Bothalia 31,1 (2001) 41

BAKER, J.G. 1906. Anthericum (Phlangiunt) recurvifolium Baker the description of three new species. Nordic Journal of Botany (Liliaeeae-Asphodeleae). In Diagnoses Africanae XIV. Kew 13: 257-280. Bulletin 1906: 15-30. OBERMEYER, A. A. 1962. A revision of the South African species of An­ KATIVU, S. 1993. In S. Kativu & I. Nordal, New combinations of thericum, Chlorophytum and Trachyandra. Bothalia 7: 669-767. African species in the genus Chlorophytum (Anthericaceae). VON POELLNITZ, K. 1946. Die Chlorophytum-aiten Tanganyikas. Nordic Journal of Botany 13: 59-65. Portugaliae acta biologica, ser. B, 1: 255-383. NORDAL, I., ERIKSEN, T.E. & FOSBY, M. 1990. Studies in the generic delimitations in Anthericaceae. Mitteilungen aus dem C. ARCHER* and S. KATIVU** Institut fur Allgemeine Botanik Hamburg 23(b): 535-559. NORDAL, I , KATIVU, S. & POULSEN, A.D. 1997. Anthericaceae. In * National Botanical Institute, Private Bag X I01, (MM) 1 Pretoria. R.M. Polhill, Flora o f tropical East Africa. Balkema, Rotterdam. ** Department of Biological Sciences, University of Zimbabwe, P.O. NORDAL, I. & THULIN, M. 1993. Synopsis of Anthericum and Box MP 167, Mount Pleasant, Harare, Zimbabwe. Chlorophytum (Anthericaceae) in the Horn of Africa, including MS. received: 2000-08-08.

ASTERACEAE

A NEW SPECIES OF FROM WESTERN CAPE. SOUTH AFRICA

Oedera epaleacea Beyers, sp. nov. Oederae fove- 0.3-0.6 mm, glabrous or sparsely pilose, may be scat­ olatae affinis sed receptaculo epaleaceo floribus disci tered with ± stalked glands; pappus crown-like, of ± con­ femineo-sterilibus stylo modo breviter fisso differt. nate scales, up to 0.6 mm long. Figure 9.

TYPE.—Western Cape, 3219 (Wuppertal): Swartrug- Diagnostic characters and relationships gens, on Farm Zeekoe-gat, southwest of the farm road about 500 m from the turnoff on the road to Katbakkies Oedera foveolata (Bremer) Anderb. & Bremer and O. Pass, 1 030 m, (-DC), 5-09-2000, Beyers 272 (NBG, sedifolia (DC.) Anderb. & Bremer are among those holo.; K, PRE, UPS, iso.). species, treated in the taxonomic revision of L'Her. by Bremer (1976), which were transferred to Erect, moderately to densely branched, rounded Oedera by Anderberg & Bremer (1991). O. epaleacea is shrub, up to 0.6 m tall, with a stout trunk up to 50 mm related to the above two species, which are recognized diam. Branches ascending-erect, glabrous, leafy, becom­ by their glandular-punctate leaves. As in O. epaleacea, ing nude and marked with leaf scars. Leaves alternate, O. foveolata, a species only known from Karoo Poort, spreading, sessile, entire, midribbed, narrowly oblong, the leaves are distinctly glandular-punctate and foveo­ 4-12 x 0.8-1.5 mm, glabrous, densely and distinctly late, and the cypselas of the ray florets are densely pilose. glandular-punctate and foveolate, mucronulate; semi- However, in O. epaleacea the receptacle is epaleate and terete in cross section. Capitula solitary, sessile, termi­ the disc florets are female sterile. O. epaleacea grows nal. heterogamous, radiate. Involucre campanulate, 4-5 intermingled with O. sedifolia, one of the more common mm wide. Involucral bracts 5- or 6-seriate, 36-47, species in the genus, but can easily be distinguished from imbricate, firm and hard basally, scarious apically, it by the glabrous branches and leaves and the female midribbed, glabrous; outer ovate, inner gradually longer, sterile disc florets. The leaves of O. epaleacea, when narrowly elliptic and apically brown, innermost narrow­ crushed have a sulphury odour. In O. sedifolia the ly obovate or narrowly oblong with a spreading, brown branches and leaves are tomentose becoming glabrous apical limb, up to 10.3 x 1.7 mm, acute. Receptacle later, the disc florets are perfect and leaves when crushed slightly convex, shortly squamose, epaleate. Ray florets give off a resinous odour. female, fertile, 13-18; tube cylindrical, 2-3 mm long, with ± stalked glands; lamina spreading, narrowly obo­ vate, 6-10 x 1.3- 2.2 mm, yellow with a dorsal, brown Distribution and ecology stripe, 4-veined, apically minutely 3-lobed; style terete, bifid. 2.5-4.0 mm long; style branches erect-spreading, Oedera epaleacea is known only from the Farm linear, 0.7- 1.5 mm long, glabrous, stylopodium not dis­ Zeekoe-gat in the Swartruggens area (Figure 10). It tinct; cypselas almost terete or angular, narrowly elliptic, occurs in arid fynbos where it grows in crevices between 1.8- 2.6 x 0.5- 0.7 mm, densely pilose; pappus crown­ rock sheets, in sandstone-derived sandy soils, in full or like. of ± connate scales, up to 0.6 mm long. Disc florets partial sun. Flowering specimens seen from June to 38^2, female sterile; corolla 3.7—5.5 x 0.6—0.9 mm, September. divided into a lower cylindrical tube with scattered, ± stalked glands on the outside and upper narrowly cyathi- Etymology form, glabrous limb; corolla lobes 5, spreading, ovate- triangular, 0.3- 0.7 mm long, dorsally gland-dotted; style The specific epithet refers to the epaleate receptacle. terete, 2.5- 5.3 mm long, shallowly cleft, style branches up to 0.3 mm long, apically rounded and papillate; sty­ Other specimens examined lopodium distinct; anthers linear, 1.4-2.2 mm long, with sterile, tlat, ovate apical appendage and sterile, flat, sub­ WESTERN CAPE.—3219 (Wuppertal): Zeekoe-gat 137, west of ulate, entire tails, 0.2- 0.4 mm long; cypselas almost Rict River, in ravine, 16-06-2000, Stobie 2 (NBG); ibid., on hill slope, terete, narrowly elliptic to narrowly oblong, 1.5-2.7 x 22-07-2000, Stobie sub NBG170457 (NBG). 42 Bothalia 31,1 (2001)

FIGURE 9.— Oedera epaleacea, Beyers 272 (NBG). A, portion of ; B, leaf (abaxial view and cross section). C-E, involucral bracts: C, outer; D, middle; E, innermost. F, disc floret; G, stamen of disc floret; H, style of disc floret; I, ray floret; J, cypsela of ray floret; K, style of ray floret.

ACKNOWLEDGEMENTS

HEIGHT ABOVE SEA LEVEL Over 1500 m I would like to thank Mrs Mary Stobie, a keen ama­ mm 900 -1500 m teur botanist, for bringing the material in for naming and S 300 - 900 m also for accompanying me to their farm to collect the I I Under 300 m type material, Prof. Kare Bremer for his expert advice, 20 0 20 40 60 80 100 km Dr Ted Oliver for the Latin diagnosis, and Mrs Inge Oliver for the line drawings.

REFERENCES

ANDERBERG, A.A. & BREMER, K. 1991. Parsimony analysis and cladistic reclassification of the Relhania generic group (Aster- aceae-). Annals of the Missouri Botanical Garden 78: 1061-1072. BREMER, K. 1976. The genus Relhania (Compositae). Opera Botan- ica 40: 1-85.

J.B.P. BEYERS*

* Compton Herbarium, National Botanical Institute, Private Bag X7, 7735 Claremont, Cape Town. FIGURE 10.—Geographical distribution of Oedera epaleacea. MS. received: 2000-10-04.