SHILAP Revista de Lepidopterología ISSN: 0300-5267 [email protected] Sociedad Hispano-Luso-Americana de Lepidopterología España Razowski, J. The Old World Hilarographini (Lepidoptera: Tortricidae) SHILAP Revista de Lepidopterología, vol. 37, núm. 147, septiembre, 2009, pp. 261-287 Sociedad Hispano-Luso-Americana de Lepidopterología Madrid, España Available in: http://www.redalyc.org/articulo.oa?id=45515238001 How to cite Complete issue Scientific Information System More information about this article Network of Scientific Journals from Latin America, the Caribbean, Spain and Portugal Journal's homepage in redalyc.org Non-profit academic project, developed under the open access initiative 261-287 The Old World Hilaro 7/9/09 15:00 Página 261 SHILAP Revta. lepid., 37 (147), septiembre 2009: 261-287 CODEN: SRLPEF ISSN:0300-5267 The Old World Hilarographini (Lepidoptera: Tortricidae) J. Razowski Abstract A previous synonimization of Thaumatographa Walsingham, 1897 with Hilarographa is confirmed. The systematic list of the Old World species of Hilarographini is provided; Embolostoma Diakonoff, 1977 and Nexosa Diakonoff, 1977 are excluded from this tribe. Thirty three species of Hilarographa Zeller, 1877 from the Oriental and Australian regions are examined of which twenty-nine are described as new: H. pahangana Razowski, sp. n., H. celebesiana Razowski, sp. n., H. baliana Razowski sp. n., H. perakana Razowski, sp. n., H. ancilla Razowski sp. n., H. meekana Razowski, sp. n., H. soleana Razowski, sp. n., H. gentinga Razowski, sp. n., H. johibradleyi Razowski, sp. n., H. fergussonana Razowski, sp. n., H. auroscripta Razowski, sp. n., H. hainanica Razowski, sp. n., H. marangana Razowski, sp. n., H. buruana Razowski, sp. n., H. bosavina Razowski, sp. n., H. sipiroca Razowski, sp. n., H. rampayoha Razowski sp. n., H. uthaithani Razowski, sp. n., H. tasekia Razowski, sp. n., H. emburonga Razowski, sp. n., H. muluana Razowski, sp. n., H. shehkonga Razowski, sp. n., H. gunongana Razowski, sp. n., H. renonga Razowski, sp. n., H. khaoyai Razowski sp. n., H. uluana Razowski, sp. n., H. obinana Razowski, sp. n., H. robinsoni Razowski, sp. n., H. calyx Razowski, sp. n. Nine new combinations are proposed (see the list of taxa). KEY WORD: Lepidoptera, Tortricidae, Hilarographini, Oriental and Australian region. Los Hilarographini del Viejo Mundo (Lepidoptera: Tortricidae) Resumen Se confirma la sinonimia previa de Thaumatographa Walsingham, 1897 con Hilarographa. Se proporciona la lista sistemática de los Hilarographini del Viejo Mundo; se excluyen de esta tribu a Embolostoma Diakonoff, 1977 y Nexosa Diakonoff, 1977. Se examinan treinta y tres especies de Hilarographa Zeller, 1877 de la región Oriental y Australiana de las que veintinueve se describen como nuevas: H. pahangana Razowski, sp. n., H. celebesiana Razowski, sp. n., H. baliana Razowski sp. n., H. perakana Razowski, sp. n., H. ancilla Razowski sp. n., H. meekana Razowski, sp. n., H. soleana Razowski, sp. n., H. gentinga Razowski, sp. n., H. johibradleyi Razowski, sp. n., H. fergussonana Razowsii, sp. n., H. auroscripta Razowski, sp. n., H. hainanica Razowski, sp. n., H. marangana Razowski, sp. n., H. buruana Razowski, sp. n., H. bosavina Razowski, sp. n., H. sipiroca Razowski, sp. n., H. rampayoha Razowski sp. n., H. uthaithani Razowski, sp. n., H. tasekia Razowski, sp. n., H. emburonga Razowski, sp. n., H. muluana Razowski, sp. n., H. shehkonga Razowski, sp. n., H. gunongana Razowski, sp. n., H. renonga Razowski, sp. n., H. khaoyai Razowski sp. n., H. uluana Razowski, sp. n., H. obinana Razowski, sp. n., H. robinsoni Razowski, sp. n., H. calyx Razowski, sp. n. Se proponen nueva nuevas combinaciones (véase la lista de taxa). PALABRAS CLAVE: Lepidoptera, Tortricidae, Hilarographini, región Oriental y Australiana. Introduction The most complete studies on the Hilarographini are by DIAKONOFF (1977b dealing with the Oriental and Australian regions; 1986 - Palaearctic region) who excluded them (DIAKONOFF 1977b) 261 261-287 The Old World Hilaro 7/9/09 15:00 Página 262 J. RAZOWSKI from Glyphipteridae. Then HEPPNER (1982) published a synopsis of the Hilarographini of the World. Both authors included in Hilarorgaphini the representatives of three different subfamilies, Chlidanotinae, Tortricinae and Olethreutinae. To this last belong a few genera provisionally placed (RAZOWSKI 2009) in Archipini. To the genuine Hilarographini belonged only Charitographa Diakonoff, 1979, Hilarographa Zeller, 1877, and Thaumatographa Walsingham, 1897. All these genera exhibit only slight differences between them. According to the original description Charitographa differs from Thaumatographa to which it is similar externally in a broad ductus bursae, lack of signa and the dense asteroid sculpture of the corpus bursae. Only this last character has not been found in other Hilarographini. Its facies, the venation and the male genitalia do not differ from those in the mentioned genus. The remaining characters mentioned by DIAKONOFF (1977b) in the descriptions of genera refer to the tribe or even to the subfamily Chlidanotinae. Meyrick sunk Thaumatographa and African Idiothauma Walsingham, 1897 as synonymies of Hilarographa. As concerns the former I am following Meyrick’s point of view. The monobasic Embolostoma Diakonoff, 1977 described to comprise E. plutostola Diakonoff, 1977 from Java, known from a single female, does not belong in this tribe, as shown by the shape of the labial palpus illustrated by its author. Another genus usually included (HEPPNER 1982, CLARKE 1969) in Hilarographini, Irianassa Meyrick, 1905, is transferable to Enarmoniini (HORAK 2006). Apart from its type species (I. sappiropa Meyrick, 1905 from Sri Lanka) and the Australian I. aetheria (Turner, 1946) three species are included in this genus by BROWN (2005). I. poecilaspis Meyrick, 1923 certainly belongs to Hilarographini but requires re-examination. Two species, Grapholitha speciosana Pagenstecher, 1900, Bismarck Archipelago (= Irianassa alcyonopa Meyrick, 1926, New Ireland) and I. uranopa Meyrick, 1927, Samoa are unknown to me. Nexosa Diakonoff, 1977 known from the Oriental and Australian regions with its four species belongs to the Archipini. Material The paper is based on the collection of the Natural History Museum, London. Systematics Preliminary list of the Old World Hilarographini Hilarographa Zeller, 1877 zapyra Meyrick, 1886, New Guinea pahangana Razowski, sp. n., Malaysia celebesiana Razowski, sp. n., Celebes oenobapta (Diakonoff, 1977), comb. n., Java crocochorista (Diakonoff, 1983), comb. n., Java phlox (Diakonoff, 1977), comb. n., Java cladara (Diakonoff, 1977), comb. n. Borneo ceramopa Meyrick, 1920, Assam, India baliana Razowski, sp. n., Bali Island, Malaysia perakana Razowski, sp. n., Perak, Malaysia caminodes Meyrick, 1905, Sri Lanka calathisca Meyrick, 1909, Assam, India mechanica Meyrick, 1909, Assam, India hermatodes Meyrick, 1909, Sri Lanka ancilla Razowski, sp. n., Bombay, India 262 SHILAP Revta. lepid., 37 (147), septiembre 2009 261-287 The Old World Hilaro 6/12/09 10:24 Página 263 THE OLD WORLD HILAROGRAPHINI meekana Razowski, sp. n., Fergusson Islands machaerophora Diakonoff & Arita, 1976, Honsyu, Japan dolichosticha (Diakonoff, 1977), comb. n., Java tornoxena (Diakonoff, 1977), comb. n., Java soleana Razowski, sp. n., Seram, Indonesia merinthias Meyrick, 1909, Assam, India gentinga Razowski, sp. n., Malaysia tornoxena (Diakonoff, 1977), comb. n., Java, Indonesia johnibradleyi Razowski, sp. n., Thailand ferox Meyrick, 1921, Java, Indonesia fergussonana Razowski, sp. n., Fergusson Islands undosa (Diakonoff, 1977), comb. n., New Guinea auroscripta Razowski, sp. n., Ambon Island, Indonesia excellens (Pagenstecher, 1900), Bismark Island, Papua New Guinea hainanica Razowski, sp. n., Hainan Island, China marangana Razowski, sp. n., Sumatra, Indonesia buruana Razowski, sp. n., Buru Island, Indonesia spermatodesma Diakonoff, 1955, New Guinea macaria (Diakonoff, 1977), comb. n., Java bosavina Razowski, sp. n., Papua New Guinea sipiroca Razowski, sp. n., Sumatra, Indonesia rampayoha Razowski, sp. n., Brunei eremnotorna Diakonoff & Arita, 1976, Honsyu, Japan aurosa Diakonoff & Arita, 1976, Yakushima Island, Japan uthaithani Razowski, sp. n., Thailand decoris Diakonoff & Arita, 1976, Kuril Islands, Russia tasekia Razowski, sp. n., Perak, Malaysia druidica (Meyrick, 1909), Assam, India. ludens Diakonoff, 1948, Buru, Indonesia temburonga Razowski, sp. n., Brunei muluana Razowski, sp. n., Sarawak leucopyrga Meyrick, 1912, Kyushyu Island, Japan shehkonga Razowski, sp. n., Hong Kong cymatodes (Diakonoff, 1983), comb. n., Sumba Island, Indonesia opistocapna (Diakonoff, 1977), New Guinea gunongana Razowski, sp. n., Sarawak renonga Razowski, sp. n., Thailand khaoyai Razowski, sp. n., Thailand obinana Razowski, sp. n., Obi Major Island, Moluccas uluana Razowski, sp. n., Obi Major Island, Moluccas robinsoni Razowski, sp. n., Brunei citharistis Meyrick, 1909, Assam, India mesostigmatias (Diakonoff, 1977), comb. n., Taiwan calyx Razowski, sp. n., Taiwan tetralina Meyrick, 1930, Solomon Islands Unplaced species: cirrhocosma Meyrick, 1930, Solomon Islands pampoecila (Thurner, 1913), Queensland, Australia SHILAP Revta. lepid., 37 (147), septiembre 2009 263 261-287 The Old World Hilaro 7/9/09 15:00 Página 264 J. RAZOWSKI Charitographa Diakonoff, 1979 micadonis (Stringer, 1930), Japan Idiothauma Walsingham, 1897 africanum Walsingham, 1897, Republic of Congo malagassicum Viette, 1958, Madagascar rigatiellum (Ghesquiere, 1940), Zaire Descriptions