British Scarabs Guide to Aphodiini (Dweller dung beetles) ACROSSUS BODILOPSIS MELINOPTERUS MELINOPTERUS MELINOPTERUS MELINOPTERUS rufipes rufa consputus prodromus punctatosulcatus sphacelatus 9-13 mm D: 0-9 5-7 mm D: 0-9 3-6 mm D- 5-7 4-7 mm D: 0-9 4-6 mm D: 7 4-6 mm D: 0-9 All habitats All habitats Well drained soils All habitats Grasslands All habitats J F M A M J J A S O N D J F M A M J J A S O N D J F M A M J J A S O N D J F M A M J J A S O N D J F M A M J J A S O N D J F M A M J J A S O N D NIMBUS NIMBUS VOLINUS ACROSSUS CHILOTHORAX obliteratus contaminatus sticticus luridus conspurcatus 4-7 mm D: 0-8 5-7 mm D: 0-9 4-6 mm D: 0, 2-9 6-9 mm D: 0-9 4-5 mm D: 0-8 Shaded habitats All habitats Shaded habitats Well drained soils Shaded habitats J F M A M J J A S O N D J F M A M J J A S O N D J F M A M J J A S O N D J F M A M J J A S O N D J F M A M J J A S O N D CHILOTHORAX CHILOTHORAX EUHEPTAULACUS EUHEPTAULACUS LABARRUS distinctus paykulli sus villosus lividus 3-6 mm D: 1-8 3-5 mm D: 1-8 4-5 mm D: 6-8 3-5 mm D: 1-8 3-6 mm D: 5-8 Prefers sandy soils Well drained soils Well drained soils Well drained soils Well drained soils J F M A M J J A S O N D J F M A M J J A S O N D J F M A M J J A S O N D J F M A M J J A S O N D J F M A M J J A S O N D BODILOIDES ESYMUS BODILOPSIS APHODIUS COLOBOPTERUS ictericus merdarius sordida foetidus erraticus 3-6 mm D: 0-9 4-5 mm D: 0-9 5-8 mm D: 0-8 5-8 mm D: 0-9 6-9 mm D: 0-9 Well drained soils Grasslands Well drained soils Grasslands Grasslands J F M A M J J A S O N D J F M A M J J A S O N D J F M A M J J A S O N D J F M A M J J A S O N D J F M A M J J A S O N D APHODIUS APHODIUS RHODAPHODIUS AGOLIINUS SIGORUS fimetarius pedellus foetens lapponum porcus 5-8 mm D: 0-9 5-8 mm D: 0-8 6-9 mm D: 0-9 4-6 mm D: 0-4 4-6 mm D: 0-8 All habitats All habitats Grasslands Upland habitats Grasslands J F M A M J J A S O N D J F M A M J J A S O N D J F M A M J J A S O N D J F M A M J J A S O N D J F M A M J J A S O N D LIMARUS PLANOLINUS PLANOLINUS OTOPHORUS PHALACRONOTHUS zenkeri borealis fasciatus haemorrhoidalis quadrimaculatus 4-6 mm D: 2-8 3-6 mm D: 0-8 3-5 mm D: 0-4 3-6 mm D: 2-9 3-4 mm D: 4-7 Shaded habitats Shaded habitats Upland, in shade Grasslands Well drained soils J F M A M J J A S O N D J F M A M J J A S O N D J F M A M J J A S O N D J F M A M J J A S O N D J F M A M J J A S O N D PLAGIOGONUS AGOLIINUS AGRILINUS AGRILINUS EUORODALUS arenarius nemoralis ater constans coenosus 3-5 mm D: 5-7 4-6 mm D: 1-3 4-6 mm D: 0-9 5-6 mm D: 1-8 3-5 mm D: 3-9 Chalk soils Upland, in shade All habitats Shaded habitats Well drained soils J F M A M J J A S O N D J F M A M J J A S O N D J F M A M J J A S O N D J F M A M J J A S O N D J F M A M J J A S O N D ESYMUS CALAMOSTERNUS LIOTHORAX LIOTHORAX ACROSSUS pusillus granarius plagiatus niger depressus 3-5 mm D: 0-9 3-5 mm D: 0, 2-9 4-5 mm D: 0-9 4-5 mm D: 7 6-9 mm D: 0-8 Grasslands Grasslands Coastal, organic debris Poolside debris All habitats J F M A M J J A S O N D J F M A M J J A S O N D J F M A M J J A S O N D J F M A M J J A S O N D J F M A M J J A S O N D TEUCHESTES AMMOECIUS OXYOMUS HEPTAULACUS Peak activity Active Absent D: Distribution fossor brevis sylvestris testudinarius 6-12 mm D: 0-9 3-4 mm D: 2 3-4 mm D: 1-9 3-4 mm D: 7 Grasslands Sand dunes Organic debris Sandy soils J F M A M J J A S O N D J F M A M J J A S O N D J F M A M J J A S O N D J F M A M J J A S O N D Many thanks to OUMNH for use of images https://www.oumnh.ox.ac.uk Once you have identified your beetle, visit britishscarabs.org species page to check the details. Don’t forget to submit your sightings to the Recording Scheme via iRecord www.britishscarabs.org (www.brc.ac.uk/irecord) v1.2.