SPANISH JOURNAL OF PALAEONTOLOGY Pseudolillia Maubeuge, 1949 (Ammonitida, Hildoceratidae) in the Lower Jurassic (Toarcian) of the NE Spain Miguel MORATILLA-GARCÍA1, Antonio GOY2* & María José COMAS-RENGIFO2 1 Jesús García Perdices, nº 3, 4ºA, 19004 Guadalajara, Spain; [email protected] 2 Departamento de Paleontología, Facultad de Ciencias Geológicas, Universidad Complutense de Madrid, José Antonio Novais, 12, 28040 Madrid, Spain; [email protected]; [email protected] * Corresponding author Moratilla-García, M., Goy, A. & Comas-Rengifo, M.J. 2017. Pseudolillia Maubeuge, 1949 (Ammonitida, Hildoceratidae) in the Lower Jurassic (Toarcian) of the NE Spain. [Pseudolillia Maubeuge, 1949 (Ammonitida, Hildoceratidae) en el Jurásico Inferior (Toarciense) del NE de España]. Spanish Journal of Palaeontology, 32 (1), 147-170. Manuscript received 1 December 2016 © Sociedad Española de Paleontología ISSN 2255-0550 Manuscript accepted 4 April 2017 ABSTRACT RESUMEN In the present paper, 147 specimens assigned to the Se han estudiado 147 ejemplares asignados al género genus Pseudolillia Maubeuge, 1949 are studied. This is a Pseudolillia Maubeuge, 1949, lo que es un número considerably high number of samples in comparison with considerablemente elevado si se compara con los conocidos those known in other geographical areas where they have en otras áreas geográfi cas donde han sido citados, como el been cited, such as northern France, the Pyrenees, the Betic norte y centro de Francia, los Pirineos, la Cordillera Bética, Range, Morocco, Portugal, Italy, Hungary and Bulgaria. The Marruecos, Portugal, Italia, Hungría y Bulgaria. Proceden six taxa described, P. murvillensis, P. hispanica, P. emiliana, de un total de 22 localidades de las cordilleras Cantábrica, P. donovani, Pseudolillia ? n. sp. (en García-Gómez & Ibérica, Isla de Mallorca y Pirineos. Se han descrito seis Rivas, 1980), and Pseudolillia ? sp., come from 22 sites in taxones: P. murvillensis, P. hispanica, P. emiliana, P. donovani, the Cantabrian and Iberian Ranges and the Isle of Majorca. Pseudolillia ? n. sp. (en García-Gómez & Rivas, 1980) y Their presence in expanded sections enabled us to situate the Pseudolillia ? sp. Su presencia en secciones expandidas, ha stratigraphic position of the genus between the upper part of permitido situar la posición estratigráfi ca del género desde the Thouarsense Chronozone (Fallaciosum Subchronozone) la parte alta de la Cronozona Thouarsense (Subcronozona to the Dispansum Chronozone (Gruneri Subchronozone), with Fallaciosum) hasta la Cronozona Dispansum (Subcronozona maximum abundance found in the Dispansum Chronozone Gruneri), encontrándose la máxima abundancia en la (Insigne Subchronozone). The origin of the genus is unclear, Cronozona Dispansum (Subcronozanona Insigne). El origen and a possible derivation from the last representatives of del género no es del todo claro, y ha sido sugerida una posible Grammoceras (close to G. striatulum) or more likely from derivación de miembros tardíos de Grammoceras (próximos a Esericeras (close to E. fascigerum) has been suggested. The G. striatulum) o más probablemente de Esericeras (próximos known palaeobiogeographic range of Pseudolillia is limited a E. fascigerum). La distribución paleobiogeográfi ca conocida to the NW-European Province and to the W of the Tethys, de Pseudolillia se limita a la provincia NO-Europea y a la and with the exception of P. emiliana, the species’ distribution parte O del Tethys y, con excepción de P. emiliana, las áreas ranges are short. In NE Spain, where the highest number de distribución de las especies son relativamente pequeñas. 148 MORATILLA-GARCÍA, GOY & COMAS-RENGIFO of known specimens comes from, P. donovani (typical of En el NE de España, de donde proceden el mayor número de the Mediterranean province) is the species presenting the especímenes conocidos, P. donovani (típica de la Provincia oldest record (Chronozone Thouarsense, Subchronozone Mediterránea) es la especie que tiene un registro más antiguo Fallaciosum), although it did not necessarily live and (Cronozona Thouarsense, Subcronozona Fallaciosum), aunque reproduce in very nearby areas. P. murvillensis (typical of se supone que no necesariamente vivía y se reproducía en the NW-European Province), was found in the Chronozone áreas muy cercanas. P. murvillensis (típica de la Provincia Dispansum (Subchronozone Insigne) together with P. emiliana NO-Europea), ha sido encontrada en la Cronozona Dispansum and P. hispanica in the eastern Pyrenees and particularly in (Subcronozona Insigne) junto con P. emiliana y P. hispanica the central sector of the Iberian Range. en los Pirineos orientales y, sobre todo, en el sector central de la Cordillera Ibérica. Keywords: Upper Toarcian, Grammoceratinae, taxonomy, palaeobiogeography, Iberian Peninsula. Palabras clave: Toarciense superior, Grammoceratinae, taxonomía, paleobiogeografía, Península Ibérica. 1. INTRODUCTION Grammoceratinae. Subsequently, García-Gómez & Rivas (1980) and Elmi & Rulleau (1990), based upon the new The genus Pseudolillia Maubeuge, 1949, from the upper species recognised, also emended the genus. These authors, Toarcian in Europe and the N of Africa, extends mainly along with Howarth (2013) maintained Pseudolillia within from Lorraine, in the NE of France, to the Atlas Mountains Grammoceratinae. in Morocco, with some occasional references in Alpine García-Gómez & Rivas (1980) studied the genus areas. The record tends to be scarce or very scarce in in the Betic Range and described two new species, most of the sites where it has been cited, and it is the Pseudolillia donovani, Pseudolillia ? n. sp., and one Iberian Peninsula, especially in the central sector of the new subspecies, Pseudolillia murvillensis hispanica. Iberian Range, where most of the known specimens were Moreover, they extended the information on the diagnosis found. In the present paper we study the available material of the genus and on the species P. emiliana. In their of the species of the genus Pseudolillia recorded in the opinion, the ribbing is not always straight on the fl ank, outcrops of the upper Toarcian in the E and N of Spain. but can present a sinusoidal trajectory similar to that of We attempt to conduct a systematic classifi cation and to Pseudogrammoceras. In relation to the coiling, although specify the precise stratigraphic position of the species. Our Maubeuge (1949) considered that the umbilicus undergoes research also aims to establish the palaeobiogeographic a relative decrease in size during ontogeny, he pointed out distribution of each of them. We paid particular attention that this occurs in some species, but not in others such to the Iberian Range, collecting new samples in its as P. donovani. Furthermore, the shape of the section central sector, especially in the localities of Ribarredonda, can also show no variation during ontogeny. The above- Huertahernando, Villar de Cobeta, Buenafuente del Sistal mentioned authors suggested that this genus presents and Turmiel in the province of Guadalajara, and in the close affi nities with Esericeras, and it is therefore related Rambla del Salto and Ariño in the province of Teruel. with Grammoceratinae, in accordance with the opinion of Also included in the present research are the specimens Donovan (1962). identifi ed by previous authors, some specimens from the Elmi & Rulleau (1990) conducted a revision of the collection of the Geominero Museum (IGME) and from the genus Pseudolillia in south-western Europe and Morocco, Asturias Jurassic Museum (MUJA) and those from the Isle which comprises all the species on our research. They of Majorca (Álvaro et al., 1989) and from the Cantabrian described and identified important material from the Range (Suárez Vega, 1974; Bernad, 1992, 1993; Goy et quarries of Lafargue in Belmont (Bas Beaujolais, Rhône, al., 1994, 2010). France), together with some specimens from Morocco, The genus Pseudolillia was created by Maubeuge the Betic Range and the Iberian Range. They also (1949) based upon one single specimen found in Murville considered highly likely that this genus belongs to the Mine (Meurtheet Moselle, Lorraine, France), which was Grammoceratinae. called Pseudolillia murvillensis. This author included Sassaroli & Venturi (2005) described a new species, Pseudolillia in the Subfamily Phymatoceratinae, as Pseudolillia apenninica, based on one single specimen would later be done by Arkell et al. (1957). Donovan found in levels of the Bonarelli Zone in the Marconessa (1962) revised the genus and included therein the species Quarry (Cingoli, Central Appenines, Italia). Metodiev Ammonites emilianus, described by Reynès (1868), (2008) studied the genus Pseudolillia in several localities of which he provisionally incorporated into the Subfamily the W of Bulgaria and cited it as coming from levels of the Fallaciosum Zone of the Bulgarian scale (= Thouarsense PSEUDOLILLIA MAUBEUGE, 1949 (AMMONITIDA, HILDOCERATIDAE) IN THE LOWER JURASSIC (TOARCIAN) OF THE NE SPAIN 149 Zone, Fallaciosum Subzone of the standard scale). Kovács (2007) in the Lusitanian Basin (Portugal), García-Gómez (2013), in his study on the Grammoceratinae of the & Rivas (1980) in the S of el Zegrí and Cerro Méndez Gerecse Mountains (Hungary) described a new species (Granada) and Caracuel et al. (2004) in Cerro de la Cruz of Pseudolillia, P. paralleliformis, from the lower part of (Sierra de Reclot Alicante). the Speciosum Zone. In addition, in the N and E of the Iberian Peninsula, this genus has been cited in general studies, among