The Nymphal Taxonomy and Systematics of Western Palaearctic Lygaeid Bugs (Hemiptera, Heteroptera, Lygaeidae), with Special Reference to the British Fauna Volume 2- Appendices Stephen Judd A thesis submitted in partial fulfilment of the requirements of Liverpooljohn Moores University for the degree of Doctor of Philosophy This research was carried out in collaboration with Liverpool Museum, National Museums and Galleries on Merseyside October 1994 APPENDICES 1- Summary of published nymphal information............ 1 2- British field 21 collecting sites ...................... 3 West Palaearctic field 32 - collecting sites ............. 4 Individuals for 46 - providing nymphs study .............. 5 Collection 47 - catalogue ................................ 6 - West Palaearctic genera not recorded during this 137 study ............................................... fifth instar 7 - Measurements of British Lygaeidae ...... 138 8 - Distribution information sources - biological 151 records centres and recording schemes ............... 9 - Distribution information sources - individual recorders and collectors ............................ 152 10 Distribution information 153 - sources - collections ...... 11 Distribution information literature 154 - sources - ....... 12 - Copy of distribution and catalogue file structure... 178 Appendix 1. Summary of Published Nymphal Information 5,4,3,2,1 - instarsj P- photograph; K- unspecified instare keyed Described Figured Keyed Lygaeidae Lygaeidae -- 5-4 De Courcey (1971) Lygaeidae -- 5-1 Dolling (1991) Lygaeidae --K Herring & Ashlock (1971) Lygaeidae --K Lawson (1959) Lygaeidae --K Leston & Scudder (1956) Lygaeidae 5-1 -K Puchkov (1969) Lygaeidae --K Slater & Saranoweki (1978a) Lygaeinae Lygaeinae - - 5-4 De Courcey (1971) Lygaeinae - - 5-3 Puchkov (1958) Lygaeinae - - K Puchkov (1969) Lygaeinae - - 5 Sweet & Slater (1961) Aethalotus horni Breddin, 1907 5-1 5-1 - Mukhopayay at a1. (1990) Arocatus Spinola, 1837 - - 5-3 Puchkov (1958) Arocatus Spinola, 1837 5-1 - K Puchkov (1969) Arocatus longiceps StAl, 1872 - - 5-3 Puchkov (1969) Arocatus melanocephalus (Fabricius, 1798) 5-1 3 K Puchkov (1969) Arocatus roeseli (Schilling, 1829) 5-1 - 5-3 Puchkov (1969) Arocatus rusticus (StAl, 1867) 5-1 5-1P Myers (1926) Aspilocoryphus fasciativentris (StAl, 1858) 3 - - Slater & Sperry (1973) Caenocoris marginatus (Thunberg, 1822) ? - - Kirkaldy (1907b) Craspedechus puichellus (Fabricius, 1794) 5-1 5-1 - Baranowski & Slater (1975) Graptostethus Distant, 1879 5-1 5-1 - Roy et al., (1989) Graptostethus sp. 5-1 - - Dhamdhere & Rawat (1967) Graptostethus sp. 5-1 5-1 - Hoffmann (1934) (Fabricius, Graptostethus servus 1787) 1 - - Slater & Sperry (1973) Haemobaphus concinnus (Dallas, 1852) 5-4,2-1? 5 K Slater & Sperry (1973) Horvathiolus delicatulus (StAl, 1855) 5-4 5 5 Slater & Sperry (1973) Horvathiolus longipilosus Slater & Sperry, 1973 5 - 5 Slater & Sperry (1973) (Pollich, Horvathiolus superbus 1779) 7 - - Horvath (1916) Horvathiolus Superbus (Pollich, 1779) 5-3 - K Puchkov (1969) Karachiocoris seidenstuckeri 5-1 5-1 - Hamid & Meher (1973) Lygaeosoma Spinola, 1837 - - 5-3 Puchkov (1958) Lygaeosoma Spinola, 1837 5-1 - K Puchkov (1969) Lygaeosoma anatolicum Seidenstricker, 1960 7 5 K Puchkov (1969) (1973) Lygaeosoma neglecta Distant, 1901 5 - 5 Slater & Sperry Lygaeosoma sardea Spinola, 1837 5-2 - K Puchkov (1969) (1969) Lygaeosoma sibiricum Seidenstricker, 1962 ? - K Puchkov (1961) Lygaeospilus Barber, 1921 - - 5 Sweet & Slater Lygaeus Fabricius, 1794 - - 5-3 Puchkov (1958) (1969) Lygaeus Fabricius, 1794 5-1 - K Puchkov Lygaeus Fabricius, 1794 -. - 5 Sweet & Slater (1961) Lygaeus equestrie (Linnaeus, 1758) 5-1 5 K Puchkov (1969) Lygaeus kalmii St&1,1874 5-1 5-1 - Simanton & Andre (1936) Lygaeus quadriguttatus (Fabricius, 1775) ? - - Froggatt (1901) Lygaeus turcicus Fabricius, 1803 5-1 - - Slater (1983a) Melanocoryphus StAl, 1872 - - 5-3 Puchkov (1958) Melanocoryphus Stal, 1872 5-1 - K Puchkov (1969) 1 Described Figured Keyed Melanocoryphus albomaculatus (Goeze, 1778) 5-1 4 K Puchkov (1969) Melanocoryphus tristrami (Douglas & Scott, 1868) 5-1 - K Puchkov (1969) Melanostethus marginatus (Thunberg, 1822) 5,1 5 5 Slater & Sperry (1973) Melanotelus villosulus (StAl, 1855) 5-1 5 5 Slater & Sperry (1973) Oncopeltus St&1,1868 - - 5 Slater (1951) Oncopeltus StAl, 1868 - - 5 Sweet & Slater (1961) Oncopeltus bergianus Kirkaldy, 1909 ? - - Berg (1883) Oncopeltus famelicus (Fabriciue, 1781) 5-3,1? 5 5 Slater & Sperry (1973) Oncopeltus fasciatus (Dallas, 1852) 5-1 - - Andre (1934) Oncopeltus fasciatus (Dallas, 1852) - 5P - Beck et al. (1958) Oncopeltue sordidus (Dallas, 1852) ? - - Froggatt (1901,1907) Orsillacis Barber, 1914 - - 5 Sweet & Slater (1961) Spilostethus furcula (Herrich-Schaeffer, 1850) 5-1 5 5 Slater & Sperry (1973) Spilostethus hospes (Fabricius, 1794) 5-1 5-1 - Hoffmann (1932) Spilostethus hospes (Fabricius, 1794) 5 - - Hoffmann (1934) Spilostethus lemniscatus (St&l, 1855) 5-4,1 5 5 Slater & Sperry (1973) SpIlostethus macilentus (StAl, 1874) 5 5 5 Slater & Sperry (1973) Spilostethus pandurus (Scopoli, 1763) 5-1 5-1 - Bhattacherjee (1959) Spilostethuspandurus (Scopoli, 1763) 5-4 - K Puchkov (1969) Spilostethus pandurus elegans (Wolff, 1802) 5-1 5 5 Slater & Sperry (1973) Spilostethus pandurus militaris (Fabricius, 1775) 5-1 5-1 - Hamid & Ahmed (1972) Spilostethus rivularis (Germar, 1837) 5-1 5 5 Slater & Sperry (1973) Spilostethus saxatilis (Scopoli, 1763) 5-2 5 K Puchkov (1969) Spilostethus taeniatus (St&l, 1865) 5-3 5 5 Slater & Sperry (1973) Spilostethus trilineatus (Fabricius, 1794) 5-4 5 5 Slater & Sperry (1973) Stenaptula angusticollis (Lindberg, 1939) 5 5 5 Slater & Sperry (1973) Tropi do thorax Bergroth, 1868 - - 5-3 Puchkov (1958) Tropidothorax Bergroth, 1868 - - K Puchkov (1969) Tropi do thorax Ieucopterus (Goeze, 1778) 5-1 - - Puchkov (1969) Orsillinae Orsillinae -- 5-4 De Courcey (1971) oraillinae 5-1 - K Puchkov (1969) BeIonochilus Uhler, 1871 -- 5 Slater (1951) Belonochilus Uhler, 1871 -- 5 Sweet & Slater (1961) Belonochilus numenius (Say, 1831) 5-1 5 - Wheeler (1984) Camptocoris rostratus Slater & Ashlock, 1980 5-4 - - Slater & Ashlock (1980) Camptocoris typus (Distant, 1918) 5- - Slater & Ashlock (1980) Lepionysius grossi Ashlock, 1967 5-1 5 5 Slater (1976a) Macroparius sg. Stil, 1872 -- 5-3 Puchkov (1958) Neseis fasciata fascists Usinger, 1942 5- K Usinger (1942) Neseis fulgida Usinger, 1942 5- K Usinger (1942) Neseis molokaiensis Usinger, 1942 5- K Usinger (1942) Neseis nitida pipturi Usinger, 1942 5- K Usinger (1942) Neseis oahuensis Usinger, 1942 1- - Usinger (1942) Neseis ochriasis maculiceps Usinger, 1942 5- K Usinger (1942) Nesomartis psammophila Kirkaldy, 1907 7- - Perkins (1913) (1958) Nithecus Horv&th, 1890 -- 5-3 Puchkov Nithecus jacobaeae (Schilling, 1829) 4-3 - - Puchkov (1969) Nysius Dallas, 1852 5-1 - K Puchkov (1969) Nyaius Dallas, 1852 -- 5 Slater (19S1) Nysius Dallas, 1852 -- 5 Sweet & Slater (1961) 2 Described Figured Keyed Nysius Uhler, angustatus 1872 -? - Chittenden (1903) Nysius angustatus Uhler, 1872 -? - Porbes (1905) Nysius angustatus Uhler, 1872 -? - Lugger (1900) Nysius coenosulus St&l, 1859 5-1 - K Usinger (1942) St&l, Nysius coensolus 1859 - 5-1 - Zimmerman (1948) Nysius delectulus Perkins, 1912 ?- - Kirkaldy (1907b) (Schilling, Nysius ericae 1829) - 4P - Burgess & Weegar (1986) Nysius ericae (Schilling, 1829) ?- - Knowlton & Wood (1943) Nysius ericae (Schilling, 1829) -5 - Milliken (1916) (Schilling, Nysius ericae 1829) -4 - Puchkov (1958) (Schilling, Nysius ericae 1829) -4 - Puchkov (1969) Nysius ericae (Schilling, 1829) -5 - Smith, J. B. (1910) Nysius fasciata Usinger, 1942 ?- - Zimmerman (1948) (Zetterstedt, Nysius groenlandicus 1840) 5-4,2-1 - - Lindberg (1935) Nysius inconspicuus Distant, 1904 11 - Miller (1934) Nysius kinbergi Usinger, 1959 -? - Zimmerman (1948) Nysius raphanus Howard, 1872 5-1 5-1 - Milliken (1918) Nysius rubescens Buchanan-White, 1881 5- K Usinger (1942) Nysius rubescens Buchanan-White, 1881 -5 - Zimmerman (1948) Nysius thymi (Wolff, 1804) -5 - Cobben (1953) Nysius thymi (Wolff, 1804) -5 - Puchkov (1969) Nysius thymi (Wolff, 1804) 5-1 4 - Puchkov & Puchkova (1956) Nysius thymi (Wolff, 1804) 5,? 5 - Sweet & Slater (1961) Nysius vinitor Bergroth, 1891 5- - Kirkaldy (1907a) Oceanides bryani Usinger, 1942 1- - Usinger (1942) Oceanides fosbergi Usinger, 1942 1- - Usinger (1942) Oceanides membranaceus Usinger, 1942 5- K Usinger (1942) Oceanides nimbatus (Kirkaldy, 1910) 5- K Usinger (1942). Oceanides nubicola (Kirkaldy, 1910) 5-1 - K Usinger (1942) Oceanides nubicola (Kirkaldy, 1910) 5-1 - - Zimmerman (1948) Orsillus Dallas, 1852 -- 5-3 Puchkov (1958) Orsillus Dallas, 1852 -- K Puchkov (1969) depressus Orsillus Dallas, 1852 5-3 - 5-4 Puchkov (1969) Orsillus maculatus (Fieber, 1861) -4 - Puchkov (1958) (Fieber, Orsillus maculatus 1861) -4 5-4 Puchkov (1969) Orsillus reyi Puton, 1871 -- 5-4 Puchkov (1969) Ortholomus St&1,1872 -- 5-3 Puchkov (1958) Ortholomus StAl, 1872 -- K Puchkov (1969) Ortholomus StAl, 1872 -- 5 Slater (1951) Ortholomus StAl, 1872 -- 5 Sweet & Slater (1961) (Herrich-Schaeffer, Ortholomus punctipennis 1838) 5-1 5-1 - Jordan (1933) Ortholomus punctipennis (Herrich-Schaeffer, 1838) 5-1 - - Puchkov (1969) Metraga nuda Buchanan-White, 1878 ?- - Kirkaldy (1907a) Metraga nuda Buchanan-White, 1878 ?- - Zimmerman (1948) Ischnorhynchinae Ischnorhynchinae -- 5-4 De Courcey (1971) Ischnorhynchinae -- K Puchkov (1969) (1961) Ischnorhynchinae -- 5 Sweet & Slater Kleidocerys Stephens, 1829 -- 5-3 Puchkov (1958) (1923) Kleidocerys resedae