A New Species of Egglestonichthys from Northern Australian Estuaries (Teleostei, Gobiidae, Gobiinae)

aqua, International Journal of Ichthyology

A new species of Egglestonichthys from northern Australian estuaries (Teleostei, Gobiidae, Gobiinae)

Helen K. Larson

Museum and Art Gallery of the Northern Territory, P.O. Box 4646, Darwin, NT 0801, Australia; Museum of Tropical Queensland, 70-102 Flinders street, Townsville, Queensland 4810, Australia. School of Marine and Tropical Biology, James Cook University, Townsville, Queensland 4811, Australia.

Received: 02 June 2013 – Accepted: 07 July 2013

Abstract midae). L’espèce la plus proche pourrait être E. bombylios A new species, that appears to belong to the gobiid genus Larson et Hoese, bien que le patron de ses papilles senso- Egglestonichthys, is described from specimens collected from rielles diffère de celles des autres membres du genre. Une the shallow coast and estuaries of Kakadu National Park in clé des quatre espèces d’Egglestonichthys connues à ce jour the Northern Territory, Australia. It is unusual among gobi- est fournie. ids in that the lower half of the pectoral-fin lacks connecting membrane between the fin rays, so that the rays are free Sommario (resembling the pectoral fin of the fishes of the family Una nuova specie, che sembra appartenere ai gobidi del Polynemidae). Its closest relative may be E. bombylios Lar- genere Egglestonichthys, è descritta sulla base di esemplari rac- son and Hoese, although its sensory papilla pattern differs colti in estuari e zone costiere poco profonde del Kakadu from others in the genus. A key to the four known species National Park nel Northern Territory, Australia. Presenta presently placed in Egglestonichthys is provided. un’insolita caratteristica tra i gobidi: la metà inferiore della pinna pettorale manca di membrana di collegamento tra i Zusammenfassung raggi, che, pertanto, risultano liberi. (ciò la rende simile alla Beschrieben wird eine neue Art, die offenbar zu den pettorale dei pesci della famiglia Polynemidae). Il suo Grundeln der Gattung Egglestonichthys gehört, nach parente più prossimo potrebbe essere E. bombylios Larson e Exemplaren, die an der flachen Küste und in den Münd- Hoese, anche se il modello di distribuzione delle papille ungsgebieten im Kakadu-Nationalpark des Nordterritori- sensoriali è diverso da tutte le altre specie del genere. Viene ums in Australien gesammelt wurden. Sie zeigen ein unter fornita una chiave di identificazione per le quattro specie Gobiiden ungewöhnliches Merkmal: die untere Hälfte der attualmente conosciute collocate in Egglestonichthys. Brustflossen hat keine verbindende Membran zwischen den Flossenstrahlen, sodass die Strahlen frei stehen (ähnlich wie INTRODUCTION bei den Fischen der Familie Polynemidae). Als nächster In 1997, during an estuarine fish survey along the Verwandter dürfte E. bombylios Larson and Hoese gelten, obwohl das Muster der Sinnes-Papillen sich von allen ande- coast of Kakadu National Park, Northern Territory, ren in der Gattung unterscheidet. Beigefügt ist ein Bes- specimens of an unusual small gobiid were obtai- timmungsschlüssel zu den vier Arten, die bisher der Gatt- ned by a small beam trawl (2 m wide mouth, 6.5 ung Egglestonichthys zugeordnet werden konnten. mm mesh body and 1.4 mm mesh cod end). Subsequent surveys in 1998, 1999 and 2001 Résumé produced additional specimens. The fish have been Une nouvelle espèce, qui semble appartenir au genre de tentatively identified as belonging to Egglestonich - Gobiidés Egglestonichthys, est décrite sur base de spécimens thys, a genus containing three other species of collectés sur la côte étroite et dans les estuaires du parc somewhat disparate appearance. Miller and Won- national de Kakadu, Territoire du Nord, Australie. Elle diffère des autres Gobiidés par le fait que la moitié infé- grat (1979) created the genus for a goby with a rieure des pectorales n’a pas de membranes entre les complex transverse papilla pattern resembling that rayons, ce qui implique que ces nageoires sont libres (évo- found in some eleotrids (e.g. Eleotris), lacking late- quant la pectorale des poissons de la famille des Polyne- ral-line sensory pores and having a wide gill open-

147 aqua vol. 19 no. 3 - 19 July 2013 A new species of Egglestonichthys from northern Australian estuaries (Teleostei, Gobiidae, Gobiinae) ing (to below preoperculum). Larson and Hoese The new species somewhat resembles E. bombylios (1997) described a second species (E. bombylios) Larson and Hoese, 1997, in general appearance, and redescribed E. melanoptera Rao, originally having reduced eyes, moderate gill opening and described as a Callogobius. Miller and Wongrat with some raised sensory papilla rows on the head, (1979) provided the only published osteological but differing in having a number of fin rays in the information for the genus (based on the type spe- lower half of the pectoral-fin free from connecting cies, E. patriciae). membranes, resembling the rays of a polynemid,

Fig. 1. Locality map for known records of Egglestonichthys ulbubunitj n. sp. Map prepared by Michael Hammer. aqua vol. 19 no. 3 - 19 July 2013 148 Helen K. Larson no scales on the cheek or opercle, as well as a sen- Museum, Sydney; NTM, Museum and Art Gallery sory papilla arrangement that is unlike other spe- of the Northern Territory, Darwin: QM, Queens- cies of Egglestonichthys. Although the new species land Museum, Brisbane; WAM, Western Aust- has several features that differ from other Egglesto- ralian Museum, Perth. nichthys, the group is so little-studied that I am reluctant to create yet another new genus based on limited information. Egglestonichthys ulbubunitj n. sp. So far this fish is known only from coastal marine Threadfin Goby and estuarine waters within Kakadu National Park (Figs 2-4, Table I) (Larson 2002) (Fig. 1). Similar beam trawl and dredging surveys have been carried out in other Holotype: NTM S.14635-005, 28 mm SL female, estuaries and harbours in the Northern Territory, off Pococks Beach, N of West Alligator Head, 1.4 but this species has not been not found. It is most m, very soft mud, beam trawl, 26 May 1998, R. interesting that this new species has not yet been Williams, G. Dally, A. Pickworth and M. Gorst. collected elsewhere in the Territory, and it is pos- Paratypes: NTM S.17348-001, 3(26.5-30), same sible that the fish has very specific habitat require- data as holotype; NTM S.14479-003, 1(31), ments. centre of West Alligator River mouth, 5.5 m, mud, beam trawl, 27 May 1997, coll. R. Williams and A. METHODS Pickworth; NTM S.14657-007, 4(10.5-20), off Measurements were taken using electronic calli- “The Rookery”, East Alligator River, 7.3 m, mud, pers and dissecting microscope. Counts and met- beam trawl, 2 June 1998, coll. R. Williams and G. hods generally follow Hubbs and Lagler (1970), Lindner; NTM S.15703-001, 1(18.5), 5 km offs- except as indicated below. Transverse scale counts hore between Field Island and Point Farewell, 9.6 backward (TRB) are taken by counting the num- m, mud, beam trawl, coll. R. Williams, S. Morr- ber of scale rows from the anal-fin origin diago- ison and D. Elphick, 12 June 2001; NTM nally upward and back toward the second dorsal- S.16024-001, 1(28), N of West Alligator Head, 16 fin base. Head length is taken to the upper attach- m, soft mud and shell grit, dredge, coll. N. Smit ment of the opercular membrane. References to and party, 21 November 2004; QM I.39101, second dorsal and anal-fin rays includes a thin flex- 4(18-24), 1 km off Pococks Beach, West Alligator ible spine in addition to the segmented rays. The Head, 5 m, mud, beam trawl, coll. A. Pickworth, segmented or branched caudal ray pattern (e.g. 9/8 M. Gorst and A. Turner, 7 June 1999; AMS or 9/7) is the number of segmented caudal rays I.46190-001, 2(24-27), off Midnight Point, South attaching to the upper and lower hypural plates Alligator River, 10 m, mud and detritus, beam respectively. Vertebral counts and other osteologi- trawl, coll. R. Williams, S. Morrison and D. Elp- cal information were obtained by X-ray. Pterygiop- hick, 12 June 2001; NTM S.15289-003, 1(19), 31 hore formula follows Birdsong et al. (1988). km upstream from mouth of South Alligator River, Institutional abbreviations are: AMS, Australian 10-11 m, mud, beam trawl, coll. G. Lindner, F.

Fig. 2. Holotype of Egglestonichthys ulbubunitj n. sp., NTM S.14635-005, 28 mm SL female. Photo by J. Larson.

149 aqua vol. 19 no. 3 - 19 July 2013 A new species of Egglestonichthys from northern Australian estuaries (Teleostei, Gobiidae, Gobiinae)

Table I. Morphometrics of Egglestonichthys ulbubunitj n. sp., expressed as percentage of SL or HL as indicated (n = 21).

Holotype Mean Maximum Minimum

Head length in SL 29.3 30.4 33.3 27.9 Head depth in HL 69.5 67.7 75.0 61.7 Head width in HL 76.8 68.5 83.0 60.7 Body depth in SL 23.9 21.2 23.9 18.4 Body width in SL 9.3 8.6 10.4 7.0 Caudal peduncle length in SL 19.6 20.5 24.6 18.3 Caudal peduncle depth in SL 13.6 11.8 13.6 10.5 Snout length in HL 23.2 23.5 27.7 20.0 Eye width in HL 15.9 15.9 18.3 13.5 Upper jaw length in HL 42.7 45.5 50.0 38.3 Interorbital width in HL 23.2 18.4 23.2 14.0 Pectoral-fin length in SL 27.9 29.0 33.7 26.7 Pelvic-fin length in SL 28.6 29.5 33.9 26.7 Caudal-fin length in SL 53.6 56.2 63.6 50.0 Longest dorsal-fin spine in SL 13.6 15.9 19.1 13.0

Baird and G. McSkimming, 6 June 1999; WAM P. South Alligator River, 12° 12’ 48.5994” 132° 24’ 33873-001, 1(33), Brooks Creek mouth, South 53.9994”, in 3.5m depth over mud, beam trawl, Alligator River, 8-9 m, mud, beam trawl, coll. G. coll. B. Pusey and M. Kennard, 27 August 2012 Lindner, F. Baird and G. McSkimming, 6 June [frozen in the field for later identification]. 1999; NTM S.14441-012, 2(20-24.5), off mouth Diagnosis: A small, slender, round-headed goby of Wildman River, 2 m, firm mud and plant detri- with small eyes, long pointed caudal fin and lower tus, beam trawl, coll. H. Larson, R. Williams, A. 3-7 pectoral-fin rays free from membrane; second Pickworth and M. Gorst, 28 May 1997. dorsal-fin rays always I,10; anal fin rays always Non-type material: NTM S.17479-001, 1(29), I,10; pectoral-fin rays 19-21; longitudinal scales

Fig. 3. Egglestonichthys ulbubunitj n. sp., diagram of head showing sensory papillae, NTM S.15703-001, 18.5 mm SL female. aqua vol. 19 no. 3 - 19 July 2013 150 Helen K. Larson

22-25; TRB 7-9; side of head naked, predorsal sca- 15*), 3-2310 (in 1). Vertebrae 10+16 (in 18). One les 9-12, extending forward to behind eyes; eyes epural (in 17*), two in one specimen. Two anal small and dorsolateral; interorbital broad and con- pterygiophores before haemal spine of first caudal vex; jaws oblique, with small caniniform teeth; vertebra (in 18). head pores absent; reduced papillae pattern on Head rounded in cross-section, length 27.9- head arranged in mostly longitudinal pattern, 33.3% (mode 30.0%) in SL; profile rounded to papillae often pointed, fleshy and may be on short slightly pointed. Depth at posterior preopercular raised ridges; eyes silver when alive, head and body margin 61.7-75.0% (mode 66.7%) in HL. Head colour pale yellowish-white, with six or seven ind- width at posterior preopercular margin 60.7- istinct dusky bands across nape and sides, and large 83.0% (mode 76.8%) in HL. Mouth terminal, oval black spot on posterior part of first dorsal fin. oblique, forming an angle of 35-40° with body Description: Based on 21 specimens, 18-33 mm axis; jaws reaching to below posterior margin of eye SL. Counts of holotype (Fig. 2), where differing or at least posterior half. Lips smooth, lower lip from paratypes, indicated by asterisk (in parenthe- free, fused on either side of broad flat chin. Upper ses where necessary); morphometrics in Table I. jaw 38.3-50.0% (mode 45.0%) in HL. Eye very First dorsal fin VI; second dorsal-fin rays always small, placed laterally, closer to snout tip than to I,10; anal-fin rays always I,10; pectoral-fin rays 19- rear margin of preopercle, 13.5-18.3% (mode 21 (usually 20 (holotype with 20 in left fin, 21 on 15.9%) in HL. Snout broad and short, 20.0- right); segmented caudal-fin rays 17; caudal-fin ray 27.7% (mode 23.2%) in HL. Interorbital very pattern 9/8 (one specimen with 9th upper ray split broad, flattened, 14.0-23.2%% (mode 19.0%) in to near base); branched caudal-fin rays 8/6 (in 10), HL. Body slender and compressed, depth at anal 8/7* (in 11), 7/6 (in 1); unsegmented (procurrent) origin 18.4-23.9%% (mode 23.9%) in SL; body caudal-fin rays 6/6 to 8/8 (usually 7/6 or 7/7, 8/7 width at origin of first dorsal 7.0-10.4% (mode in holotype); longitudinal scale count 22-25 7.0%) in SL. Caudal peduncle length 20.0-24.6% (modally 23*); TRB 7-9 (modally 8*); predorsal (mode 20.0%) in SL. Caudal peduncle depth 10.5- scale count 9-12 (modally 9; 11 in holotype); circ- 13.6% (mode 13.6%) in SL. umpeduncular scales 12*-13 (modally 12). Gill No mental fraenum; single transverse row of rakers on outer face of first arch 3+11 (1), 4+11 fleshy papillae present on chin. Anterior naris in (1), 5+11 (1). Pterygiophore formula 3-22110* (in broad, very short tube, close behind upper lip.

Fig. 4. Egglestonichthys ulbubunitj n. sp., showing fresh dead colour pattern; photograph taken through microscope; whole fish could not be viewed; NTM S.17479-001, 29 mm SL, South Alligator River. N.B. some small yellowish patches of org- anic material still adhere to the specimen. Photo courtesy of Charles Darwin University.

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Posterior naris rounded to oval, placed above and ral-fin rays branched, upper and lowermost rays forward of anterior margin of eye. Gill opening unbranched; lowermost three to seven (usually five moderate, extending forward to just under opercle to six, three in only one specimen) rays free from or to mid-opercle, but not reaching posterior preo- membrane (tips of branches of each individual percular margin. Anterior edge of shoulder girdle branched ray joined by membrane). Pelvic-fins (cleithrum) smooth. Gill rakers on outer face of large (reaching anus), oval, disc-shaped, fraenum first arch without spines, fleshy and slender, deep; fin length 26.7-33.9% (mode 28.6%) in SL. somewhat shorter than gill filaments; rakers on Caudal fin very long and pointed, 50.0-63.6% inner face of first arch short, stubby, without fine (mode 53.6%) in SL. spines; outer rakers on other arches similar to inner Live coloration: Only a few notes were face rakers. Tongue short, blunt to slightly taken when the holotype was collected: “translucent rounded. grey fat-headed goby with small silver eyes and ind- Outer row teeth in upper jaw small, sharp and istinct grey cross-hatched bands across back”. Fig- pointed, in three rows across front and two along ure 4 shows the freshly dead non-type specimen. side; outermost row teeth larger, more curved, with Coloration in alcohol: Background largest two to six teeth on either side of symphysis. colour yellowish white, with six or seven short faint Outer row teeth in female slightly smaller than brownish saddles crossing back; first on nape those in male. Lower jaw with two to three rows of similarly sized, small sharp teeth, none particularly large; outermost row teeth may be more upright and less curved. Scales on head and body all cycloid. Predorsal scales small, reaching forward to close behind eyes. Cheek and operculum without scales. Pectoral base naked or with few large scales. Prepelvic area with moderately-sized scales, may be embedded. Head pores absent. Sensory papillae pattern on head in pattern as shown diagrammatically in Figure 3; appearing longitudinal but with few rows that appear transverse (and with some individual variation); all specimens show some damage to sensory papillae due to abrasion during trawling and subsequent handling. Papillae on head large, fleshy and often with pointed tips, in some specimens, papillae appear as two forms: vertically oriented papillae on short raised ridges, horizontally oriented papillae small and oval to round, within rounded fleshy depressions. In some specimens, papillae barely visible above skin, and can be seen through partly transparent flesh as short ridges; papillae on under- side of head usually discernible above skin. Genital papilla in male short, small, flattened, with blunt to slightly pointed tip. Genital papilla in females very short, bulbous. First dorsal-fin low, rounded, spines barely reaching first element of second dorsal-fin origin; third or fourth dorsal spine longest, 13.0-19.1% in SL. Fig. 5. Typical beam-trawl catch on the South Alligator Second dorsal and anal-fins long, pointed posteri- River, Northern Territory, after 20 minutes’ haul with 2 orly, posteriormost rays of these fins usually reach- metre beam-trawl. Rex Williams is holding an adult Strea- ing onto caudal fin. Pectoral-fin relatively narrow, mer Threadfin, Parapolynemus verekeri (Polynemidae), pointed, 26.7-33.7% (mode 30.0%) in SL; pecto- which co-occurs with E. ulbubunitj. Photo by H. K. Larson. aqua vol. 19 no. 3 - 19 July 2013 152 Helen K. Larson before first dorsal fin and seventh at caudal-fin base nal Owner Jonathan Nadji. He stated: “My family’s (two saddles under second dorsal fin may coalesce); clan area includes the area where this fish (goby saddles usually joining short staggered oblique pale threadfin) was discovered. Our clan, Ulbu Bunidj, brownish bars or blotches along midside of body; shares this area with Djindibi. I would like this fish within brownish saddles, scale margins narrowly to be named after our clan, which is Ulbu Bunitj”. outlined with slightly darker pigment (Fig. 4). Suggested common name: Threadfin Goby. Head and chin evenly speckled with fine brown; Remarks: This species will not key out to Egg- nape paler, may show one to three indistinct lestonichthys in Larson and Murdy’s 2001 key to brownish bars crossing from opercle to opercle. western Pacific gobiid genera. In fact, anyone Indistinct short brown line from front margin of attempting to use this key for any species of Egg- eye to middle to upper jaw. Remainder of under- lestonichthys will not succeed as couplet 35b is side of head, breast and belly whitish. incorrect, unfortunately (it states that head pores First dorsal-fin translucent, anteriorly with faint are present, when this is not so in Egglestonichthys; brownish wedge-shaped mark extending up from writing a working key to 76 often poorly defined second dorsal saddle; distinct black round to oval genera is difficult). spot on posterior part of fin extending from fourth As noted by Larson and Hoese (1997), there is to sixth spine. Second dorsal-fin translucent to some variation of features among the described faintly brownish, fin margin narrowly unpigmen- species of this genus. Egglestonichthys ulbubunitj ted. Anal-fin translucent, with variable amounts lacks spines on the gill rakers, as does E. bombylios. and intensity of brown speckling along basal half of The sensory papillae pattern on the head of E. fin, usually more prominent on posterior part of ulbubunitj is not like that of any of the other spe- fin. Pectoral-fin translucent, with faint wedge of cies, in that the transverse pattern is reduced and brown speckling on upper half, an extension of an varies somewhat between specimens, although the indistinct brown bar along bases of pectoral-fin papilla structures themselves (fleshy, partly raised rays and rear part of pectoral-fin base. Pelvic-fins on ridges) do resemble those of E. bombylios. Of translucent. Caudal-fin translucent with faint the four known species of the genus, only E. pat- brownish speckling basally and often along riciae and E. melanoptera are similar to each other membranes (mostly on lower half of fin). (and could be conspecific; fresh, undamaged Distribution: Known only from turbid estuarine material is needed). Neither E. bombylios nor E. and coastal habitats in the Alligator Rivers region, ulbubunitj greatly resemble the other two. The Northern Territory. paucity of specimens makes further analysis diffi- Comparisons: This is a distinctive fish, and is cult – one known specimen of E. patriciae, four unlikely to be confused with any other gobiid, each of E. melanoptera and E. bombylios as comp- especially when the pectoral fins are examined. It ared to 21 of E. ulbubunitj, the smallest species does not closely resemble the other species of the (N.B. have just learned that 30 specimens of E. genus Egglestonichthys; indeed, all four described melanoptera have recently been collected off India; species are rather different from each other in body pers. comm. D. Ray, Zoological Survey of India, form and colour pattern (see Remarks and dicho- West Bengal). It is possible that more specimens tomous key below). are waiting among the trawled fish identified only Ecology: All specimens were obtained by 2-metre as “Gobiidae” in museums. beam trawl or small dredge over (and occasionally Further material of all species will confirm whet- in) very soft to firm mud (occasionally with broken her these four odd goby taxa truly belong together shell and plant detritus such as drifts of leaves), at as a group or not. Genetic analyses and further depths of 1-16 m. Water quality varied with the morphological examination will undoubtedly help tide, from 25.8-28.4°C and 28.0-30.3‰ salinity. disentangle them. Although it is not entirely satis- This small goby co-occurs with estuarine fishes factory to assign this new species to a genus to such as small polynemids, juvenile sciaenids, which it may not belong, it at least flags the pro- engraulids and cynoglossids in this soft-substrate blem and indicates that much work remains to habitat (Larson 1999). be done on all those small “trawl gobies” presently Etymology: Named ulbubunitj, to be treated as a waiting to be examined in collections world-wide. noun in apposition, for the Ulbu Bunidj clan in Arnhem Land, at the suggestion of senior Traditio-

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Key to the species of Egglestonichthys with us in collecting these peculiar fish during the Kakadu National Park estuarine fish monitoring 1. Pelvic frenum present, may be thin (and easily surveys from 1997 to 2002. Their willingness and damaged); body pale to brownish with darker good humour made the difficult task of beam- bands or blotches; mouth small to large ……2 trawling in deep mud habitats that much more 1a. Pelvic frenum absent; fins and body generally enjoyable. plain dark brown to black; mouth large and reaching to well below eye...... E. melanoptera (Rao, 1971) REFERENCES 2. Lower 3-7 pectoral-fin rays free from fin BIRDSONG, R. S., MURDY, E. O. & PEZOLD, F. L. 1988. A membrane; caudal-fin elongate and pointed, study of the vertebral column and median fin osteology much longer than head; lateral scales 22-25 ... in gobioid fishes with comments on gobioid relations- ...... E. ulbubunitj n. sp. hips. Bulletin of Marine Science 42 (2): 174-214. 2a. All pectoral-fin rays contained within mem- HUBBS, C. L. & LAGLER, K. F. 1970. Fishes of the Great brane; caudal fin not elongate and pointed; Lakes Region. University of Michigan Press: Ann Arbor. with at least 30 lateral scales or more ...... 3 LARSON, H. K. 1999. Report to Parks Australia North, on the estuarine fish inventory of Kakadu National Park, 3. Papilla rows on head raised on fleshy ridges Northern Territory, Australia. Museum and Art Gallery especially around jaws, anterior cheek and of the Northern Territory Research Report No. 5. 50 pp. snout; lateral scales 31-35; eyes small (9-13 in LARSON, H. K. 2002. Report to Parks Australia North, on HL)..... E. bombylios Larson and Hoese, 1997 estuarine fish monitoring of Kakadu National Park, 3a. Papilla rows on head not raised on fleshy rid- Northern Territory, Australia. Museum and Art Gallery ges; lateral scales 40; eyes moderate (3.5 in of the Northern Territory, unpublished report. 42 pp. HL).... E. patriciae Miller and Wongrat, 1979 LARSON, H. K. & HOESE, D. F. 1997. A new species of Egglestonichthys (Pisces; Teleostei; Gobiidae) from the Indo-West Pacific, with discussion of the species of the ACKNOWLEDGEMENTS genus. The Beagle, Records of the Museums and Art Galler- My many thanks to Rex Williams and Gavin ies of the Northern Territory 13: 45-52. Dally of the Museum and Art Gallery of the Nort- LARSON, H. K. & MURDY, E. O. 2001. Gobiidae. Gobies. hern Territory for their considerable help. Thanks Pp 3578-3603. In: Carpenter, K. E. and Niem, V. H. to Charles Darwin University staff for photograp- (eds) FAO species identification guide for fishery purposes. hing the freshly dead colouring of the small non- The living marine resources of the western Central Pacific. paratype specimen. Many thanks also to Garry Volume 6. Bony fishes part 4 (Labridae to Latimeriidae). Linder of Parks Australia North for all his assist- FAO, Rome. MILLER, P. J. & WONGRAT, P. 1979. A new goby (Teleos- ance, especially with logistics; Parks Australia tei: Gobiidae) from the South China Sea and its signific- North staff, in particular Mick Gorst, Anna Pick- ance for gobioid classification. Zoological Journal of the worth, Buck Salau and Andrew Wellings, and Linnean Society 67: 239-257. Traditional Owners Joseph Bishop, Leanne Alder- RAO, V. V. 1971. New gobioids from Godavari estuary. son and Kenny Wauchope; all of whom worked Journal of the Zoological Society of India 23 (1): 39-54.

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