A revision of the genus Dillenia by R.D. Hoogland (Flora Malesiana, Leyden) (Pro-issued separately 16.1. 19:52) Contents Pago Introduction 2 Dillenia L. 1753 . 3 6 History of the genus Relationships 6 within the 8 Relationships genus Morphological notes H 1. Stilt-roots 11 2. Petiolar wings 11 3. Vernation 11 12 4. Hcterophylly 5. Inflorescence 12 6. Corolla 12 7. Androoeiuin 12 14 8. Gynoecium 15 9. Fruit and seed Anatomy 15 Geographical distribution 15 Ecology 15 1' Vernacular names Uses 17 Miscellaneous notes 17 18 Key to the species 21 Species-descriptions and notes 134 Doubtful species Excluded species 135 136 Invalidly published names Index to collectors' numbers 137 Index to scientific names 142 Introduction The is an extension of revision of the present paper my Malaysian of the Dillenia L. Wormia Rottb. inserted in species genus ( included) the in the Flora revision of the Dilleniaceae Malesiana ser. I, vol. 4, in December 1951. A critical revision of part 3, pp. 141—174, published whole the genus has never been published before; the unfortunate result of this has been that the delimitation of Dillenia and Wormia, usually as has distinct genera, been based on different characters by various authors. The extension of the revision for the Flora Malesiana so as to include the extra-malaysian species enabled me to study a number of species, the which confirmed me in that the characters knowledge of certainly my idea been on which Dillenia and Wormia had separated before are certainly not the primary characters, to be used in the taxonomic treatment of the genus. All specimens and literature mentioned in this work have been examined indicated by me, unless otherwise; excepted are the specimens of the U.S. National Herbarium., of which I have only examined those collections, of which no duplicates were available from other herbaria. Particulars, not to be taken from the herbarium specimens themselves, taken such as habit, height, diameter, colour, etc., have been from the the collectors’ notes and, as far as reliable, from literature, and are in- in if serted the descriptions; there are contradictory data, they are dis- cussed under the Notes. A great amount of material, kindly, lent by various herbaria to the Rijksherbarium, Leyden, made it possible to mc to carry out this study. In the citation of the specimens these herbaria are indicated by the in Chronica Botanica 142— abbreviations, proposed (by Lanjouw 5, 1939, pp. 150; with a single addition (LAB) they are: A Arnold Anborotum Herbarium, Harvard University, Jamaica Plain, Mass.. BM Department of Botany, British Museum (Natural History), London. BE Herbier du Jardin Botanique de l'fitat, Bruxelles. BRI Botanical Museum and Herbarium, Brisbane. BRfSL Botanical Institute, Wroclaw (formerly Breslau). BZ Herbarium Bogoricnse, Bogor, .Java. C Universitetets Botaniskc Museum, Copenhagen. CAL The Royal Botanic Garden and Herbarium, Calcutta. CGE Botanical Museum and Herbarium of the University, Cambridge (England). DD Forest Research Institute Herbarium, Dehra Dun. E Herbarium of the Royal Botanic Garden, Edinburgh. PI Erbario del Instituto Botanico dell' Universita, Firenzc. G Institut de Botanique SystiSmatique de l'Universite, Geneve. GH Gray Herbarium, Harvard University, Cambridge, Mass.. GL Botanical Department Herbarium, Glasgow. GRO Botanisch Laboratorium der Rijksuniversiteit, Afdeling Plantensystcmatiek, Groningen. R. I). Hocx; land : A revision of the genus Dillenia 3 IPI Herbarium of the Imperial Forestry Institute, Oxford. K Herbarium, Royal Botanic Gardens, Kcw, Surrey. L Rijksherbarium, Leiden. LAE The Papua and New Guinea Herbarium, Lae. LINN Herbarium of the Linnean Society of London, London. M Botanischo Anstalten, Miinchen. MEL Botanic Gardens and National Herbarium, Melbourne. MICH Herbarium of the University of Michigan, Ann Arbor, Mich.. MO Herbarium of the Missouri Botanical Garden, St Louis, Mo.. NY New Herbari um of the York Botanical Garden, New York, N. Y.. OXF Oxford University Herbarium, Oxford. P Laboratoire do Phanerogamic du Musfium National d'Histoire Naturelle. Paris. Naturhistoriska Riksmuseet, Stockholm. SING Herbarium of the Botanic Gardens, Singapore. Botanisch Museum en Herbarium, Utrecht. UC University of California Herbarium, Berkeley, Calif.. UPS Botaniska Muscet, Uppsala. US United States National Herbarium, Washington, B.C.. I wish here indebtedness directors to express ray to the and keepers of these herbaria for their valuable help in putting their specimens at my disposal. A special word of thanks is due to the directors and staffs of those herbaria and libraries which 1 have personally visited, for the hospitality offered and assistance rendered to viz. me during my stay, at Brussels, Cambridge, Kew, Condon, Oxford, and Paris. I also am highly indebted to the "Ponds Vollenhoven", which enabled me to visit Paris and Brussels, and to the British Council, which offered visit me a grant enabling me to the English herbaria mentioned above and made the arrangements for this visit. My most sincere thanks are due to Prof. Dr H, J. Lam, Director of the Rijksherbarium, Leyden, for his support and for the hospitality shown to me in his and institute, to Prof. Dr C. G. G. J. van Steenis, director °f the Foundation Flora Malesiana, Leyden, for the freedom he allowed me to extend revision of the my Malaysian species of Dillenia to the present monograph. To both of them and to the members of the staffs of the Rijksherbarium and the Flora Malesiana 1 feel greatly indebted.' The figures are from the able hands of the draughtsman H. J. T. Tammel. DILLENIA L. 1753 J Iovt. Mai. A Fam. d. PI. 364. Syalita Bheedo, 3, 1683, p. 39; dans., 2, 17C3, p. Dillenia Hort. PI. Gen. PI. L., Cliff., 1737, p. 321; L., Sp. 1, 1753, p. 536; L., C(l- 239; 5, 1754, p. Tliunb., Trans. Linn. Soc. 1, 1791, p. 198; DC., Syst. 1, 1818, P- Prod. 1, 1824, 435; DC., p. 75; Buck.-11am., Trans. Linn. Soc. Lond. 15, 182(1, p. 99; G. Don, Gen. Ilist. Diehl. PI. FI. Ind. ed. .1, 1831, p. 77; Koxb., Carey 2, 1832, p. (.'DO; Prod. Fl. Wight & Arn., Pen. Ind. Or. Hook.f. & Thorns., Fl. Ind. 1, 1834, p. 5; 1, Fl. Ind. !855, p. 69; Miq., Bat. 1, 2, 1859, p. 11; Miq., Fl. Ind. Bat. Suppl. 1, 1860, 1>- 619; Benth. & Gen. PI. Handb. Ind. Fl. 152, 618, Hook.f., 1, 1862, p. 13; Drury, 1, 1864, Adansonia Ann. Mas. Bot. Bat. p. 9; Baill., 6, 1865'—6, p. 281 ; Miq., Lugd. 4, 1868, Hist. d. PI. Hook.f. & Fl. Br. Ind. p. 79; Baill., 1, 1867—9, p. 110; Thorns., 1, 1872, For. Fl. N.W. & C. For. Fl. Br. Burma p. 36; Brandis, India, 1874, p. 1; Kurz, 1, Fl. For. Cochincln 5 Fam. & 1877, p. 18; Pierre, 1, 1879, pi. —14; Vidal, Sinops. Gen. P], Lea. Filip., 1883, p. 24; Martelli in Becc., Malesia 3, 1886, p. .154; King, J- As. Soc. Fl. Ned. Beng. 58, II, 1889, p. 366; Bocrl., Handl. Ind. 1, 1890, p. 8; Watt, Diet. Prod. India Handb. Fl. Keen. 3, 1890, p. 112; Trimen, C'eyl. 1, 1893, p. 12; Koord. & Booms. in & Nat. Pfl. Val., Bijdr. 1 Java, 1894, p. 160; Gilg Engl. Prantl, Lam. Bot. Jahrb. 3, 6, 1895, p. 123; Priteel, 24,'1897, p. 354; Brandis, Ind.-Forester 4 BLUMBA VOL. VII, No. 1, 1952 Man. Ind. Timb. 2nd 26, 19'00, p. 429; Gamble, od., 1902, p. 4; Duthie, Fl. Upp. PI. Ind. Fin. & Gang. 1, 1903, p. 20; Brandis, Trees, 1906, p. 3; Gagncp., Bull. Soc. & Bot. Fr. Mem. 4, 1906, p. 5; Moll Janssonius, Mikrogr. Holz. Java Baumarten 1, Fin. & Fl. Gen. Ind.-Ch. 1906, p. 71; Gagnep., 1, 1907, p. 16; llid]., J. Str. Br. K. A. S. Sclioolfl. 54, 1910, p. 7; Back., Java, 1911, p. 10; Svedelius, Svensk Bot. Tidskr. Exk. Fl. Java Mus. J. 5, 1911, p. 152.; Koord., 2, 1912, p. 601; Ridl., Saraw. Fl. Pres. Madras Bot. Bihar & Orissa 1, .1913, p. 70; Gamble, 1, 1915, p. 7; Haines, Bibl. En. Born. 2, 1921, p. 6; Merr., PL, 1921, p. 382; Diels, Bot. Jahrb. 57, 1922, Fl. Mai. Pen. En. Fl. p. 436; Ridl., 1, 1922, p. 10; Merr., Philip. PI. 3, 1923, p. 59; For. Fl. Andam. Fl. Siam. En. Parkinson, Isls, 1923', p. 71; Craib, 1, 1925, p. 21; & Werderm. im & Prantl, Nat. Pfl. Fam. 2nd Gilg Engl. ed., 21, 1925, p. 35; Foxworthy, Mai. For. Ree. 147; Pears. & Comm. Timb. India 3, 1927, p. Brown, 1, 1932, p. 1; & Fl. Assam Diet. Kanjilal, Kanjilal, Das, 1, 1934, p. 10; Burk., Econ. Prod. Mai. Pen., Ind. Forester Fl. Gen. Ind.-Ch. 1935, p. 80S; Parkinson, 61, 1935, p. 450; Gagnep., Suppl. S. 1, 1938, p. 20; Com., Gard. Bull. S. 10, 1939, p. 3; Corn., Wayside Trees Malaya, 1940, Bekn. Fl. Java fam. p. 203; Back., (em. ed.) 4, 1942, 80, p. 4; Masamune, En. Phan. Fl. Nutt. PI. Born., 1942, p. 462; Masamune, Ivainantensis, 1943, p. 205; Heync, Indon., Fl. Mai. 154. 1950, p. 1071; Hoogl., I, 4, 1951, p. Wormia Samiml. Vidensk. Selsk. Skrift. Vormia Rottb., Nye 2, 1783, p. 531, non Fam. d. PI. Wormia Nat. Selsk. Adans., 2, 1763, p. 248, nec Vahl, iSkrift. Kjocb. 6, Prod. G. Gen. 1810, p. 104; DC., Syst. 1, 1818, p. 433; DC., 1, 1824, p. 75; Don, Hist. Dich'l. PI. Ilook.f. & Fl. Ind. Fl. Ind. 1, 1'831, p. 76; Thorns., 1, 1855, p. 66; Miq., Bat. 1, 10; Fl. Ind. Bat. 1, Benth. & 2, 1859, p. Miq., Suppl. 1860, p. 152, 618, 619; Gen. PI. AdansOnia Ann. Mus. Hook.f., 1, 1S62, p. 13; Baill., 6, 1865—6, p. 281; Miq., Bot. Bat. 4, Hist. d. PL 1867 Ilook.f. & Lugd. 1868, p.