THE UNIVERSITY OF KANSAS PALEONTOLOGICAL CONTRIBUTIONS ARTICLE 60 (ECHINODERMATA 12) LOWER PENNSYLVANIAN (MORROWAN) CRINOIDS FROM ARKANSAS, OKLAHOMA, AND TEXAS RAYMOND C. MOORE and HARRELL L. STRIMPLE The University of Kansas, Lawrence; The University of Iowa, Iowa City The University of Kansas Paleontological Institute THE UNIVERSITY OF KANSAS PUBLICATIONS JUNE 22, 1973 UKPAII, 1-54 ( 173) THE UNIVERSITY OF KANSAS PALEONTOLOGICAL CONTRIBUTIONS Article 60 (Echinodermata 12), 84 Pages, 7 Figures, 23 Plates, 1 Table LOWER PENNSYLVANIAN (MORROWAN) CRINOIDS FROM ARKANSAS, OKLAHOMA, AND TEXAS RAYMOND C. MOORE and HARRELL L. STRIMPLE The University of Kansas, Lawrence; The University of Iowa, Iowa City CONTENTS PAGE PAGE ABSTRACT 7 Genus Puramphicrinus Strimple & INTRODUCTION 7 Moore, 1971 37 Previous studies of Morrowan crinoids 8 P. magnus Moore & Strimple, Stratigraphie setting 9 new species 38 Summary of Morrowan stratigraphie units Subclass Camerata Wachsmuth & Springer, 1885 39 in Arkansas and Oklahoma 11 Order Monobathrida Moore & Laudon, 1943 39 Age relationships of Morrowan crinoids 12 Suborder Tanaocrinina Moore, 1952 39 Some morphological terms 12 Superfamily Periechocrinitacea Austin & Modifications in form of inadunate cups 13 Austin, 1843 (Ubaghs, 1953) 39 Phylogeny 13 Family Paragaricocrinidae Moore & Discrimination of superfarnilies 14 Laudon, 1942 40 Classi fication of flexible and inadunate crinoids 15 Genus Megaliocrinus Moore & Acknowledgments 31 Laudon, 1942 40 SYSTEMATIC DESCRIPTIONS 31 M. aplatus Moore & Laudon, 1942 40 Designations of specimens 31 Suborder Glvptocrinina Moore, 1952 40 Class Crinoidea Miller, 1821 31 Superfamily Platycrinitacea Austin & Subclass Flexibilia Zittel, 1895 32 Austin, 1843 (Ubaghs, 1953) 41 Order Sagenocrinida Springer, 1913 32 Family Platycrinitidae Bassler, 1938 41 Superfamily Lecanocrinacea Springer, 1913 Genus Platycrinites Miller, 1821 41 (Moore & Strimple, new superfamily) 32 P. species 41 Family Mespilocrinidae Jaekel, 1918 32 Subclass Inadunata Wachsmuth & Springer, 1885 .. 41 Genus Cibolocrinus Weller, 1909 32 Order Cladida Moore & Laudon, 1943 41 C. tumidus Moore & Plummer, 1938 32 C. spinosus Moore & Strimple, Suborder Cyathocrinina Bather, 1899 42 new species 33 Superfamily Codiacrinacea Bather, 1890 C. circulus Moore & Strimple, (Lane, 1967) 42 new species 33 Family Codiacrinidae Bather, 1890 42 SubfamilyC. bellus Codiacrininae Moore & Strimple, Bather, 1890 new species 33 Genus Lecythiocrinus White, 1879 42 Family Calycocrinidae Moore & Strimple, L. asymmetricus Moore & new family 34 Strimple, new species 42 Genus Calycocrinus Wanner, 1916 34 Suborder Poteriocrinina laekel, 1918 42 C. symmetricus Moore & Strimple, Superfamily Scytalocrinacea Moore & Laudon, new species 34 1943 (Moore & Strimple, new superfamily) .... 42 C. furnish: Moore & Strimple, Family Scytalocrinidae Moore & new species 35 Laudon, 1943 43 Superfamily Sagenocrinitacea Bassler, 1938 Genus Scytalocrinus Wachsmuth & (Moore & Strimple, new superfamily) 35 Springer, 1879 43 Family Dactylocrinidae Bather, 1899 36 S. crassibrachiatus Moore & Strimple, Genus Zenocrinus Moore & Strimple, new species 43 new genus 36 S. sansabensis Moore & Plummer, 1940 44 Z. Zeus Moore & Strimple, new species .. 36 Genus Phacelocrinus Kirk, 1940 44 Family Euryocrinidae Moore & Strimple, P. rose: (Moore & Plummer, 1938) new family 37 Moore & Stritnple, new combination .. 44 PAGE PAGE P. brevis Moore & Strimple, Genus Metacromyocrinus Strimple, 1961 55 new species 44 M. papulosus (Moore & Plummer, Genus Morrotvcrinus Moore & 1938) Moore & Strimple, Plummer, 1938 45 new combination 55 Family Spaniocrinidae Moore & M. gillumi Strimple, 1966 55 Laudon, 1943 45 Genus Goleocrinus Strimple & Genus Spaniocrinus Wanner, 1924 45 Watkins, 1969 56 S. species 45 G. vadosus (Washburn), 1968, Superfamily Mollocrinacea \Vanner, 1916 Moore & Strimple, new combination .. 56 (Moore & Strimple, new superfamily) 45 Genus Dicromyocrinus Jaekel, 1918 56 Family Mollocrinidae Wanner, 1916 45 D optimus Strimple, 1951 57 Genus Strongylocrinus Wanner, 1916 45 D. medius Moore & Strimple, S. hansoni Moore & Strimple, new species 57 new species 46 D. subaplatus Moore & Strimple, S. ornatus Moore & Strimple, new species 59 new species 46 Genus Paracromyocrinus Strimple, 1966 59 Superfamily Lophocrinacea Bather, 1899 P. oklahomensis (Moore & (Moore & Scrimple, new superfamily) 46 Plummer), 1938 59 Family Laudonocrinidae Moore & Strimple, P.? species 61 new family 47 Superfamily Erisocrinacea Wachsmuth & Genus Anchicrinus Strimple & Springer, 1886 (S. A. Miller, 1890) (Moore & Watkins, 1969 47 Strimple, new superfamily) 61 A. plan ulatus Moore & Strimple, Family Erisocrinidae Wachsmuth & Springer, new species 47 1886 (S. A. Miller, 1890) 61 A. rugosus (Strimple, 1961) Moore & Genus Erisocrinus Meek & Strimple, new combination 48 Worthen, 1865 61 E.Genus Athlocrinus Moore & Plummer, tvapanucka (Strimple), 1961 61 Family1940 48 Paradelocrinidae Knapp, 1969 62 A. dejustus Moore & Strimple, Genus Atokacrinus Knapp, 1969 62 new species 49 A. tumulosus Moore & Strimple, Family Stellarocrinidae Strimple, 1961 49 new species 62 Heliosocrinus Strimple, 1961 49 Genus Family Arkacrinidae Knapp, 1969 62 Moore & Strimple, H. neotosus Genus Arkacrinus Knapp, 1969 63 50 new species A. dubius (Mather), 1915 63 50 H. species A. constrictus Moore & Strimple, Kirk, 1942 50 Family Pachylocrinidae new species 63 Genus Plum mericrinus Moore & Laudon, 1943 50 Family Diphuicrinidae Strimple & P. expansus Moore & Strimple, Knapp, 1966 64 new species 50 Genus Diphuicrinus Moore & 64 Superfamily Agassizocrinacea S. A. Miller, Plummer, 1938 Moore & 1938 64 1890 (Moore & Strimple, new superfamily) .... 51 D. croneisi Plummer, D. faustus Moore & Strimple, Family Agassizocrinidae S. A. Miller, 1890 .... 51 Genus Paragassizocrinus Moore & new species 65 mammi fer Plummer, 1940 51 D. Moore & Strimple, P. kendrickensis Strimple & new species 65 Knapp, 1966 52 D. pentanodus Moore & Strimple, 65 Family Ampelocrinidae Kirk, 1942 52 new species Genus Ampelocrinus Kirk, 1942 53 Family Catacrinidae Knapp, 1969 65 A. convexus (Strimple, 1940) Moore & Genus Palmerocrinus Knapp, 1969 66 Strimple, new combination 53 P. prof undus Moore & Strimple, Family Anobasicrinidae Strimple, 1961 53 new species 66 Genus Anobasicrinus Strimple, 1961 53 P. kesslerensis Moore & Strimple, A. obscurus Strimple, 1961 53 new species 67 Superfamily Cromyocrinacea Bather, 1890 Genus Endelocrinus Moore & (Moore & Strimple, new superfamily) 54 Plummer, 1940 67 Family Cromyocrinidae Bather, 1890 E. matheri (Moore & Plummer), 1938 67 (Jaekel, 1918) 54 Superfamily Pirasocrinacea Moore & Laudon, Genus Cromyocrinus Trautschold, 1867 54 1943 (Moore & Strimple, new superfamily) .... 68 C. grandis Mather, 1915 54 Family Pirasocrinidae Moore & Laudon, 1943 .. 68 PAGE PAGE Genus Lasanocrinus Moore & A. politus Moore & Strimple, Plummer, 1940 68 hansoni 46 L. daileyi (Strimple), 1940 68 A. normalis Moore & Strimple, L. nodatus Moore & Strimple, new species 74 new species 69 Genus Stenopecrinus Strimple, 1961 75 L. nod us Moore & Strimple, S. ornatus Moore & Strimple, new species 69 new species 75 L. multinodulus Moore & Strimple, S. species 75 new species 69 Genus Sciadiocrinus Moore & L. strigosus (Moore & Plummer), Plummer, 1938 75 1938 69 S. cascus Moore & Strimple, L. minutus Moore & Strimple, new species 75 new species 70 S. crassacanthus? Moore & Genus Utharocrinus Moore & Plummer, 1938 76 Plummer, 1938 71 Genus Perimestocrinus Moore & U. pentanodus (Mather), 1915 71 Plummer, 1938 76 Genus Metutharocrinus Moore & P. teneris Moore & Plummer, 1938 76 Strimple, new genus 71 Superfamily Texacrinacea Moore & Strimplc, newM. cockei Moore & Strimple, superfamily 76 new species 72 Family Cymbiocrinidae Strimple & Watkins,M. spinifer Moore & Strimple, 1969 77 new species 72 Genus Proallosocrinus Moore & M. undulatus Moore & Strimple, Strimple, new genus 77 new species P. glenisteri Moore & Strimple, Genus Affinocrinus Knapp, 1969 73 new species 77 A. grandis Moore & Strimple, REFERENCES 78 new species 73 A. progressus Moore & Strimple, LOCALITY REGISTER 78 new species 73 INDEX 80 ILLUSTRATIONS FIGURE PAGE PLATE FACING PAGE 1. Correlation of Lower Pennsylvanian (Morrow- bensis; 2a-d, Cromyocrinus grandis; 3. Paragassi- an) deposits in Arkansas and Oklahoma 10 zocrinus kendrickensis; 4a-d, Strongylocrinus 2. Interpretation of orientation of cup and stem of hansoni 46 Calycocrinus (luring life 26 6. Inadunate crinoids: 1-4, Anchicrinus planulatus; 3, 4. Paramphicrinus oklahomaensis 38 5a-c, Athlocrinus dejustus; 6a-c, Affinocrinus 5. Cups and arms of Goleocrinus, Steno pecrinus, normalis 47 and Palmerocrinus 58 7. Inadunate crinoids: 1-5, Anchicrinus rugosus 48 6. Cup of Lasanocrinus strigosus 70 8. Inadunate crinoids: la,b, Atfinocrinus progres- 7. Cup of Affinocrinus politus 74 sus, and 2a-c, A. normalis; 3a,b, Heliosocrinus sp., and 4a-c, H. neotosus; 5a-c, Dicromyocrinus PLATE FACING PAGE medius 49 All Morrowan crinoids. 9. Inadunate crinoids: la-d, Cromyocrinus grandis; 1. Flexible crinoid Calycocrinus: la-d, C. tumidus; 2a-c, Proallosocrinus glenisteri; 3a-d, Plummeri- 2a-e, C. spinosus; 3a-d, 4a-d, C. be//us 34 crinus expansus; 4a,b, Lasanocrinus nodatus 52 2. Flexible crinoids: la-d, Cibolocrinus circulus, 10. Inadunate crinoids: la-d, Metacromyocrinus pap- and 4, C. be//us; 2a-d, Calycocrinus furnishi, and ulosus; 2-4, Paracromyocrinus oklahomensis 53 3a-d, C. symmetricus 35 11. Inadunate crinoids: 1,4, Dicromyocrinus sub-