Monograph of the Little Slit Shells

VOLUME 1 Introduction

VOLUME 2 Larocheidae Depressizonidae Sutilizonidae Temnocinclidae

Daniel L. Geiger Introduction Monograph of the Little Slit Shells

Daniel L. Geiger

Santa Barbara Museum of Natural History Monographs Number 7 Studies in Biodiversity Number 5

© 2012 Santa Barbara Museum of Natural History 2559 Puesta del Sol Road Santa Barbara, CA 93105-2936 USA http://www.sbnature.org

Library of Congress Cataloging-in-Publication Data

Geiger, Daniel L. Monograph of the little slit shells / Daniel L. Geiger. -- 1st ed. p. cm. -- (Studies in biodiversity ; no. 5) Includes bibliographical references and index. ISBN 978-0-936494-45-6 (alk. paper) 1. Scissurellidae. 2. Archaeogastropoda. I. Title. QL430.5.S25G45 2012 594'.32--dc23 2012015342

ISBN (13) 978-0-936494-45-6

First Edition

© All rights reserved. This book may not be reproduced in whole or in part for any purpose whatsoever, without written permission from the publisher (Santa Barbara Museum of Natural History).

Entire layout and design by Daniel L. Geiger. Images by Daniel L. Geiger, unless otherwise noted. Typeset in Optima 18 point (sans serif headers) and Arno Pro 11 point (most text) 10 point (specimen records), and 9 point (synonymies). Printed and bound in China by Everbest Printing Company through FCI Print Group.

2 D. L. Geiger: Monograph of the Little Slit Shells About the author

Daniel L. Geiger grew up in Basel, Switzerland, where he started shell collecting in Gymnasium (high school). He soon specialized in abalone (Haliotidae) as an active

member of the Swiss Société internationale de Conchyliologie and the German Club Introduction Conchylia. He received his Diplom in organismal biology with a Nebenfach (minor) in paleontology from the University of Basel, Switzerland; thesis work was carried out as a visiting post-graduate student at the University College of Swansea, Wales, Great Britain, co-advised by David G. Senn and John Ryland. Subsequently he earned his Ph.D. in marine biology from the University of Southern California, Los Angles (USC), co-advised by Russel L. Zimmer (USC)

and James H. McLean (Los Angeles County Museum of Natural History: LACM). Scissurellidae His dissertation on systematics and evolution of abalone received the 2000 Annual Award by the Malacological Society of London. After a W. M. Keck post-doctoral fellowship in molecular systematics at LACM, and teaching appointments as an adjunct assistant professor at USC, he took a position at the Santa Barbara Museum of Natural History in 2005, where he now is curator of malacology. Geiger is interested in global systematic assessments of basal marine snails,

with special emphasis on small-sized groups. He has published two monographs Scissurellidae on abalone (Geiger & Poppe, 2000; Geiger & Owen, 2012) and two key works on micromollusks (Geiger et al., 2007; Geiger & Ruthensteiner, 2008), amongst many other contributions. Field work has taken him from boreal St. Kilda to tropical Papua New Guinea and anywhere in between. He is senior subject editor of with Zootaxa, associate editor of Molluscan Research, member of the editorial board of Malacologia, and member of the IUCN

Mollusc Specialist Group. Scissurellidae Sukashitrochus When not spending time on snails, he enjoys photography, gardening with California native plants, growing orchids, playing the viola d’amore, and spending quality time in the kitchen and above all with LACM ichthyologist Christine E. Thacker. Satondella Scissurellidae Scissurellidae Incisura Scissurellidae Scissurellidae

Fore-edge quick index

The fore-edge of the volume shows edge markings indicating the position of the Coronadoa various parts. It permits the reader to quickly find the beginning of the desired Scissurellidae section. D. L. Geiger: Monograph of the Little Slit Shells 3 Introduction Table of contents: Volume 1

About the author ...... 3 Fore-edge quick index ...... 3 Table of contents ...... 4 Acknowledgements ...... 13 Introduction ...... 15 Why little slit shells?...... 15 History ...... 16 Discovery, composition ...... 16 Genera included ...... 18 Anatomical characters ...... 20 Phylogeny, classification ...... 20 Small size ...... 22 Preservation ...... 23 Taxon concepts ...... 25 ...... 25 Genera ...... 25 Families ...... 26 Species groups ...... 26 Applied ...... 27 finlayi ...... 27 Anatoma pseudoequatoria ...... 28 Taxonomic actions taken here ...... 29 Organization of the descriptions ...... 30 Species name ...... 30 Chresonymy ...... 30 Type material ...... 31 Type species ...... 31 Other genera/species ...... 31 Species removed from ...... 31 Type locality ...... 31 Etymology ...... 31 Description ...... 32 Comparisons ...... 32 Distribution ...... 32 Geologic Range ...... 32 Fossil...... 32 Specimen records ...... 32 Literature records ...... 33 Remarks ...... 33 Distribution maps ...... 33 Collecting and curation ...... 33 Collecting ...... 33 Curation ...... 36 Sampling effort ...... 36 Government regulations ...... 37 Specimen preparation ...... 38

4 D. L. Geiger: Monograph of the Little Slit Shells Type specimens ...... 38 Shells ...... 38 Radula ...... 39 External anatomy ...... 39 Photography and digital imaging ...... 40

SEM ...... 40 Introduction Light microscopy ...... 40 SLR photography and digital imaging ...... 40 Distribution maps ...... 41 Characters ...... 41 Shell shape ...... 41 Protoconch ...... 41 Protoconch ...... 42 Protoconch varix ...... 43 Shape of protoconch margin ...... 43 Teleoconch I/II ...... 43 Shoulder ...... 41 Base ...... 43 Shell sculpture ...... 43 ...... 44 Umbilical pouch ...... 45 ...... 45 Slit and selenizone ...... 45 Start of selenizone ...... 45 Suture ...... 45 Sutsel ...... 46 ...... 46 Radula ...... 46 Rachidian tooth ...... 46 Lateral teeth ...... 44 Marginal teeth ...... 44 External anatomy ...... 47 Discussion of characters ...... 47 Protoconch ...... 47 Protoconch in systematics ...... 48 Teleoconch ...... 49 Umbilicus ...... 50 Slit and selenizone ...... 50 Coloration ...... 52 Radula ...... 52 Lateral tooth 5 ...... 53 Radula variability ...... 54 Operculum ...... 55 External anatomy ...... 57 Internal anatomy ...... 57 Aberrant specimens ...... 58 Occasional confusions ...... 59 Distribution and diversity patterns ...... 59 Bathymetric distribution ...... 59

D. L. Geiger: Monograph of the Little Slit Shells 5 Geographic distribution ...... 57

Introduction Diversity patterns ...... 61 Fossils ...... 63 Biology ...... 63 Food and feeding ...... 66 Habitat ...... 66 Reproduction, life history ...... 66 Predators ...... 67 Symbionts ...... 67 Conservation ...... 68 Supplementary information ...... 68 Glossary of terms ...... 69 Collection abbreviations ...... 73 Abbreviations ...... 76 Voucher repository ...... 76 Thumbnail identification ...... 77 List of species by global regions...... 89 Definitions of regions ...... 89 1–Mediterranean ...... 90 2–NE Atlantic ...... 90 2–West Africa ...... 90 4–South Africa ...... 90 5–Indian Ocean ...... 91 6–Indo-Malayan Archipelago ...... 91 7–Temperate Australia ...... 93 9–New Zealand ...... 93 10–NW Pacific ...... 94 10–Central Pacific ...... 94 11–NE Pacific ...... 95 12–Panamic ...... 95 13–Peruvian ...... 96 14–Magellanic ...... 96 15–Caribbean ...... 96 17–NW Atlantic ...... 96 18–Arctic ...... 97 19–Antarctic ...... 97 Indices of species and genus-level taxa...... 98 Index of species-level taxa ...... 99 Index of genus-level taxa ...... 111 Unavailable names ...... 112 Scissurella monfortii Orbigny, 1824 [unavailable hypothetical name] ...... 112 Scissurella tenera Monterosato, 1878 [unavailable name, nomen dubium] ...... 112 Scissurella crispata subangulata Monterosato, 1878 [unavailable name] ...... 113 Scissurella dorbignyi Scacchi, 1836 [unavailable junior primary homonym] ...... 113 Scissurella borealis “Sowerby” [nomen nudum] ...... 114 Scissurella rubiginosa “Hutton” nomen[ nudum] ...... 114 Schismope californica “Bartsch” [nomen nudum] ...... 114 Schismope trivialis auct. [Sinezona beddomei] ...... 115 Scissurella mediteranea auct./Monterosato ...... 115

6 D. L. Geiger: Monograph of the Little Slit Shells Scissurella athleenae Pilsbry & McGinty, ms. [unavailable, unpublished, nomen nudum] ...... 116 Scissurella costata var. semicostata auct. (non Burnay & Rolán, 1990) [unpublished] ...... 116 Praescissurella indopacifica Lozouet, 1986 [unavailable, unpublished, nomen nudum] ...... 116 Scissurella (Sukashitrochus) subitana Lozouet, 1986 [unavailable, unpublished] ...... 116 Anatoma alba Bandel, 1998 [nomen nudum] ...... 117

Pagodella Bandel, 1998 [nomen nudum, junior homonym] ...... 117 Introduction Scissurella reunionensis auct. [nomen nudum] ...... 117 Species erroneously described/classified in Scissurellidae...... 118 Anatomus indicus Montfort, 1810 [Annelida: Polychaeta: Serpulidae: Spirorbis sp.] ...... 118 Pleurotomaria sismondai Goldfuss, 1844 † [Pleurotomariidae] ...... 119 Leptomaria niloticiformis (von Schlotheim, 1820) † [Pleurotomariidae] ...... 120 Schismope lacuniformis Watson, 1886 [Macromphalina: Vanikoridae?] ...... 120 Pseudorbis caliana (Dall, 1919) [Skeneidae] ...... 122 Zardinitrochus Bandel, 1998 † ...... 123 Temnotropis Laube, 1869 † ...... 123 “Temnotropis” suessi Klipstein in Kittl, 1891 † incertae[ sedis: Fossariidae?] ...... 123 Scissurella minuta Turton, 1932 [nomen dubium] ...... 124 Atlanta peronii Lesueur, 1817 [Heteropoda: Atlantidae] ...... 125 Color Plate 1. Colorations of Anatomidae shells and ...... 126 Color Plate 2. Colorations of scissurellid shells and animals ...... 128 Color Plate 3. Live Scissurellidae from Madagascar and the Philippines ...... 130 Color Plate 4. Live scissurellids from Vanuatu ...... 132 Color Plate 5. Histology ...... 134 Scissurellidae Gray, 1847 ...... 136 Scissurella d’Orbigny, 1824 ...... 137 Scissurella alexandrei Montouchet, 1972 [nomen dubium] ...... 142 Scissurella alto Geiger, 2003 ...... 144 Scissurella azorensis Nolt, 2008 ...... 147 Scissurella bountyensis Powell, 1933 ...... 150 Scissurella cebuana (Bandel, 1998) ...... 153 Scissurella clathrata Strebel, 1908 ...... 158 Scissurella coronata Watson, 1886 [nomen dubium] ...... 166 Scissurella costata d’Orbigny, 1824 ...... 167 Scissurella cyprina Cotton & Godfrey, 1938 ...... 185 Scissurella dalli Bartsch, 1903 [nomen dubium] ...... 191 Scissurella dohrniana (Dunker, 1861) [nomen dubium] ...... 192 ? Scissurella electilis Montouchet, 1972 ...... 193 Scissurella evaensis Bandel, 1998 ...... 194 Scissurella georgica Davolos & Moolenbeek, 2006 ...... 201 Scissurella hoernesi Semper, 1865 [nomen dubium] ...... 203 Scissurella jucunda Smith, 1890 ...... 204 Scissurella kaiserae Geiger, 2006 ...... 212 Scissurella koeneni Semper, 1865 [nomen dubium] ...... 217 Scissurella lobini (Burnay & Rolán, 1990) ...... 218 Scissurella lorenzi Geiger, 2006 ...... 222 Scissurella manawatawhia Powell, 1937 ...... 226 Scissurella mantelli Woodward, 1859 [nomen dubium] ...... 230 Scissurella maraisorum Geiger, 2006 ...... 232 Scissurella mirifica (A. Adams, 1862) ...... 235

D. L. Geiger: Monograph of the Little Slit Shells 7 Scissurella morretesi Montouchet, 1972 ...... 252

Introduction Scissurella obliqua Watson, 1886 ...... 254 Scissurella ornata May, 1908 ...... 257 Scissurella petermannensis Lamy, 1910 ...... 260 Scissurella phenax new species ...... 266 Scissurella prendrevillei Powell, 1933 ...... 270 Scissurella quadrata Geiger & Jansen, 2004 ...... 278 Scissurella redferni (Rolán, 1996) ...... 285 Scissurella regalis Geiger & Marshall, 2012 ...... 289 Scissurella reticulata Philippi, 1853 ...... 291 Scissurella rota Yaron, 1983 ...... 296 Scissurella skeneoides new species ...... 304 Scissurella spinosa Geiger & Jansen, 2004 ...... 307 Scissurella staminea (A. Adams, 1862) ...... 317 Scissurella sudanica Bandel, 1998 ...... 321 Scissurella supraplicata Smith, 1875 ...... 327 Scissurella xandaros new species ...... 330 Scissurella aliceae Schnetler, Lozouet & Pacaud, 2000 † ...... 334 Scissurella annulata Ravn, 1933 † ...... 335 Scissurella apudornata (Laws, 1935) † ...... 336 Scissurella bituminata Beets, 1943 † ...... 339 Scissurella condita Laws, 1939 † ...... 340 Scissurella cossmanni Depontaillier, 1881 † ...... 343 Scissurella depontaillieri Cossmann, 1879 † ...... 345 Scissurella eocaenica Bandel, 1998 † ...... 351 Scissurella gruendeli (Bandel, 1998) † ...... 352 Scissurella hedegaardi Bandel, 1998 † ...... 353 Scissurella marchmontensis Sohl, 1992 † ...... 354 Scissurella peyrerensis Bandel, 1998 † ...... 357 Scissurella ravni (Schnetler, Lozouet & Pacaud, 2000) † ...... 359 Scissurella szaboi Conti & Monari, 1991 † ...... 360 Scissurella tinaeformis Amitrov, 1996 † ...... 361 Scissurella transylvanica Reuss, 1860 † ...... 363 Sinezona Finlay, 1926 ...... 367 Sinezona bandeli Marshall, 2002 ...... 372 Sinezona beddomei (Petterd, 1884) ...... 376 Sinezona brevis (Hedley, 1904) ...... 381 Sinezona brucei new species ...... 386 Sinezona calumnior new species ...... 389 Sinezona carolarum Geiger & McLean, 2010 ...... 395 Sinezona chilensis new species ...... 399 Sinezona cingulata (O. G. Costa, 1861) ...... 404 Sinezona concinna (Sowerby, 1831) [nomen dubium, nomen oblitum] ...... 419 Sinezona confusa Rolán & Luque, 1994 ...... 420 Sinezona costulata Geiger & Sasaki, 2009 ...... 428 Sinezona danieldreieri Geiger, 2008 ...... 432 Sinezona doliolum Herbert, 1986 ...... 438 Sinezona enigmatica Geiger & Marshall, 2012 ...... 441 Sinezona ferriezi (Crosse, 1867) ...... 444

8 D. L. Geiger: Monograph of the Little Slit Shells Sinezona finlayi (Laws, 1948) ...... 457 Sinezona garciai Geiger, 2006 ...... 459 Sinezona globosa Geiger, 2006 ...... 462 Sinezona hawaiiensis Geiger & McLean, 2010 ...... 466 Sinezona insignis (Smith, 1890) ...... 468

Sinezona insularis Simone, 2009 ...... 473 Introduction Sinezona iota (Finlay, 1926) ...... 474 Sinezona kayae Geiger & McLean, 2010 ...... 478 Sinezona laqueus (Finlay, 1926) ...... 482 Sinezona levigata (Iredale, 1908) ...... 489 Sinezona macleani Geiger, 2006 ...... 495 Sinezona marrowi new species ...... 500 Sinezona mechanica Geiger & Marshall, 2012 ...... 504 Sinezona milleri Geiger & Sasaki, 2009 ...... 508 Sinezona miranda (A. Adams, 1862) [nomen dubium] ...... 510 Sinezona modesta (A. Adams, 1862) [nomen dubium] ...... 511 Sinezona mouchezi (Vélain, 1877) ...... 513 Sinezona norfolkensis new species ...... 516 Sinezona pacifica (Oliver, 1915) ...... 519 Sinezona paumotuensis (Garrett, 1872) nomen[ dubium] ...... 525 Sinezona pauperata Powell, 1933 ...... 526 Sinezona platyspira Geiger & Marshall, 2012 ...... 530 Sinezona plicata (Hedley, 1899) ...... 533 Sinezona rimuloides (Carpenter, 1865) ...... 549 Sinezona semicostata Burnay & Rolán, 1990 ...... 556 Sinezona singeri Geiger, 2006 ...... 560 Sinezona subantarctica (Hedley, 1916) ...... 564 Sinezona wanganellica Geiger & Marshall, 2012 ...... 570 Sinezona wileyi Geiger, 2008 ...... 573 Sinezona zimmeri Geiger, 2003 ...... 578 Sinezona deshayesi (Munier-Chalmas, 1862) † ...... 582 Sinezona haliotimorpha (Bandel, 1998)† ...... 585 Sinezona koruahina (Laws, 1936) † ...... 587 Sinezona malloryi (Squires & Goeddert, 1996) † ...... 589 Sinezona praecedens Lozouet, 1998 † ...... 591 Sinezona terquemi (Deshayes, 1865) † ...... 593 Sinezona tertia (Laws, 1940) † ...... 594 Sukashitrochus Habe & Kosuge, 1964 ...... 598 Sukashitrochus atkinsoni (Tenison-Woods, 1877) ...... 600 Sukashitrochus carinatus (A. Adams, 1862) ...... 612 Sukashitrochus dorbignyi (Audouin, 1826) ...... 616 Sukashitrochus estotiensis Lozouet, 1999 ...... 624 Sukashitrochus lyallensis (Finlay, 1926) ...... 627 Sukashitrochus morleti (Crosse, 1880) ...... 632 Sukashitrochus pulcher (Petterd, 1884) ...... 646 Sukashitrochus kaiparaensis (Laws, 1941) † ...... 651 Sukashitrochus ngatutura (Laws, 1936) † ...... 653 Satondella Bandel, 1998 ...... 657 Satondella azonata Geiger & Marshall, 2012 ...... 659

D. L. Geiger: Monograph of the Little Slit Shells 9 Satondella bicristata Geiger & Marshall, 2012 ...... 661

Introduction Satondella brasiliensis (Mattar, 1987) ...... 663 Satondella cachoi Luque, Geiger & Rolán, 2011 ...... 668 Satondella danieli Segers, Swinnen & Abreu, 2009 ...... 673 Satondella dantarti Luque, Geiger & Rolán, 2011 ...... 676 Satondella goudi new species ...... 679 Satondella minuta Bandel, 1998 ...... 682 Satondella senni Geiger, 2003 ...... 684 Satondella tabulata (Watson, 1886) ...... 687 Satondella kondoi (Ladd, 1970) † ...... 692 Incisura Hedley, 1904 ...... 694 Incisura auriformis Geiger & Jansen, 2004 ...... 695 Incisura lytteltonensis (Smith, 1894) ...... 698 Incisura remota (Iredale, 1924) ...... 704 Incisura rosea (Hedley, 1904) ...... 709 Incisura geoffreyi (Laws, 1940) † ...... 713 Coronadoa Bartsch, 1946 ...... 717 Coronadoa demisispira Geiger & McLean, 2010 ...... 718 Coronadoa hasegawai Geiger & Sasaki, 2009 ...... 720 Coronadoa simonsae Bartsch, 1946 ...... 724

Volume 2

Anatomidae McLean, 1989 ...... 732 Anatoma Woodward, 1859 ...... 733 Anatoma aedonia (Watson, 1886) ...... 739 Anatoma africanae (Barnard, 1963) ...... 742 Anatoma agulhasensis (Thiele, 1925)...... 744 Anatoma alta (Watson, 1886) ...... 752 Anatoma alternatisculpta Geiger & McLean, 2010 ...... 756 Anatoma amoena (Thiele, 1912) ...... 760 Anatoma amydra Geiger & Marshall, 2012 ...... 764 Anatoma argentinae (Zelaya & Geiger, 2007) ...... 771 Anatoma aspera (Philippi, 1844) ...... 774 Anatoma atlantica (Bandel, 1998) ...... 788 Anatoma aupouria (Powell, 1937) ...... 792 Anatoma australis (Hedley, 1903) ...... 796 Anatoma austrolissa Geiger & Sasaki, 2008 ...... 802 Anatoma bathypacifica (Geiger & McLean, 2010) ...... 806 Anatoma bertheloti (d’Orbigny, 1840) [nomen dubium] ...... 809 Anatoma biconica new species ...... 812 Anatoma boucheti Geiger & Sasaki, 2008 ...... 817 Anatoma breveprima new species ...... 819 Anatoma concinna (A. Adams, 1862) ...... 823 Anatoma conica (d’Orbigny, 1841) ...... 828 (Fleming, 1828) ...... 832

10 D. L. Geiger: Monograph of the Little Slit Shells Anatoma disciformis (Golikov & Sirenko, 1980) ...... 848 Anatoma emilioi Geiger, 2011 ...... 851 Anatoma equatoria (Hedley, 1899) ...... 854 Anatoma euglypta (Pelseneer, 1903) ...... 862 Anatoma eximia (Seguenza, 1877) ...... 872

Anatoma finlayi (Powell, 1937) ...... 877 Introduction Anatoma flemingi (Marshall, 2002) ...... 883 Anatoma flexidentata Geiger & Sasaki, 2008 ...... 889 Anatoma fujikurai Sasaki, Geiger & Okutani, 2010 ...... 895 Anatoma globulus new species ...... 901 Anatoma gunteri (Cotton & Godfrey, 1933) ...... 905 Anatoma hyposculpta new species ...... 909 Anatoma indonesica Bandel, 1998 ...... 913 Anatoma janetae Geiger, 2006 ...... 917 Anatoma janusa new species ...... 923 Anatoma japonica (A. Adams, 1862) ...... 926 McLean, 1970 ...... 942 Anatoma kelseyi (Dall, 1905) ...... 948 Anatoma kopua Geiger & Marshall, 2012 ...... 956 Anatoma lamellata (A. Adams, 1862) ...... 960 Anatoma lyra (Berry, 1947) ...... 964 Anatoma maxima (Schepman, 1908) ...... 972 Anatoma megascutula Geiger & Marshall, 2012 ...... 978 Anatoma micalii new species ...... 982 Anatoma munieri (Fischer, 1862) ...... 986 Anatoma orbiculata new species ...... 996 Anatoma parageia Geiger & Sasaki, 2009) ...... 1001 Anatoma peruviana (Geiger & McLean, 2010) ...... 1004 Anatoma philippinica (Bandel, 1998) ...... 1008 Anatoma planapex new species ...... 1012 Anatoma plicatazona Geiger & McLean, 2010 ...... 1017 Anatoma porcellana new species ...... 1020 Anatoma proxima (Dall, 1927) ...... 1026 Anatoma pseudoequatoria (Kay, 1979) ...... 1035 Anatoma quadraxialis new species ...... 1045 Anatoma rainesi Geiger, 2003 ...... 1047 Anatoma rapaensis Geiger, 2008 ...... 1054 Anatoma regia (Mestayer, 1916) ...... 1060 Anatoma rhynchodentata new species ...... 1065 Anatoma rolani Geiger & Fernández-Garcés, 2010 ...... 1071 Anatoma sagamiana (Okutani, 1964) ...... 1075 Anatoma schanderi Høisaeter & Geiger, 2011 ...... 1081 Anatoma schioettei Høisaeter & Geiger, 2011 ...... 1086 Anatoma shiraseae Numanami & Okutani, 1990 ...... 1089 Anatoma sinuosa new species ...... 1091 Anatoma soyoae (Habe, 1951) ...... 1095 Anatoma tabulata (Barnard, 1964) ...... 1097 Anatoma tangaroa Geiger & Marshall, 2012 ...... 1101 Anatoma tenuis (Jeffreys, 1877)...... 1104

D. L. Geiger: Monograph of the Little Slit Shells 11 Anatoma tenuisculpta (Seguenza, 1877) ...... 1109

Introduction Anatoma tobeyoides Geiger & Jansen, 2004 ...... 1118 Anatoma umbilicata (Jeffreys, 1883)...... 1122 Anatoma weddelliana (Zelaya & Geiger, 2007) ...... 1127 Anatoma yaroni Herbert, 1986 ...... 1131 Anatoma zancliformis new species ...... 1136 Anatoma gephyra Maxwell, 1992 † ...... 1141 Anatoma miocenica (Laws, 1939) † ...... 1143 Anatoma muelleri new species † ...... 1147 Anatoma philippiana (Semper, 1865) † ...... 1149 Anatoma tongaensis Ladd, 1970 † ...... 1154 Sasakiconcha Geiger, 2006 ...... 1157 Sasakiconcha elegantissima Geiger, 2006 ...... 1158 Larocheidae Finlay, 1927 ...... 1162 Larochea Finlay 1927 ...... 1162 Larochea miranda Finlay, 1927 ...... 1163 Larochea secunda Powell, 1937 ...... 1168 Larochea scitula Marshall, 1993 ...... 1171 Larochea spirata Geiger & Marshall, 2012 ...... 1174 Larocheopsis Marshall, 1993 ...... 1177 Larocheopsis amplexa Marshall, 1993 ...... 1177 Larocheopsis macrostoma Geiger & Marshall 2012 ...... 1180 Trogloconcha Kase & Kano, 2002 ...... 1182 Trogloconcha christinae Geiger, 2003 ...... 1183 Trogloconcha lamellinodosa new species ...... 1185 Trogloconcha lozoueti Geiger, 2008 ...... 1190 Trogloconcha ohashii Kase & Kano, 2002 ...... 1194 Trogloconcha tesselata Kase & Kano, 2002 ...... 1197 Trogloconcha marshalli (Lozouet, 1998) † ...... 1201 Depressizonidae Geiger, 2003 incertae sedis ...... 1203 Depressizona Geiger, 2003 ...... 1204 Depressizona axiosculpta Geiger, 2009 ...... 1205 Depressizona exorum Geiger, 2003 ...... 1207 Sutilizonidae McLean, 1989 Vetigastropoda incertae sedis: Lepetodriloidea? ...... 1210 Sutilizona McLean, 1989 ...... 1211 Sutilizona pterodon Warén & Bouchet, 2001 ...... 1211 Sutilizona theca McLean, 1989 ...... 1215 Sutilizona tunnicliffae Warén & Bouchet, 2001 ...... 1217 Temnocinclidae McLean, 1989 Vetigastropoda incertae sedis: Lepetodriloidea? ...... 1220 Temnocinclis McLean, 1989 ...... 1221 Temnocinclis euripes McLean, 1989 ...... 1221 Temnozaga McLean, 1989 ...... 1224 Temnozaga parilis McLean, 1989 ...... 1224 Triassurella Nützel & Geiger, 2006 † ...... 1228 Triassurella carnica Nützel & Geiger, 2006 † ...... 1230 Triassurella goederti Kiel, Campbell & Gaillard, 2010 † ...... 1231 Problem specimens...... 1232 Literature cited ...... 1238 Index ...... 1281

12 D. L. Geiger: Monograph of the Little Slit Shells