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Una Nuova, Eccezionale Specie Appartenente Al Genere a New, Very Interesting Species Belonging to the Genus Corynopuntia Knuth

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Una Nuova, Eccezionale Specie Appartenente Al Genere a New, Very Interesting Species Belonging to the Genus Corynopuntia Knuth Una nuova eccezionale specie appartente al genere Corynopuntia Knunt: Corynopuntia guccinii D. Donati 115 Davide Donati Una nuova, eccezionale A new, very interesting specie appartenente al species belonging to the genere genus Corynopuntia Knuth: Corynopuntia guccinii D. Donati sp. nov. Riassunto: durante l'esplorazione di una valle Abstract: during the exploration of a valley del Coahuila meridionale, Messico, l'autore ha in southern Coahuila, Mexico, the Author scoperto un membro del genere Corynopuntia discovered a very interesting member of Knuth dalle caratteristiche eccezionali, qui the genus Corynopuntia Knuth, described descritto come C. guccinii species nova. here as C. guccinii species nova. 116 Davide. Donati Prefazione Preface Considerando: Considering that: - i risultati ottenuti nei recenti studi basati sulla - in 2002 WALLACE and DICKIE, on the basis sequenza del DNA da WALLACE & DICKIE, e of DNA sequence, and STUPPY, on the sullo studio microcarpologico da STUPPY, dove basis of seed micromorphology, proposed viene proposto di ascrivere nuovamente al a new classification of the subfamily Opun- rango generico molti generi prima inglobati in tioideae, ascribing once more at generic Opuntia; rank many opuntioid genera; - lo studio genetico di GRIFFITH, nel quale si - in 1999 ANDERSON included Corynopuntia dimostra come un genere Grusonia sensu in an enlarged Grusonia, but GRIFFITH Anderson comprendente Corynopuntia Knuth e demonstrated that this generic recombina- Micropuntia Daston, sia parafiletico, e si ripri- tion involve a paraphyletic Grusonia, and stinano i suddetti generi Corynopuntia e Micro- reinstated Corynopuntia and Micropuntia; puntia; - HUNT et al. followed partially GRIFFITH stu- - che HUNT et al. accettano la proposta di GRIF- dies, accepting his conclusion but including FITH di ripristinare il genere Corynopuntia; Micropuntia in Corynopuntia; in questo studio si seguirà la classificazione in this study GRIFFITH classification will be proposta da GRIFFITH. assumed. La scoperta The discovery Lo stato messicano di Coahuila, nella sua por- The south-western portion of the state of zione sud occidentale, è caratterizzato da vaste Coahuila, Mexico, is characterized by high pianure alluvionali che intervallano imponenti mountain ranges, divided by wide alluvial catene montuose. Tali pianure sono caratterizza- plains. Such plains are typically very arid, te da suoli argilloso-gessosi, allagati in caso di with loamy-gypseous soil, sometimes floo- forti precipitazioni. Il clima è decisamente arido. ded by heavy rainstorm. Una nuova eccezionale specie appartente al genere Corynopuntia Knunt: Corynopuntia guccinii D. Donati 117 Durante l'esplorazione During the explora- di alcune di queste tion of one of these valli a nord dell'abitato valleys, North of di San Pedro de las San Pedro de las Colonias, l'autore ha Colonias, I found a incontrato un taxon taxon belonging to ascrivibile al genere the genus Coryno- Corynopuntia dalle puntia, with excep- caratteristiche eccezio- tional characteri- nali: stics: - enormi dimensioni, i - very large, with cespi possono supera- groups more than 3 re i 3 metri di diame- m wide and more tro ed un'altezza di than 40 cm tall; oltre 40 cm; - often semi erect - portamento inizial- when young, pro- mente semi eretto, strate when old; solo successivamente very elongate prostrato; joints, sometimes - articoli molto allungati, more than 15 cm lunghi talvolta oltre 15 long and 4 cm in cm e 4 cm di diametro; diam.; - 4 (8) spine centrali - 4 (- 8) central spi- giallo rosate, piane, nes, flattened, yel- lunghe oltre 10 cm; Olotipo/Holotipe lowish pink, more - radice napiforme; than 10 cm long; - fiore rosso vinaccia, con gola giallo verdastra. - napiform root; Considerando che: - red flower, with yellow-greenish throat. - le dimensioni di questo taxon ne fanno la più Considering that: grande Corynopuntia orientale e uno dei più gran- - this taxon is the biggest eastern Coryno- di taxa del genere, con taglia nettamente maggio- puntia and one of the biggest taxa of the re rispetto alle specie geograficamente vicine, ma genus, larger than any in the area, compa- comparabile solo con le grandi Corynopuntia a rable only with the big north-western Cory- distribuzione nord-occidentale, come C. emoryi ( nopuntias such as C. emoryi (Engelm.) Engelm.) M.P. Griffith, C. kunzei (Rose) M.P. M.P. Griffith, C. kunzei (Rose) M.P. Griffith, Griffith, C. invicta (Brandegee) Knuth; C. invicta (Brandegee) Knuth; - il suo portamento giovanile è sovente peculiare; - its early, often erect growth is peculiar in - il colore rosso del fiore deve essere considera- the genus; to una caratteristica eccezionale, condiviso nel - the red colour of the flower should be con- genere Corynopuntia solo con C. vilis (Rose) H. sidered as a very interesting character, Robinson, che però presenta una tonalità molto shared in the genus only with C. vilis diversa, magenta intenso, ed una morfologia (Rose) H. Robinsonw, which shows decisamente distinta; nevertheless magenta flower and a very - il taxon in questione condivide simpatricamen- distinct morphology; te l'habitat con ben 3 specie di Corynopuntia, - the taxon is sympatric with three other quali C. moelleri (Berg.) Knuth, C. bulbispina Corynopuntias, C. moelleri (Berg.) Knuth, (Engelm.) Knuth, C. grahamii (Engelm.) Knuth, C. bulbispina (Engelm.) Knuth, C. grahamii senza che si presenti alcun ibrido tra i vari taxa; (Engelm.), but no hybrids were detected. è parere dell'autore che tale taxon debba perciò I think that this taxon should be described essere descritto come species nova. as species nova. 118 Davide. Donati Corynopuntia guccinii D. Donati sp. nov. Diagnosi latina/Latin diagnosis Corynopuntia guccinii D. Donati sp. nov. Descriptio: frutex formans turmas in latitudinem plus quam 3 m et in altitudinem plus quam 40 cm patentes, quae a densis catenis articulorum, initio habitu erecto, deinde ad terram proiecto, consti- tutae sunt; radix rapacea sed tenua; articuli magni et elongati, usque ad 15 cm longi et 4 cm dia- metientes; spinae mediae 4 (interdum 8) usque ad 10 cm longae, planae, scabrae, pungentes, luteo- roseae; spinae radiales 6-12, usque ad 20 mm longae, albae; glochidia pauca, albescentia; flores lati 4-5 cm, rubido colore, fauce luteo-subviridi, stamina lutea, receptaculum tuberculatum gerens mul- tas areolas glochidiis et spinis subtilibus instructas; fructus maturi exarescentes, clavati, spinosi, 5 cm longi, perianthii residua persistentia gerentes, semina suborbicularia 3,5-6 mm diametientia, compressa, cinerea. Typus: MEXICO, COAHUILA: Mpio. San Pedro de las Colonias, ad septentrionem San Pedro, in planitiis alluviis, 850 m, iunii 2009, Hinton et al. 29067 (holotypus Herbarium Geo. B. Hintonii; Isotypus MEXU). Distributio: endemica rei publicae mexicanae Coahuila, in alluvionibus argillosis, ubi cum C. bulbispi- na, C. grahamii, C. moelleri, Coryphantha werdermannii, Echinocactus horizonthalonius, etc. crescit. Descrizione: pianta cespitosa, costituente grup- Description: body cespitose, forming pi larghi oltre 3 metri e alti oltre 40 cm; formata groups over 3 m wide and more than 40 cm da dense catene di articoli, a portamento inizial- tall; formed by dense, initially erect and mente eretto, poi prostrato; radice sottile ma fit- after prostrate chains of joints; root tonante; articoli grandi ed allungati, lunghi fino napiform but thin; segments big and elon- a 15 cm, 4 cm di diametro; spine centrali 4 (8 gate, 15 X 4 cm; central spines 4 (someti- talvolta), lunghe fino a 10 cm, appiattite, scabre, mes 8), 5-10 cm long, generally flattened, pungenti, giallo rosate; spine radiali 6-12, lun- very pungent, rough, yellow-pinkish; radial ghe fino a 20 mm, bianche; glochidi biancastri, spines 6-12, approx. 5-20 mm long, whi- non numerosi; fiore ampio 4-5 cm, rosso vinac- tish; glochids small, whitish, few; flower 4 to cia con gola giallo-verdastra, stami gialli, ovario 5 cm broad, red, with light yellow-greenish tubercolato con numerose areole ornate da glo- throat, filaments yellow, receptacle slightly chidi e sottili spine; frutto secco a maturazione, tuberculate, with white glochids and thin, clavato, spinoso, 5 cm di lunghezza, con residui long spines; fruit dry, clavate, spiny, 5 cm del perianzio persistenti; semi pseudocircolari long, with persistent perianth residuals; 3,5-6 mm in diametro, schiacciati, beige. seeds subcircular, 3,5-6 mm diam., flatte- Tipo: MESSICO. COAHUILA: Mpio. San ned, beige. Pedro de las Colonias, nord di San Pedro, in Type: MEXICO. COAHUILA: Mpio. San pianure alluvionali, 850 m, giugno 2009, Hin- Pedro de las Colonias, North of San Pedro ton et al. 29067 (olotipo Herbarium di Geo. B. de las Colonias, on alluvial plains, 850 m, Hinton; Isotipo MEXU). June 2009, Hinton et al. 29067 (holotype Distribuzione: endemica dello stato messicano Herbarium of Geo. B. Hinton; Isotype di Coahuila, tipica delle pianure alluvionali MEXU). argillose, dove vive con C. bulbispina, C. Distribution: endemic of the Mexican state grahamii, C. moelleri, Coryphantha werder- of Coahuila, on loamy, alluvial plains, mannii, Echinocactus horizonthalonius, ecc. where it grows with C. bulbispina, C. graha- Dedicata a Francesco GUCCINI: incontrai la mii, C. moelleri, Coryphantha werderman- pianta ascoltando la canzone “Incontro”, ren- nii, Echinocactus horizonthalonius, etc. dendomi subito conto che, più di chiunque altra Dedicated to Mr. Francesco GUCCINI: his succulenta, “non perdona e tocca”. Non potevo words made me thinking many times during
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