, and Studies section of topics discussed are the origin of a terrestrial catalog), students in ocean sciences are flora, the evolution of the seed plants, and the Related Courses in Other Departments required to take four advanced Courses from origin and adaptive radiation of the an- Introductory Mycology (Plant Pathology 309) the following list to fulfill their major giosperms. Marine Botany: Ecology of Marine Plants (Biological requirements: BIOPL 740 Plant Biology Seminar Sciences [BIOSM] 449) 1) BIOEE 373 Biology of the Marine Fall and spring. No credit (no official Mycology Conferences (Plant Pathology 649) Invertebrates registration). Required of graduate students 2) BIOEE 457 Limnology doing work in plant biology. Sem, F 11:15. Physiological Plant Ecology, Lectures and 3) BIOEE 462 Marine Ecology Staff. Laboratory (Biological Sciences [BIOEE] 466 Lectures on current research in plant biology, and 468) 4) BIOEE 478 Ecosystem Biology presented by visitors and staff. Phytomycology (Plant Pathology 709) 5) BIOES 490 Topics in Marine Biology BIOPL 741 Problems in Plant Cell and Plant Ecology and Population Biology, Lectures and 6) BIOSM 303 Ecology of Marine Fishes Molecular Biology Laboratory (Biological Sciences [BIOEE] 463 7) BIOSM 308 Marine Microbial Ecology Fall. 2 credits. Limited to first- and second- and 465) 8) BIOSM 309 Climates and Ecosystems year graduate students in the Plant Cell Plant Ecology Seminar (Biological Sciences 9) BIOSM 329 Ecology of Animal Behavior and Molecular Biology Program. Disc TBA. [BIOEE] 669) Staff. 10) BIOSM 364 Field Marine Science Plant Cytogenetics Laboratory (Plant Breeding 446) An introduction to the research literature in 11) BIOSM 365 Underwater Research plant molecular and cellular biology through Teaching Experience (Biological Sciences 12) BIOSM 374 An Introduction to Field weekly problem sets and discussions. [BIO G] 498) Ornithology BIOPL 742 Current Topics in Plant Undergraduate Research in Biology (Biological 13) BIOSM 375 Field Marine Biology and Molecular Biology Sciences [BIO G] 499) Ecology Fall or spring. 1 credit. Enrollment is 14) BIOSM 413 Experimental Marine Ecology limited. Primarily for graduate students, 15) BIOSM 418 Tropical Marine Science with preference given to majors or minors COURSES IN MARINE SCIENCE 16) BIOSM 449 Seaweeds, Plankton and in plant molecular biology; written Seagrass permission of instructor required for Cornell offers an extensive listing of under­ 17) BIOSM 376 Marine Invertebrate Zoology undergraduates. S-U grades only. Sem, graduate courses in marine science. 1 hour each week TBA. Staff. (note: not the same as BIOEE 373) Undergraduates interested in pursuing studies 18) BIOSM 477 Marine Vertebrates BIOPL 743 Current Research in Plant in marine science are encouraged to explore 19) EAS 375 Sedimentology and Stratigraphy Cell and Molecular Biology the undergraduate specialization in Marine Fall. 1 credit. Limited to graduate students; Biology offered through the Division of 20) EAS 455 Geochemistry written permission from a member of the Biological Sciences, the undergraduate 21) EAS 475 Special Topics in Oceanography Plant Cell and Molecular Biology Program specialization in Ocean Sciences offered 22) EAS 479 Paleobiology or by permission of coordinator required through the Science of Earth Systems Program, 23) NTRES 306 Coastal and Oceanic Law and for undergraduates. Disc TBA. Staff. and the summer program of courses offered Policy An introduction for graduate students to the by the Shoals Marine Laboratory. Further 24) NTRES 417 Wetland Resources research being conducted by Cornell faculty in information on these programs can be found the Plant Cell and Molecular Biology Program. at the Cornell Marine Programs Office, G14 Sea Semester Stimson Hall, or on their web site, BIOPL 745 Current Topics in BIOSM 366 SEA: Introduction to www.sml.cornell.edu. Systematics Oceanography Fall. 1 credit. Limited to graduate students, Undergraduate Specialization in Marine BIOSM 367 SEA: Introduction to Maritime except by permission of instructor. S-U Biology and Oceanography Studies grades optional. Disc, T 12:20. Bailey Biological Sciences majors in the ecology and Hortorium staff. BIOSM 368 SEA: Introduction to Nautical evolutionary biology program of study have A seminar with presentations and discussion Science the option of specializing their program of by students of original research papers in BIOSM 369 SEA: Practical Oceanography I systematic biology. study in the area of marine biology. This specialization is intended for students with BIOSM 370 SEA: Practical Oceanography II interests in understanding the unique aspects of organismal biology in the marine environ­ These courses must be taken concurrently. ment. In addition to fulfilling the major and Special program run by the Sea Education 174 BIOLOGICAL SCIENCES - 2001-2002

Association. Contact Marine Programs Office BIOSM 160 The Oceanography of the populations and communities, and analysis of (607-255-3717) for more details. Gulf of Maine current research emphasizing theory and its Summer. 4 credits. Limited to 24 students. potential uses in fisheries’ management. Lab Students in both marine science specializa­ A special 2-week course offered aboard a and field activities emphasize collection and tions are exposed to an integrated program of SEA vessel and at Cornell’s Shoals Marine analysis of data from the Gulf of Maine and study, emphasizing a natural progression of Laboratory (SML) on an island off adjacent estuarine habitats. formal course work combined with ample Portsmouth, N.H. For more details or an opportunities for practical field experience. application, contact the SML office, G14 BIOSM 308 Marine Microbial Ecology Stimson Hall or the Sea Education Summer. 4 credits. Prerequisite: 1 year of Association office at P.O. Box 6, Woods college-level biology. S-U grades optional. Hole, MA, 02543. Daily lees, labs, and A special 2-week course offered at Shoals SHOALS MARINE LABORATORY fieldwork for 2 weeks. SML faculty. Marine Laboratory (SML), on an island off Portsmouth, N.H. For more details or an (BI0SM) An exciting opportunity to explore the offshore and near-coastal environments of the application, consult the SML office, G14 G14 Stimson Hall, 255-3717 Gulf of Maine for pre-college and first-year Stimson Hall. Daily lees and fieldwork for 2 weeks. SML staff. The objective of the Shoals Marine Laboratory non-science majors. Students spend 10 days aboard the Sea Education Association’s sailing This course examines the fundamental role of (SML) is to provide undergraduates and other marine microbial communities in the function interested adults a unique opportunity to vessels round trip between Woods Hole, Mass., and the Isles of Shoals via Georges of the biosphere. Lectures survey bacterial, explore marine sciences in an island setting protozoan, and micrometazoan assemblages noted for its biota, geology, and history. SML Bank and the Gulf of Maine. Besides operating the ship, students study the many from Arctic to deep sea vent communities. has established a national reputation for Laboratory exercises cover several principal excellence and has become North America’s characteristics of this unique ocean environ­ ment. Following the sea component, students techniques of field microbial ecology and largest marine field station focusing on explore the rich marine microbial environment undergraduate education. spend seven days at the Shoals Marine Laboratory collecting data characteristic of the surrounding the Isles of Shoals. The summer population of Appledore Island Isles of Shoals coastal environment. BIOSM 309 Climates and Ecosystems is limited to about one hundred people at any Summer. 4 credits. Prerequisite: 1 year of one time. Participants and faculty members BIOSM 161 Introduction to Field Marine Science college-level biology; background can literally and figuratively immerse preferred in physics/physical geography. themselves in their explorations, free from Summer. 4 credits. S-U grades optional. A special 2-week course offered in coopera­ S-U grades optional. A special 2-week distractions common to most academic course offered at Cornell’s Shoals Marine institutions. Because SML is a residential tion with Rider University at Cornell’s Laboratory (SML) on an island off facility, a sense of community develops that Shoals Marine Laboratory (SML) on an Portsmouth, N.H. For more details or an makes courses and seminars at SML outstand­ island off Portsmouth, N.H. Apply directly application consult the SML office, G14 ing educational and intellectual experiences. to Rider University, College of Continuing Stimson Hall. Daily lees, labs, and Participants leam from and exchange ideas Studies. Contact Dr. Richard Alexander for fieldwork for 2 weeks. SML faculty. with a wide range of specialists whose application and information at: A study of the fundamentals of organism- primary interests are marine but whose [email protected], (609) 895- environment interaction developed through perspectives often differ, providing fertile 5422. Offered alternate years. defining and measuring abiotic factors ground for lively discussions. This course allows students who are not biology majors to experience the breadth of including solar radiation, temperature, Credit courses at Shoals Marine Laboratory are the marine sciences under field conditions at atmospheric moisture, precipital wind, and full-time, intensive learning experiences. an island laboratory. Aspects of biology, currents. On-site exploration of the dynamics Courses may be taken sequentially, but not geology, earth science, chemistry, and physics of meteorology and the role of abiotic and concurrently. A typical day combines lecture are included. Specific topics include: beach, biotic factors in the life of coastal and marine sessions, laboratory and field work, field trips salt marsh, tidal mud flat, tide pool, and plants and animals including humans. to nearby islands and the mainland, and benthic offshore environments; identification collecting and research excursions aboard the BIOSM 329 Ecology of Animal Behavior of marine plants and animals; chemical and (also BIONB 329) Laboratory’s 47-foot research vessel, Jo h n M. physical oceanography; marine geology; and Summer. 4 credits. Prerequisite: 1 year of K ingsbury. Field experience is an integral ecology of kelp beds and urchin barrens. component of all courses, using Appledore’s introductory college biology. Recom­ extensive intertidal and subtidal zones, wading BIOSM 204 Biological Illustration mended: course work in ecology, bird rookeries, and seabird colonies. Faculty, Summer. 2 credits. A special 1-week psychology, or behavior. S-U grades optional. A special 2-week course offered drawn from Cornell University, the University course offered at Cornell’s Shoals Marine of New Hampshire, and other leading Laboratory (SML) on an island off at Cornell’s Shoals Marine Laboratory academic institutions, are selected not only Portsmouth, N.H. For more details or an (SML) on an island off Portsmouth, N.H. based on their academic excellence, but also application, consult the SML office, G14 For more details and an application, on their teaching ability in the field. In Stimson Hall. Daily sessions for 1 week. consult the SML office, G14 Stimson Hall. addition, there are numerous guest lecturers SML faculty. Daily lees, labs, and fieldwork for 2 weeks. including engineers, coastal planners, and General discussion of scientific publishing, SML faculty. specialists from private industry, government, illustration labeling, color techniques, and The ecological significance of behaviors of coastal organisms, with emphasis on field and and the academic community. printing processes. The course provides the scientist or science student a chance to laboratory research methods. Lectures and The Ithaca campus functions of the Shoals experience several illustration techniiques with readings address the major subareas of Marine Laboratory are centered in the Cornell the goal of obtaining an overview of scientific behavior (communication, orientation, social Marine Programs Office, G14 Stimson Hall. and wildlife illustrations. The student may behavior, foraging, predator avoidance, and The office serves as an advising center for choose a single technique to explore in depth. sensory mechanisms). Each student engages in students interested in the marine sciences, Course size is limited so that individual short-term behavioral observation and maintains a browsing library with updated attention can be emphasized. prepares a research proposal for studying a information on graduate study and career problem within the course subject area. opportunities as well as on marine programs BIOSM 303 Ecology of Marine Fishes at other institutions, and administers the SEA Summer. 4 credits. Prerequisite: 1 year of [BIOSM 363 Marine Biology for Teachers Semester, a 17-credit program offered in college-level biology. SCUBA certification Summer. 3 or 4 credits (4-credit option: cooperation with the Sea Education Associa­ recommended, not required. S-U grades additional 4 days for individual research). tion (SEA). optional. A special 2-week course offered Primarily for teachers, grades 6 through 12, but open to others with teaching experi­ The following marine sciences courses are at Shoals Marine Laboratory (SML), on an island off Portsmouth, N.H. For more ence. Prerequisite: 1 year of introductory currently administered by the Cornell Marine college biology. S-U grades optional. A Programs Office. details or an application, consult the SML office, G14 Stimson Hall. Daily lees and special 10-day course offered at Cornell’s fieldwork for 2 weeks. SML staff. Shoals Marine Laboratory (SML), on an This course presents principles, models, and island off Portsmouth, N.H. For more methods for analysis of dynamics of fish details or an application, consult the SML SHOALS MARINE LABORATORY 175

office, G14 Stimson Hall. Daily lees, labs, An introduction to field ornithology focusing BIOSM 413 Experimental Marine Ecology and fieldwork for 10 days. Offered on the biology, ecology, and behavior of the Summer. 6 credits. Prerequisite: 1 year of alternate years. Not offered 2001-2002. avifauna on the Isles of Shoals. The course college level biology; experience in SML faculty. focuses on field work designed to observe ecology or physiology recommended. S-U Designed to give an overview of living marine and study many concepts frequently taught in grades optional. A special 3-week course organisms (algae, invertebrates, fishes, marine the classroom setting including territoriality, offered at Cornell’s Shoals Marine mammals, and shorebirds) and of the breeding biology, and survivorship. Students Laboratory (SML) on an island off environment they inhabit. Fieldwork is learn and apply numerous ornithological field Portsmouth, N.H. For more details or an emphasized. Occasional lectures and films methods including various census techniques, application, consult the SML office, G14 deal with additional topics such as coastal- territory mapping, banding, behavioral Stimson Hall. Daily lees, labs, and zone problems, marine fisheries, economics of observations, and creating a field notebook. fieldwork for 3 weeks. SML faculty. marine organisms, and educational resources An introduction to the physiological ecology of the marine environment.] BIOSM 375 Field Marine Biology and and functional morphology of marine plants Ecology (FMBE) BIOSM 364 Field Marine Science (FMS) and animals, with emphasis on selected algal Summer. 6 credits. Prerequisites: 1 full year and invertebrate examples from the Gulf of Summer. 6 credits. Prerequisite: 1 year of of college level biology. S-U grades Maine. Topics covered include photosynthesis college biology. S-U grades optional. A optional. A 4-week course offered at in the marine environment; respiration in special 4-week course offered twice each Cornell’s Shoals Marine Laboratory (SML), intertidal organisms; carbohydrates, proteins, summer at Cornell’s Shoals Marine on an island off Portsmouth, N.H. For and lipids as nutrients in the sea; acclimation Laboratory (SML), on an island off more details or an application, consult the and tolerance of tide-pool biota; and Portsmouth, N.H. For more details or an SML Office, G-14 Stimson Hall. Daily lees, biological responses to competition and application, consult the SML Office, G14 labs, and fieldwork for 4 weeks. SML grazing. Field and laboratory exercises explore Stimson Hall. Daily lees, labs, and faculty. principles and procedures used to characterize fieldwork for 4 weeks. 3 core faculty Designed for students seeking an introduction the physical, chemical, and biotic environment members assisted by up to 15 visiting to the marine sciences and marine ecology; of intertidal and shallow subtidal organisms, lecturers, including representatives of FMBE emphasizes field work in natural including determination of temperature, light, governmental agencies. SML faculty. habitats. Examines aspects of the biology and salinity, oxygen and nutrient levels, and in Designed for the student who desires an initial ecology of marine organisms, including vivo functional analyses of metabolic overview of the marine sciences, this course intertidal plants and invertebrates, fishes, phenomena. The process of scientific emphasizes living material in natural habitats. marine mammals and birds, biological investigation is the predominant theme of the Most of the course work is concerned with the oceanography, and human impacts on the course. biology of intertidal plants and animals, marine environment. FMBE places a special biological oceanography, ichthyology, and emphasis on the ecology of the intertidal BIOSM 418 Tropical Marine Science fisheries. Attention is also given to introduc­ zone, and ecological, evolutionary and Summer. 8 credits and 4 credits of tory physical and chemical oceanography and physiological adaptations of marine organ­ research. Limited to 12 students. Prerequi­ marine geology. Marine ecology and the isms. Students may not take FMBE after taking sites: 1 year college-level biology; effects of human activity on the marine FMS (BIOSM 364). recognized SCUBA certification; medical environment are included. Students apply this exam; and permission of instructor. A BIOSM 376 Marine Invertebrate Zoology knowledge by conducting a transect study special 8-week course offered in Akumal, toward the end of the course. FMS places Summer. 6 credits. Prerequisite: 1 year of Mexico. For more details, contact Shoals emphasis on ichthyology, fisheries biology, introductory biology and permission of Marine Laboratory, G-14 Stimson Hall, general oceanography (biological, physical, instructors. S-U grades optional. A special 255-3717. For certified divers only. Lee/ and chemical) and marine geology. FMBE 3-week course offered at Cornell’s Shoals lab, 2 weeks; 6 weeks monitoring study (BIOSM 375) places an additional emphasis Marine Laboratory (SML), on an island off and individual research projects, including on ecology, especially in the intertidal zone; Portsmouth, N.H. For more details or an data analysis on computers. SML faculty. ecological, evolutionary and physiological application, consult the SML office, G14 In addition to lectures and laboratories adaptations of marine organisms; and field Stimson Hall. Daily lees, labs, and covering the basic principles of coral reef experiments. fieldwork for 3 weeks. Offered alternate ecology, students participate in a coral reef years. SML faculty. BIOSM 365 Underwater Research monitoring survey. Following two weeks of An introduction to the biology and evolution course work, students engage in independent Summer. 4 credits. Prerequisites: 1 year of of the major invertebrate phyla, concentrating research projects. college-level biology, recognized scuba on marine representatives. Emphasis is placed certification, and a medical examination. on the evolution of form and function, and BIOSM 449 Seaweeds, Plankton and S-U grades optional. A special 2-week the ecology, behavior, physiology, chemical Seagrass: the Ecology and course offered at Cornell’s Shoals Marine ecology, and natural history of invertebrates. Systematics of Marine Plants Laboratory (SML), on an island off Appledore Island’s unique location provides Summer. 4 credits. Prerequisite: BIOSM Portsmouth, N.H. For more details or an an excellent venue for the study of freshly 364 or 1 year of introductory biology. S-U application, consult the SML office, G14 collected and in situ representatives of most grades optional. A special 2-week course Stimson Hall. Daily lees and fieldwork for of the major phyla. offered at Cornell’s Shoals Marine 2 weeks. Team-taught by 3 faculty Laboratory (SML), on an island off members with occasional guest lecturers. BIOSM 402 Marine Pollution Portsmouth, N.H. For more details or an Not for recreational divers. Summer. 4 credits. Prerequisites: 1 year of application, consult the SML office, G14 Course covers the philosophy of research, college-level biology and chemistry or Stimson Hall. Daily lees, labs, and hypothesis testing and experimental design, permission of instructor. S-U grades fieldwork for 2 weeks. SML faculty. sampling methods, various underwater optional. A special 2-week course offered An overview of the major marine algal groups, techniques, diving physics and physiology, at Cornell’s Shoals Marine Laboratory including aspects of anatomy, morphology, and use of dive tables. Emphasis is on subtidal (SML), on an island off Portsmouth, N.H. development, life histories, physiology, and ecological research. Requirements include For more details or an application, consult use. Laboratories and fieldwork emphasize critical evaluation of several journal articles the SML office-, G14 Stimson Hall. Daily relationships between distribution and major and production of a research proposal. lees, labs, and fieldwork for 2 weeks. environmental parameters and involve student Offered alternate years. SML faculty. projects. BIOSM 374 Field Ornithology (An An introduction to marine pollutants; their Introduction) sources and control/treatment; the effects of EAS 475 Special Topics in Summer. 4 credits. Prerequisite: 1 year of marine pollution upon coastal ecosystems; Oceanography: Satellite Remote college-level biology. S-U grades optional. and federal and state water pollution Sensing in Biological Oceanography A special 2-week course offered at Shoals regulatory programs. Laboratory includes Summer. 6 credits. Prerequisites: 1 course Marine Laboratory (SML), on an island off training in field collection of water samples, in oceanography and/or marine biology, Portsmouth, N.H. For more details or an measurement and modeling of effluent plume or permission of the instructor. Strong application, consult the SML office, G14 dispersion, and measurement of microbial computer skills are desired. S-U grades Stimson Hall. Daily lees and fieldwork for indicators of water quality, dissolved nutrients, optional. A special 4-week course offered 1 weeks. SML staff. BOD, dissolved oxygen, and toxicity. at Cornell’s Shoals Marine Laboratory (SML), on an island off Portsmouth, N.H. 176 BIOLOGICAL SCIENCES - 2

and on campus at Cornell University. For two credits with a two-hour discussion per Laboratory (SML), on an island off more details or an application, consult the week. Offered at the Shoals Marine Laboratory Portsmouth, N.H. For more details or an SML office, G14 Stimson Hall. Daily lees, (summer) for one credit with four one-hour application, consult the SML office, G14 labs, and fieldwork for 4 weeks. Offered discussions per iveek for four weeks. Stimson Hall. Daily lees, labs, and alternative years. SML faculty. fieldwork for 1 week. SML faculty. BIOSM 495 Research Methods in Marine This course examines the complete history of “Remote Sensing” provides hands-on research Biology experience in hydrologic optics and satellite the Isles of Shoals from Paleozoic intrusion, Summer. 1 credit. Prerequisite: concurrent remote sensing to advanced undergraduate deformation and metamorphism to recent enrollment in BIOSM 499, or permission of and beginning graduate students. The course glaciation, sea-level change and wave erosion. instructor. Primarily for undergraduates. A is comprised of four principal parts, each Students will learn basic surveying and special 8 week course offered at the Shoals taught by a separate team of instructors at two mapping techniques using the Brunton Marine Laboratory (SML). For more details different locations: Part 1 (nine days) will be compass, poplevel/stadia rod, autolevel and or an application, consult the SML office, conducted at Shoals Marine Laboratory and GPS. Field efforts will focus on creating a G14 Stimson Hall. Weekly seminars for 8 aboard the R/V Kingsbury in waters surround­ series of thematic maps depicting island weeks. J. G. Morin and M. J. Shulman. ing the Isles of Shoals. Part 1 will be devoted characteristics such as topography, bedrock Seminar course on research methodology, to the theory and measurement of seawater geology and structure, vegetation, and land experimental design, statistical analyses, and optical properties, emphasizing the depen­ use patterns. scientific writing. The course is designed to dency of apparent optical property on assist students in the research they are NTRES 306 Coastal and Oceanic Law chlorophyll and dissolved organic matter conducting while enrolled in BIOSM 499. and Policy ■concentrations. Parts 2-4 (19 days) will be Summer. 2 credits. A special 1-week conducted at the Science of Earth Systems’ BIOSM 499 Research in Biology course offered at Cornell’s Shoals Marine computer laboratory on the Cornell campus. Summer. Credits variable (2 credits/7 days Laboratory (SML) on an island off Part 2 will cover satellite remote sensing of the on site). For more details and an applica­ Portsmouth, N.H. For more details or an apparent optical properties of seawater with tion, consult the SML Office, G14 Stimson application, consult the SML office, G14 an emphasis on processing SeaWiFS data Hall. Stimson Hall. Daily lees and disc for 1 using SeaDAS software and IDL programming Section A: Independent Biological week. SML faculty. language. Part 3 addresses satellite remote Research: Independent study with a member Intended for people interested in careers in sensing of physical oceanographic processes management of marine or coastal resources or that influence ecosystem dynamics with an of the Shoals Marine Laboratory core faculty, based on student faculty interest and available in the natural sciences. Subjects include law emphasis on AVHRR-derived sea-surface and policy related to ocean dumping, marine temperature and SSM/I-derived ocean winds. facilities. A short proposal of research must be sent with application materials. sanctuaries, environmental impact statements, Part 4 is devoted to independent projects; water and air pollution, fisheries management, student will attempt to integrate SeaWiFS, Research Experiences for Undergraduates offshore gas and oil production, and territorial AVHRR, and SSM/1 data in order to address (REU) jurisdiction. Lectures on the status and history questions of biological-physical interactions. 0 credit. The National Science Foundation of law are accompanied by discussion of (NSF) Research Experiences for Under­ BIOSM 477 Marine Vertebrates relevant policy and analysis of the efficacy of graduates (REU) program provides support Summer. 6 credits. Prerequisites: a course various legal techniques. A case study that for undergraduates to pursue supervised, in vertebrate biology. S-U grades optional. requires extensive use of the laboratory’s independent research projects at the library and personnel is assigned. The week A special 3-week course offered at Shoals Marine Laboratory. Nine students Cornell’s Shoals Marine Laboratory (SML), concludes with a mock hearing. will be selected from a competitive, on an island off Portsmouth, N.H. For national pool to participate in the eight- NTRES 417 Wetland Resources more details or an application, consult the week summer program. For more Summer. 2 credits. Prerequisite: 1 year of SML office, G14 Stimson Hall. Daily lees, information and an application, please college-level biology. A special 1-week labs, and fieldwork for 3 weeks. SML contact the SML office, G14 Stimson Hall, course offered at Cornell’s Shoals Marine faculty. or view SML’s web site at: Laboratory (SML), on an island off Topics in marine vertebrate biology emphasiz­ www.sml.comell.edu Portsmouth, N.H. For more details or an ing laboratory studies, field collections or application, consult the SML office, G14 observations, and readings from the current [ARKEO 319 Archaeology Underwater Stimson Hall. Daily lees, labs, and literature. Topics covered include: systematics Summer. 2 credits. Prerequisites: recog­ fieldwork. SML faculty. of fishes of the Gulf of Maine; elasmobranch nized scuba certification and a medical An examination of coastal and adjacent physiology; interpretation of life history and examination required for students freshwater wetlands from historic, destruction, parameters from otolith microstructure; teleost engaging in underwater research; also and preservation perspectives, including fresh- skeletomuscular structure and function; open to non-divers. A special 1-week and salt-marsh ecology and management. population biology and the contemporary course offered at Cornell’s Shoals Marine Field trips to selected examples of the Gulf of Maine fishery; Mesozoic marine Laboratory (SML), on an island off wetlands under discussion and follow-up reptiles; the biology of sea turtles in cold Portsmouth, N.H. For more details or an laboratories emphasize successional features, water; coloniality in sea birds; avian adapta­ application, consult the SML office, G14 plant identification and classification, and tions to life at sea; evolution and systematics Stimson Hall. Daily lees, labs, and examination of the dominant insect and of marine mammals; diving physiology; and fieldwork for 1 week. Offered alternate vertebrate associations. ecology and conservation of existing marine years. Not offered 2001-2002. SML faculty. mammal populations. Dissection of vertebrate An introduction to the subject and a review of BIOEE 373 Biology of the Marine animals is a part of one or more laboratory this contemporary subdiscipline of archaeol­ Invertebrates sessions. ogy. The approach of the course is practical, Fall (but taken in the previous summer at the Shoals Marine Laboratory [SML]). 4 BIOSM 490 Topics in Marine Biology with a strong potential for actual on-site experience in search, site recognition, survey, credits. Limited to 30 students. Prerequi­ Summer. 1-2 credits. Prerequisite: 1 year and recording. The course also covers the site: 1 year of introductory biology for of introductory biology and permission of history and development of the subject, the majors. Permission of faculty required. instructors. S-U grades optional. A special legal aspects of underwater research, and the 2-week, full-time course. Daily and course offered at Cornell’s Shoals Marine worldwide potential of the field. Since any evening lectures, laboratories, and field Laboratory (SML), on an island off archaeological research project involves a work. Offered alternate years. Portsmouth, N.H. and on campus at great deal more than digging, the course C. D. Harvell, J. G. Morin, SML faculty. Cornell University. For more details, provides ample opportunities for those who An introduction to the biology and evolution consult the SML office, G14 Stimson Hall. are interested in the subject but are not divers of the major invertebrate phyla, concentrating Daily lees, labs, and fieldwork for 3 weeks. or sufficiently experienced in scuba.] on marine representatives. In addition to the SML faculty. evolution of form and function, lectures cover Seminar courses on selected topics in marine EAS 213 Marine and Coastal Geology aspects of ecology, behavior, physiology, biology. Students and faculty explore recent Summer. 2 credits. Prerequisite: an chemical ecology, and natural history of research through reading of the primary introductory course in geology or invertebrates. By being taught at the Shoals literature. The course may also include a lab permission of instructor. A special 1-week Marine Laboratory, students are exposed to a or field trips. Offered in spring semester for course offered at Cornell’s Shoals Marine wealth of marine and terrestrial invertebrates FACULTY 177

in their natural habitats. Regular field architecture, ship construction, marine Anderson, John M., Ph.D., New York U. Prof. excursions allow an excellent opportunity to engineering systems, and the physics of sail Emeritus, Molecular Biology and Genetics study freshly collected and in situ representa­ are taught from their bases in astronomy, Angert, Esther R., Ph.D., Indiana U. Asst. Prof., tives of most of the major phyla. mathematics, and physics. Provides the Microbiology BIOSM 366-372 SEA Semester theoretical foundation for the navigation, Bates, David M., Ph D., U. of California at Los seamanship, and engineering that students Angeles. Prof., Plant Biology (Bailey In cooperation with the Sea Education employ at sea. Hortorium) Association (SEA), the Shoals Marine Beyenbach, Klaus W., Ph.D., Washington State Laboratory offers a semester-length sequence U. Prof., Physiology/Veterinary Physiology) of courses designed to provide college Sea Component (six weeks) Bruns, Peter J., Ph.D., U. of Illinois. Prof. undergraduates with a thorough academic, Courses 369, 370 and 372 take place aboard Emeritus, Molecular Biology and Genetics scientific, and practical understanding of the the R/V W estw ard, a 125-foot steel auxiliary- Cade, Thomas J., Ph.D., U. of California at Los sea. This sequence is repeated approximately powered staysail schooner built in 1961, or Angeles. Prof. Emeritus, Ecology and once every two months throughout the year. the R/V Corwith Cramer, a 134-foot steel Evolutionary Biology Students spend the first half of SEA Semester auxiliary-powered brigantine built in 1987 for Calvo, Joseph M., Ph.D., Washington State U. (a six-week shore component) in Woods SEA. Both ships normally put to sea with a William T. Keeton Professor in Biological Hole, MA, receiving instruction in oceanogra­ ship’s company of 34. The professional staff of Sciences, Molecular Biology and Genetics phy, nautical science, and maritime studies. nine includes the captain, the chief scientist, Chabot, Brian F., Ph.D., Duke U. Prof., The second half of SEA Semester (a six-week three science watch officers, three deck watch Ecology and Evolutionary Biology sea component) is spent at sea aboard the R/V officers, an engineer, and a steward. In Clayton, Roderick K., Ph.D., California Inst, of W estw ard or the R/V Corwith Cramer. addition, one or more visiting investigators are Technology. Prof. Emeritus, Plant Biology Enrollment is open to both men and women frequently aboard. Up to 24 students round Crepet, William L., Ph.D., Yale U. Prof., Plant judged capable of benefiting from SEA out the complement. Biology (Bailey Hortorium)* semester; a student must have successfully BIOSM 369 SEA Practical Oceanography I Daniel, Louise J., Ph.D., Cornell U. Prof. completed at least one college-level 4 credits. Prerequisite: BIOSM 366. Emeritus, Molecular Biology and Genetics laboratory science course (or its equivalent) Theories and problems raised in the shore Davies, Peter J., Ph.D., U. of Reading in order to be admitted to SEA Semester or component are tested in the practice of (England). Prof., Plant Biology SEA Summer Session. No prior sailing oceanography at sea. Students are introduced Davis, Jerrold I., Ph.D., U. of Washington. experience Is necessary. Cornell students to the tools and techniques of the practicing Assoc. Prof., Plant Biology (Bailey enrolled in the SEA Semester must take the oceanographer. During lectures and watch Hortorium) entire sequence. standing, students are instructed in the Dhondt, Andre A., Ph.D., Ghent State U. For more information, contact the Shoals operation of basic oceanographic equipment; (Belgium). Edwin H. Morgens Professor of Marine Laboratory office, G14 Stimson Hall, or in the methodologies involved in the Ornithology, Ecology and Evolutionary call SEA directly at 800-552-3633. Program collection, reduction, and analysis of Biology/Laboratory of Ornithology costs are to be paid in place of regular Cornell oceanographic data; and in the attendant Dondero, Norman C., Ph.D., Cornell U. Prof. tuition and fees: tuition for the entire 17-credit operations of a sailing oceanographic research Emeritus, Microbiology SEA Semester, approximately $14,000 which vessel. Doyle, Jeffrey J., Ph.D., Indiana U. Prof,, Plant includes room and board at SEA. BIOSM 370 SEA Practical Biology (Bailey Hortorium) Dress, William J., Ph.D., Cornell U. Prof. Instructors for the SEA Semester include Oceanography II Emeritus, Plant Biology (Bailey Hortorium) faculty of the Sea Education Association and 4 credits. Prerequisites: BIOSM 368 and Eisner, Thomas, Ph.D., Harvard U. Jacob the Woods Hole Oceanographic Institution 369. Gould Schurman Professor, Neurobiology and others. Building on the experience of Practical and Behavior* Oceanography I, students assume increasing Emlen, Stephen T., Ph.D., U. of Michigan. responsibility for conducting oceanographic Shore Component (six weeks) Jacob Gould Schurman Professor, Neurobi­ research and overseeing operations of the BIOSM 366 SEA Introduction to ology and Behavior vessel. The individual student is ultimately Oceanography Feeny, Paul P., Ph.D., Oxford U. (England). responsible directly to the chief scientist and 3 credits. Prerequisite: concurrent Prof., Ecology and Evolutionary Biology/ the master of the vessel for the safe and enrollment in BIOSM 367 and 368. Entomology orderly conduct of research activities and A survey of the characteristics and processes Fitzpatrick, John W., Ph.D., Princeton U. Prof., related operations of the vessel. Each student of the global ocean. Oceanographic concepts Ecology and Evolutionary Biology/ undertakes an individual research project are introduced and developed from their Laboratory of Ornithology designed during the shore component. bases in biology, physics, chemistry, and Flecker, Alexander S., Ph.D., U. of Maryland. geology. Provides a broad background in BIOSM 372 SEA Practical Asst. Prof., Ecology and Evolutionary oceanography with special attention to areas Oceanography III Biology pertinent to the subsequent cruise. Guest Summer. 3 credits'. Prerequisites: BIOSM Fox, Thomas D., Ph.D., Harvard U. Prof., lecturers from the Woods Hole research 366, 367, and 368. Molecular Biology and Genetics community interpret current trends and Theories and problems raised in class are Fu, Jianhua, Ph.D., U. Pittsburgh. Asst. Prof., activities in this rapidly evolving field. tested in the practice of oceanography at sea. Molecular Biology and Genetics Students develop individual projects to be During lectures and watch standing, students carried out at sea. Ghiorse, William C., Ph.D. Rensselaer are instructed in the operation of basic Polytechnic Inst. Prof., Microbiology BIOSM 367 SEA Introduction to Maritime oceanographic equipment, in the methodolo­ Gibson, Jane, Ph.D., U. of London (England). Studies gies involved in the collection, analysis, and Prof. Emeritus, Molecular Biology and 3 credits. Prerequisite: concurrent reduction of oceanographic data, and in the Genetics enrollment in BIOSM 366 and 368. attendant operations of sailing an oceano­ Goldberg, Michael L., Ph.D., Stanford U. Prof., An interdisciplinary consideration of our graphic research vessel. Group research Molecular Biology and Genetics relationship with the marine environment. projects are completed. Hanson, Maureen R., Ph.D., Harvard U. Prof., Covers the elements of maritime history, law, Molecular Biology and Genetics/Liberty literature, and art necessary to appreciate our Hyde, Prof., Plant Bioiogy marine heritage and to understand the Harrison, Richard G., Ph.D., Cornell U. Prof., political and economic problems of contem­ FACULTY ROSTER Ecology and Evolutionary Biology porary maritime affairs. Harris-Warrick, Ronald M., Ph.D., Stanford U. New York State College of Agriculture Prof., Neurobiology and Behavior BIOSM 368 SEA Introduction to Nautical Harvell, C. Drew, Ph.D., U. of Washington. Science and Life Sciences Prof., Ecology and Evolutionary Biology Adler, Kraig K., Ph.D., U. of Michigan. Prof., 3 credits. Prerequisites: concurrent Hay, Anthony, Ph.D., U. of California. Asst. Neurobiology and Behavior enrollment in BIOSM 366 and 367. Prof., Microbiology Alani, Eric E., Ph.D., Harvard U. Assoc. Prof., An introduction to the technologies of Helmann, John D., Ph.D., U. of California at Molecular Biology and Genetics operation at sea. The concepts of navigation Berkeley. Assoc. Prof., Microbiology (piloting, celestial and electronic), naval BIOLOGICAL SCIENCES - 2001-2002

Henry, Susan, Ph.D., U. of California Berkley. Rose, Jocelyn, Ph.D., U. of California at Davis. Comstock, Jonathan P., Adjunct Assoc. Prof., Prof., Molecular Biology and Genetics and Asst. Prof., Plant Biology Boyce Thompson Institute/Ecology and Dean CALS Russell, James B., Ph.D., U. of California at Evolutionary Biology Hopkins, Carl D., Ph.D., Rockefeller U. Prof., Davis. Prof., Microbiology Foote, Robert H., Jacob Gould Schurman Prof. Neurobiology and Behavior Seeley, Jr., Harry W., Ph.D., Cornell U. Prof. Emeritus, Animal Science/Physiology Howarth, Robert W., Ph.D., Massachusetts Emeritus, Microbiology Giovannoni, James G., Adjunct Asst. Prof., Inst, of Technology/Woods Hole Oceano­ Shalloway, David I., Ph.D., Massachusetts Inst, USDA Science and Education Administra- graphic Institution. David R. Atkinson of Technology. Greater Philadelphia Prof., tion/Plant Biology Professor in Ecology and Environmental Molecular Biology and Genetics Hodge, Kathie, Asst. Prof. Plant Pathology/ Biology, Ecology and Evolutionary Biology/ Shapleigh, James P., Ph.D., U. of Georgia. Plant Biology (Bailey Hortorium) Earth and Atmospheric Sciences Asst. Prof., Microbiology Kochian, Leon V., Adjunct Prof., USDA Ingram, John W., Ph.D., U. of California at Spanswick, Roger M., Ph.D., U. of Edinburgh Science and Education Administration/Plant Berkeley. Prof. Emeritus, Plant Biology (Scotland). Prof., Plant Biology Biology (Bailey Hortorium) Thiel, Daniel J., Ph.D., Cornell U. Asst. Prof., Korf, Richard P., Prof. Emeritus, Plant Jagendoif, Andre T., Ph D., Yale U. Liberty Molecular Biology and Genetics Pathology/Plant Biology (Bailey Hortorium) Hyde Bailey Professor of Plant Physiology Tye, Bik-Kwoon, Ph.D., Massachusetts Inst, of Kresovich, Stephen, Prof., Plant Breeding/ Emeritus, Plant Biology Technology. Prof., Molecular Biology and Plant Biology Kemphues, Kenneth J., Ph.D., Indiana U. Genetics Liebherr, James K., Assoc. Prof., Entomology/ Prof., Molecular Biology and Genetics Uhl, Charles H., Ph.D., Cornell U. Prof. Plant Biology (Bailey Hortorium) Kingsbury, John M., Ph.D., Harvard U. Prof. Emeritus, Plant Biology Mason, Hugh S., Adjunct Asst. Prof., Boyce Emeritus, Plant Biology Uhl, Natalie W., Ph.D., Cornell U. Prof. Thompson Institute/Plant Biology Kraus, Lee, Ph.D., U. of Illinois. Asst. Prof., Emeritus, Plant Biology (Bailey Hortorium) McClure, Polley A., Prof., Information Molecular Biology and Genetics Van Wijk, Klaas J., Ph.D., Groningen U., The Technologies/Ecology and Evolutionary Lis, John T., Ph.D., Brandeis U. Prof., Netherlands. Asst. Prof., Plant Biology Biology Molecular Biology and Genetics Vogt, Volker M., Ph.D., Harvard U. Prof., McCouch, Susan R., Assoc. Prof., Plant Loew, Ellis R., Ph.D., U. of California at Los Molecular Biology and Genetics Breeding/Plant Biology Angeles. Prof., Physiology/Veterinary Walcott, Charles, Ph.D., Cornell U. Prof., Richmond, Milo E., Assoc. Prof., USDI Fish Physiology! Neurobiology and Behavior and Wildlife Service/Natural Resources/ Lovette, IrbyJ., Ph.D., U. of Pennsylvania. Wayne, Randy O., Ph.D., U. of Massachusetts. Ecology and Evolutionary Biology Asst. Prof., Ecology and Evolutionary Assoc. Prof., Plant Biology Rossman, Michael J., Adjunct Prof., Purdue U./ Biology/Laboratory of Ornithology Winans, Stephen C., Ph.D., Massachusetts Inst. Molecular Biology and Genetics Luckow, Melissa A., Ph.D., U. of Texas at of Technology. Assoc. Prof., Microbiology Stem, David B., Adjunct Prof., Boyce Austin. Assoc. Prof., Plant Biology (Bailey Winkler, David W., Ph.D., U. of California at Thompson Institute/Plant Biology Hortorium) Berkeley. Assoc. Prof., Ecology and Tanksley, Stephen, Prof., Plant Breeding/ MacDonald, Russell E., Ph.D., U. of Michigan. Evolutionary Biology Liberty Hyde Bailey Prof., Plant Biology Prof. Emeritus, Molecular Biology and Wu, Ray, Ph.D., U. of Pennsylvania. Prof., Thompson, John F., Adjunct Prof., USDA Genetics Molecular Biology and Genetics Science and Education Administration/Plant MacIntyre, Ross J., Ph.D., Johns Hopkins U. Zahler, Stanley A., Ph.D., U. of Chicago. Prof. Biology Prof., Molecular Biology and Genetics Emeritus, Molecular Biology and Genetics Vehrencamp, Sandra, Ph.D., Cornell U. Prof., Madsen, Eugene L., Ph.D., Cornell U. Asst. Zinder, Stephen H., Ph.D., U. of Wisconsin. Neurobiology and Behavior/Library of Prof., Microbiology Prof., Microbiology Natural Sounds Marks, Peter L., Ph.D., Yale U. Prof., Ecology Wheeler, Quentin D., Prof., Entomology/Plant and Evolutionary Biology Other Teaching Personnel Biology (Bailey Hortorium) McCune, Amy R., Ph.D., Yale U. Assoc. Prof., Baustian, Mark D., Ph.D., Cornell U. Lecturer, Ecology and Evolutionary Biology Physiology College of Arts and Sciences Morin, James G., Ph.D., Harvard U. Prof., Blankenship, James E., M.S., Cornell U. Sr. Adkins-Regan, Elizabeth, Ph D., U. of Ecology and Evolutionary Biology Lecturer, Molecular Biology and Genetics Pennsylvania. Prof., Neurobiology and Mortlock, Robert P., Ph.D., U. of Illinois. Prof. Calvo, Rita A., Ph.D., Cornell U. Sr. Lecturer, Behavior/Psychology Emeritus, Microbiology Molecular Biology and Genetics Aquadro, Charles F., Ph.D., U. of Georgia. Nasrallah, June B., Ph.D., Cornell U. Prof., Ecklund, P. Richard, Ph.D., Oregon State U. Prof., Molecular Biology and Genetics/ Plant Biology Sr. Lecturer, Neurobiology and Behavior Ecology and Evolutionary Biology Nasrallah, Mikhail E., Ph.D., Cornell U. Prof., Ely, Susan, Ph.D., Tufts U. Lecturer, Molecular Bass, Andrew H., Ph.D., U. of Michigan. Prof., Plant Biology Biology and Genetics Neurobiology and Behavior Naylor, Harry B., Ph.D., Cornell U. Prof. Glase, Jon C., Ph.D., Cornell U. Sr. Lecturer, Blackler, Antonie W., Ph.D., U. of London Emeritus, Microbiology Neurobiology and Behavior (England). Prof., Molecular Biology and Niklas, Karl J., Ph.D., U. of Illinois. Prof., Plant Land, Bruce, Ph.D., Cornell U. Sr. Lecturer, Genetics Biology Neurobiology and Behavior* Booker, Ronald, Ph.D., Princeton U. Assoc. Nixon, Kevin C., Ph.D., U. of Texas at Austin. McFadden, Carol H., Ph.D., Cornell U. Sr. Prof., Neurobiology and Behavior Assoc. Prof., Plant Biology (Bailey Lecturer, Physiology Bretscher, Anthony P., Ph D., Leeds U. Hortorium) Merkel, Susan, M.S., Cornell U. Lecturer, (England). Prof., Molecular Biology and Owens, Thomas G., Ph D., Cornell U. Assoc. Microbiology Genetics Prof., Plant Biology Nivison, Helen T., Ph.D., U. of California at Brown, William J., Ph.D., U. of Texas Health Paolillo, Dominick J., Jr., Ph.D., U. of Davis. Lecturer, Molecular Biology and Science Center at Dallas. Prof., Molecular California at Davis. Prof. Emeritus, Plant Genetics Biology and Genetics Biology Rehkugler, Carole M., M.S., Cornell U. Sr. Capranica, Robert R., Sc.D., Massachusetts Parthasarathy, Mandayam V., Ph.D., Cornell U. Lecturer, Microbiology Inst, of Technology. Prof. Emeritus, Prof., Plant Biology! Shulman, Myra J., Ph.D., U. of Washington. Sr. Neurobiology and Behavior Quaroni, Andrea, Ph D., U. of Pavia (Italy). Res. Assoc., Ecology and Evolutionary Chen, Rey-Huei, Ph.D., Harvard U. Asst. Prof., Prof., Physiology/Veterinary Physiology! Biology Molecular Biology and Genetics Reeve, H. Kern, Ph D., Cornell U. Assoc. Prof., Silva, Thomas, Ph.D., Cornell U. Lecturer, Deitcher, David, Ph.D., Harvard Med. School. Neurobiology and Behavior Plant Biology Asst. Prof., Neurobiology and Behavior Ellner, Stephen P., Ph.D., Cornell U. Prof., Roberts, Jeffrey W., Ph.D., Harvard U. Robert Joint Appointees J. Appel Professor of Cellular and Molecular Ecology and Evolutionary Biology Biology, Molecular Biology and Genetics Bloom, Stephen E., Assoc. Prof., Poultry and Feigenson, Gerald W., Ph.D., California Inst, Rodriguez, Eloy, Ph D., U. of Texas. Prof., Avian Sciences/Biological Sciences of Technology. Prof., Molecular Biology Plant Biology (Bailey Hortorium) Bradbury, Jack, Ph.D., Rockefeller. Prof., and Genetics Root, Richard B., Ph D., U. of California at Neurobiology and Behavior/Library of Finlay, Barbara, Ph.D., Massachusetts Inst, of Berkeley. Prof., Ecology and Evolutionary Natural Sounds Tech. Prof., Neurobiology and Behavior/ Biology/Entomology Brutnell, Thomas, Prof., Plant Breeding/Plant Psychology Biology FACULTY 179

Geber, Monica A., Ph.D., U. of Utah. Assoc. Other Teaching Personnel ‘Joint appointment with the College of Arts Prof., Ecology and Evolutionary Biology Berry, John, Ph.D., Cornell U. Post Doc, Plant and Sciences. Gibson, Quentin H., Ph.D./D.Sc., Queen’s U. Biology (Bailey Hortorium) (Northern Ireland). Greater Philadelphia Uoint appointment with the College of Eberhard, Carolyn, Ph.D., Boston U. Sr. Veterinary Medicine. Professor Emeritus in Biological Sciences, Lecturer, Plant Biology Molecular Biology and Genetics Johnson, Bruce R., Ph.D., Boston U. Sr. (Joint appointment with the College of Greene, Harry W., Ph.D., U. of Tennessee. Lecturer, Neurobiology and Behavior Agriculture and Life Sciences. Prof., Ecology and Evolutionary Biology Sjjoint appointment with the College of Hairston, Nelson G., Jr., Ph.D., U. of Washing­ Joint Appointees Engineering. ton. Frank H. T. Rhodes Professor of Levin, Simon A., Adjunct Prof., Princeton U./ Environmental Science, Ecology and Ecology and Evolutionary Biologyf Evolutionary Biology Likens, Gene E., Adjunct Prof., Institute of Halpern, Bruce P., Ph D., Brown U. Prof., Ecosystem Studies/Ecology and Evolution­ Neurobiology and Behavior/Psychology ary Biology Hedin, Lars O., Ph.D., Yale U. Assoc. Prof., Ecology and Evolutionary Biology College of Veterinary Medicine Heppel, Leon A., Ph D., U. of California at Gasteiger, Edgar L., Ph.D., U. of Minnesota. I Berkeley. Prof. Emeritus, Molecular Biology Prof. Emeritus, Physiology/Veterinary and Genetics Physiology Hess, George P., Ph.D., U. of California at Kotlikoff, Michael I., Ph.D., U. California at Berkeley. Prof., Molecular Biology and Davis. Prof., Veterinary Physiology/ Genetics Physiology Hinkle, Peter C., Ph.D., New York U. Prof., Tapper, Daniel N., Ph.D., Cornell U. Prof. Molecular Biology and Genetics Emeritus, Physiology/Veterinary Physiology Howland, Howard C., Ph.D., Cornell U. Prof., Wasserman, Robert H., Ph.D., Cornell U. Neurobiology and Behavior/Biomedical James Law Prof. Emeritus, Physiology/ Sciences Veterinary Physiology/Nutritional Sciences'^ Hoy, Ronald R., Ph.D., Stanford U. Prof., Neurobiology and Behavior Other Teaching Personnel Huffaker, Tim C., Ph D., Massachusetts Inst, of Concannon, Patrick W., Ph D., Cornell U. Sr. Technology. Assoc. Prof., Molecular Biology Res. Assoc. Emeritus, Veterinary Physiol- and Genetics ogy/Physiology Kennedy, Kenneth A. R., Ph.D., U. of California at Berkeley. Prof., Ecology and Joint Appointees Evolutionary Biology Fortune, Joanne E., Ph.D., Cornell U. Prof., Leonard, Samuel L., Ph.D., U. of Wisconsin. Veterinary Physiology/Physiology Prof. Emeritus, Molecular Biology and Gilmour, Robert F., Ph.D., SUNY Upstate Genetics Medical Center. Assoc. Prof., Veterinary Linster, Christine, Ph.D., Pierre and Marie Physiology/Physiology Curie U. Asst. Prof., Neurobiology and Houpt, Katherine A., Prof., Veterinary Behavior Physiology/Physiology McCobb, David, Ph.D., U. of Iowa. Asst. Prof., Houpt, T. Richard, Prof., Veterinary Physiol- Neurobiology and Behavior ogy/Physioldgy MacDonald, June M. Fessenden, Ph.D., Tufts Nathanielsz, Peter W., James Law Prof., U. Assoc. Prof. Emeritus, Molecular Biology Veterinary Physiology/Physiology and Genetics/Program on Science, Robertshaw, David, Ph.D., Glasgow U. Technology, and Society (Scotland). Prof., Veterinary Physiology/ McFarland, William N., Ph.D., U. of California Physiology at Los Angeles. Prof. Emeritus, Ecology and Wootton, John F., Prof., Veterinary Physiol­ Evolutionary Biology ogy/Physiology Nicholson, Linda, Ph.D., Florida State U. Asst. Prof., Molecular Biology and Genetics College of Engineering Podleski, Thomas R., Ph.D., Columbia U. Prof. Emeritus, Neurobiology and Behavior Joint Appointees Power, Alison G., Ph.D., U. of Washington. Cisne, John L., Assoc. Prof., Geological Prof., Ecology and Evolutionary Biology/ Sciences/Biological Sciences Science and Technology Studies Webb, Watt W., Prof., Applied and Engineer­ Provine, William B., Ph.D., U. of Chicago. ing Physics/Biological Sciences Charles A. Alexander Professor of Biological Sciences, Ecology and Evolutionary Biological Sciences Biology/History Joint Appointees Seeley, Thomas D., Ph.D., Harvard U. Prof., Neurobiology and Behavior Snedeker, Suzanne M., Asst. Prof., Center for Sherman, Paul W., Ph.D., U. of Michigan. the Environment/Biological Sciences Prof., Neurobiology and Behavior Division of Nutritional Sciences Turgeon, Robert, Ph.D., Carleton U. (Canada). Prof., Plant Biology Joint Appointees Wallace, Bruce, Ph.D., Columbia U. Prof. Arion, William J., Prof., Nutritional Sciences/ Emeritus, Molecular Biology and Genetics Molecular Biology and Genetics Whitlock, Kathleen E., Ph.D., U. Washington Seattle. Asst. Prof., Molecular Biology and Bensadoun, Andre, Prof., Nutritional Sciences/ Physiology Genetics Kazarinoff, Michael N., Assoc. Prof., Nutri­ Wilson, David B., Ph.D., Stanford U. Prof., Biochemistry, Molecular Biology and tional Sciences/Molecular Biology and Genetics Genetics Wright, Lemuel D., Ph.D., Oregon State Coll. Wolfner, Mariana F., Ph.D., Stanford U. Prof., Prof. Emeritus, Nutritional Sciences/ Molecular Biology and Genetics Molecular Biology and Genetics Zamudio, Kelly R., Ph.D., U. of Washington. Asst. Prof., Ecology and Evolutionary Biology