Family: Moraceae Taxon: Artocarpus nitidus Synonym: Artocarpus lingnanensis Merr. Common Name: ma hat Artocarpus parvus Gagnep. hung kwai muk Artocarpus sampor Gagnep. kwai muk Questionaire : current 20090513 Assessor: Assessor Designation: EVALUATE Status: Assessor Approved Data Entry Person: Assessor WRA Score 3 101 Is the species highly domesticated? y=-3, n=0 n 102 Has the species become naturalized where grown? y=1, n=-1 103 Does the species have weedy races? y=1, n=-1 201 Species suited to tropical or subtropical climate(s) - If island is primarily wet habitat, then (0-low; 1-intermediate; 2- High substitute "wet tropical" for "tropical or subtropical" high) (See Appendix 2) 202 Quality of climate match data (0-low; 1-intermediate; 2- High high) (See Appendix 2) 203 Broad climate suitability (environmental versatility) y=1, n=0 n 204 Native or naturalized in regions with tropical or subtropical climates y=1, n=0 y 205 Does the species have a history of repeated introductions outside its natural range? y=-2, ?=-1, n=0 n 301 Naturalized beyond native range y = 1*multiplier (see y Appendix 2), n= question 205 302 Garden/amenity/disturbance weed n=0, y = 1*multiplier (see n Appendix 2) 303 Agricultural/forestry/horticultural weed n=0, y = 2*multiplier (see n Appendix 2) 304 Environmental weed n=0, y = 2*multiplier (see n Appendix 2) 305 Congeneric weed n=0, y = 1*multiplier (see n Appendix 2) 401 Produces spines, thorns or burrs y=1, n=0 n 402 Allelopathic y=1, n=0 403 Parasitic y=1, n=0 n 404 Unpalatable to grazing animals y=1, n=-1 405 Toxic to animals y=1, n=0 n 406 Host for recognized pests and pathogens y=1, n=0 y 407 Causes allergies or is otherwise toxic to humans y=1, n=0 n 408 Creates a fire hazard in natural ecosystems y=1, n=0 n 409 Is a shade tolerant plant at some stage of its life cycle y=1, n=0 y 410 Tolerates a wide range of soil conditions (or limestone conditions if not a volcanic island) y=1, n=0 Print Date: 9/11/2013 Artocarpus nitidus (Moraceae) Page 1 of 7 411 Climbing or smothering growth habit y=1, n=0 n 412 Forms dense thickets y=1, n=0 n 501 Aquatic y=5, n=0 n 502 Grass y=1, n=0 n 503 Nitrogen fixing woody plant y=1, n=0 n 504 Geophyte (herbaceous with underground storage organs -- bulbs, corms, or tubers) y=1, n=0 n 601 Evidence of substantial reproductive failure in native habitat y=1, n=0 n 602 Produces viable seed y=1, n=-1 y 603 Hybridizes naturally y=1, n=-1 604 Self-compatible or apomictic y=1, n=-1 605 Requires specialist pollinators y=-1, n=0 606 Reproduction by vegetative fragmentation y=1, n=-1 n 607 Minimum generative time (years) 1 year = 1, 2 or 3 years = 0, >3 4+ years = -1 701 Propagules likely to be dispersed unintentionally (plants growing in heavily trafficked y=1, n=-1 areas) 702 Propagules dispersed intentionally by people y=1, n=-1 y 703 Propagules likely to disperse as a produce contaminant y=1, n=-1 n 704 Propagules adapted to wind dispersal y=1, n=-1 n 705 Propagules water dispersed y=1, n=-1 706 Propagules bird dispersed y=1, n=-1 y 707 Propagules dispersed by other animals (externally) y=1, n=-1 708 Propagules survive passage through the gut y=1, n=-1 y 801 Prolific seed production (>1000/m2) y=1, n=-1 n 802 Evidence that a persistent propagule bank is formed (>1 yr) y=1, n=-1 n 803 Well controlled by herbicides y=-1, n=1 804 Tolerates, or benefits from, mutilation, cultivation, or fire y=1, n=-1 805 Effective natural enemies present locally (e.g. introduced biocontrol agents) y=-1, n=1 Designation: EVALUATE WRA Score 3 Print Date: 9/11/2013 Artocarpus nitidus (Moraceae) Page 2 of 7 Supporting Data: 101 2003. Wu, Z.Y./Raven,P.H./Hong, D.Y. (eds.). [Is the species highly domesticated? No] No evidence Flora of China. Vol. 5 (Ulmaceae through Basellaceae). Science Press & Missouri Botanical Garden Press, Beijing & St. Louis 102 2013. WRA Specialist. Personal Communication. NA 103 2013. WRA Specialist. Personal Communication. NA 201 2003. Wu, Z.Y./Raven,P.H./Hong, D.Y. (eds.). [Species suited to tropical or subtropical climate(s) 2-High] "Mixed forests. Flora of China. Vol. 5 (Ulmaceae through Guangdong, Guangxi, Hainan, S Hunan, S Yunnan [cultivated in Cambodia, Basellaceae). Science Press & Missouri Botanical Thailand, and N Vietnam]." Garden Press, Beijing & St. Louis 202 2003. Wu, Z.Y./Raven,P.H./Hong, D.Y. (eds.). [Quality of climate match data 2-High] Flora of China. Vol. 5 (Ulmaceae through Basellaceae). Science Press & Missouri Botanical Garden Press, Beijing & St. Louis 203 1960. Jarrett, F. M.. Studies in Artocarpus and [Broad climate suitability (environmental versatility)? Elevation range almost allied genera, IV. A revision of Artocarpus sub- exceeds 1000 m] "in evergreen and mixed forest, and in savannah, to 3000 ft.; genus Pseudojaca (concluded). Journal of the Siam, Indochina, southern China (Kwangtung, Hainan); cultivated, especially in Arnold Arboretum. 41(2): 111-140. Tonkin." 203 1996. Schulte, A./Hans-Friedrich Schöne, D. [Broad climate suitability (environmental versatility)? Presumably No] "Artocarpus (eds.). Dipterocarp Forest Ecosystems: Towards dadah, Artocarpus nitidus, and Artocarpus rigidus are common trees especially in Sustainable Management. World Scientific, the lowlands and hills up to 600 m altitude." [Presumably restricted to lower London, UK elevation, tropical climates] 203 2013. Nickrent, D.L./Barcelona, J./Pelser, [Broad climate suitability (environmental versatility)? No. Species restricted to low P./Molina, J.E./Callado, J.R.. Co's Digital Flora of elevation tropics] "Lowland primary and secondary forests, in regions with rather the Philippines. http://www.philippineplants.org/ marked dry seasons, 0 150m. Four other subspecies extend from Assam to continental SE Asia, S China to W Malesia (Sumatra, Peninsular Malaysia, and Borneo)." 204 2003. Wu, Z.Y./Raven,P.H./Hong, D.Y. (eds.). [Native or naturalized in regions with tropical or subtropical climates? Yes] "Mixed Flora of China. Vol. 5 (Ulmaceae through forests. Guangdong, Guangxi, Hainan, S Hunan, S Yunnan [cultivated in Basellaceae). Science Press & Missouri Botanical Cambodia, Thailand, and N Vietnam]." Garden Press, Beijing & St. Louis 205 1992. Flynn, T./Ragone, D.. Specimen Details for [Does the species have a history of repeated introductions outside its natural Artocarpus nitidus subsp. lingnanensis (Merr.) range? No] "USA - Hawaii - Maui - East Maui, Hana District, cultivated the Jarrett [BISH 634527]. Bishop Museum, National Tropical Botanical Garden's Kahanu Garden." [Sparingly cultivated in the http://nsdb.bishopmuseum.org/5681216E-DE32- Hawaiian Islands] 4BB3-9DA8-2C1D15BABF3D 205 2005. Staples, G.W./Herbst, D.R.. A Tropical [Does the species have a history of repeated introductions outside its natural Garden Flora - Plants Cultivated in the Hawaiian range? No evidence] "Locally grown plants belong to subsp. lingnanensis (Merrill) Islands and Other Tropical Places. Bishop F.M. Jarrett, native from southern China throughout Southeast Asia." … "Trees are Museum Press, Honolulu, HI cultivated in Asia for their edible fruit, and the bark and roots are added to betel. Kwai muk is said to have been introduced to Hawaii around 1960 from southern China. It is sparingly grown elsewhere." 301 2012. Randall, R.P.. A Global Compendium of [Naturalized beyond native range? No evidence elsewhere] Weeds. 2nd Edition. Department of Agriculture and Food, Western Australia 301 2013. Kaneholani, L.. KISC Field Crew [Naturalized beyond native range? Possibly Yes] "may be Artocarpus nitidus var. Supervisor. Pers. Comm. 09 Sep 2013. lingnanensis" … "there are lots of seedling popping up around her home in the Wailua/Kapahi area" [Showing signs of naturalization on Kauai] 302 2012. Randall, R.P.. A Global Compendium of [Garden/amenity/disturbance weed? No evidence] Weeds. 2nd Edition. Department of Agriculture and Food, Western Australia 303 2012. Randall, R.P.. A Global Compendium of [Agricultural/forestry/horticultural weed? No] No evidence Weeds. 2nd Edition. Department of Agriculture and Food, Western Australia 304 2012. Randall, R.P.. A Global Compendium of [Environmental weed? No] No evidence Weeds. 2nd Edition. Department of Agriculture and Food, Western Australia 305 2012. Randall, R.P.. A Global Compendium of [Congeneric weed? No. Several Artocarpus species are listed as naturalized, but Weeds. 2nd Edition. Department of Agriculture none are regarded as a serious agricultural or environmental weed] and Food, Western Australia Print Date: 9/11/2013 Artocarpus nitidus (Moraceae) Page 3 of 7 401 2003. Wu, Z.Y./Raven,P.H./Hong, D.Y. (eds.). [Produces spines, thorns or burrs? No] "Trees to 17 m tall. Petiole 0.5-1.5 cm; leaf Flora of China. Vol. 5 (Ulmaceae through blade oblongelliptic to obovate-elliptic, 7-15 × 3-7 cm, thinly leathery, base Basellaceae). Science Press & Missouri Botanical cuneate to ± rounded, margin entire or irregularly shallowly toothed, apex Garden Press, Beijing & St. Louis mucronate to caudate; secondary veins 6-10 on each side of midvein, adaxially prominent." 402 2003. Fujii, Y./Parvez, S. S./Parvez, [Allelopathic? Unknown for A. nitidus. Artocarpus heterophyllus & Artocarpus altilis M.M./Ohmae, Y./Iida, O.. Screening of 239 tested for allelopathic properties in lab conditions, but incisory activity not greater medicinal plant species for allelopathic activity than the mean] using the sandwich method. Weed Biology and Management. 3: 233–241. 403 2003. Wu, Z.Y./Raven,P.H./Hong, D.Y. (eds.). [Parasitic? No] "Trees to 17 m tall." [Moraceae] Flora of China. Vol. 5 (Ulmaceae through Basellaceae). Science Press & Missouri Botanical Garden Press, Beijing & St.