ISSN 0006 8241 = Bothalia Bothalia A JOURNAL OF BOTANICAL RESEARCH Vol. 39,2 Oct. 2009 TECHNICAL PUBLICATIONS OF THE SOUTH AFRICAN NATIONAL BIODIVERSITY INSTITUTE PRETORIA Obtainable from the South African National Biodiversity Institute (SANBI), Private Bag X101, Pretoria 0001, Republic of South Africa. A catalogue of all available publications will be issued on request. BOTHALIA Bothalia is named in honour of General Louis Botha, first Premier and Minister of Agriculture of the Union of South Africa. This house journal of the South African National Biodiversity Institute, Pretoria, is devoted to the furtherance of botanical science. The main fields covered are taxonomy, ecology, anatomy and cytology. Two parts of the journal and an index to contents, authors and subjects are published annually. Three booklets of the contents (a) to Vols 1–20, (b) to Vols 21–25, (c) to Vols 26–30, and (d) to Vols 31–37 (2001– 2007) are available. STRELITZIA A series of occasional publications on southern African flora and vegetation, replacing Memoirs of the Botanical Survey of South Africa and Annals of Kirstenbosch Botanic Gardens. MEMOIRS OF THE BOTANICAL SURVEY OF SOUTH AFRICA The memoirs are individual treatises usually of an ecological nature, but sometimes dealing with taxonomy or economic botany. Published: Nos 1–63 (many out of print). Discontinued after No. 63. ANNALS OF KIRSTENBOSCH BOTANIC GARDENS A series devoted to the publication of monographs and major works on southern African flora.Published: Vols 14–19 (earlier volumes published as supplementary volumes to the Journal of South African Botany). Discontinued after Vol. 19. FLOWERING PLANTS OF AFRICA (FPA) This serial presents colour plates of African plants with accompanying text. The plates are prepared mainly by the artists at the South African National Biodiverity Institute. Many botanical artists have contributed to the series, such as Fay Anderson, Peter Bally, Auriol Batten, Gillian Condy, Betty Connell, Stella Gower, Rosemary Holcroft, Kathleen Lansdell, Cythna Letty (over 700 plates), Claire Linder-Smith and Ellaphie Ward-Hilhorst. The Editor is pleased to receive living plants of general interest or of economic value for illustration. From Vol. 55, twenty plates are published at irregular intervals. An index to Vols 1–49 is available. FLORA OF SOUTHERN AFRICA (FSA) A taxonomic treatise on the flora of the Republic of South Africa, Lesotho, Swaziland, Namibia and Botswana. the FSA contains descriptions of families, genera, species, infraspecific taxa, keys to genera and species, syn- onymy, literature and limited specimen citations, as well as taxonomic and ecological notes. Contributions to the FSA also appear in Bothalia. PALAEOFLORA OF SOUTHERN AFRICA A palaeoflora on a pattern comparable to that of the Flora of southern Africa. Much of the information is pre- sented in the form of tables and photographic plates depicting fossil populations. Now available: Molteno Formation (Triassic) Vol. 1. Introduction. Dicroidium, 1983, by J.M. & H.M. Anderson. Molteno Formation (Triassic) Vol. 2. Gymnosperms (excluding Dicroidium), 1983, by J.M. & H.M. Anderson. Prodromus of South African Megafloras. Devonian to Lower Cretaceous, 1985, by J.M. & H.M. Anderson. Obtainable from: A.A. Balkema Marketing, Box 317, Claremont 7735, RSA. Towards Gondwana Alive. Promoting biodiversity and stemming the Sixth Extinction, 1999, by J.M. Anderson (ed.). Heyday of the gymnosperms: systematics and biodiversity of the Late Triassic Molteno fructifications, 2003, by J.M. Anderson & H.M. Anderson. Strelitzia 15. Brief history of the gymnosperms: classification, biodiversity, phytogeography and ecology, 2007, by J.M. Anderson, H.M. Anderson & C.J. Cleal. Strelitzia 20. Molteno ferns: Late Triassic biodiversity in southern Africa, 2008, by H.M. Anderson & J.M. Anderson. Strelitzia 21. SANBI BIODIVERSITY SERIES A series of occasional reports on projects, technologies, workshops, symposia and other activities initated by or executed in partnership with SANBI. BOTHALIA A JOURNAL OF BOTANICAL RESEARCH Volume 40,1 Scientific Editor: G. Germishuizen Technical Editor: B.A. Momberg 2 Cussonia Avenue, Brummeria, Pretoria Private Bag X101, Pretoria 0001 ISSN 0006 8241 May 2010 Editorial Board D.F. Cutler Royal Botanic Gardens, Kew, UK B.J. Huntley South African National Biodiversity Institute, Cape Town, RSA P.H. Raven Missouri Botanical Garden, St Louis, USA M.J.A. Werger University of Utrecht, Utrecht, The Netherlands Acknowledgements to referees Allemann, Dr J. University of the Free State, Bloemfontein, RSA. Archer, Mrs C. South African National Biodiversity Institute, Pretoria, RSA. Archer, Dr R. South African National Biodiversity Institute, Pretoria, RSA. Briggs, Dr B. Royal Botanic Gardens, Sydney, Australia. Brummitt, Dr R.K. Royal Botanic Gardens, Kew, England, UK. Burrows, J.E. Box 710, 1120 Lydenburg, RSA. Claßen-Bockhoff, Prof. R. J. Gutenberg-Universität Mainz, Mainz, Germany. Craib, C. Box 67142, Bryanston, 2021 Sandton, RSA. Crouch, Prof. N.R. South African National Biodiversity Institute, Durban, RSA. Ellery, Prof. W.N. Rhodes University, Grahamstown, RSA. Feuillet, Dr C. Smithsonian Institution, Washington DC, USA. Forster, Dr P.I. Queensland Herbarium, Brisbane Botanic Gardens, Australia. Glen, Dr H.F. South African National Biodiversity Institute, Durban, RSA. Goldblatt, Dr P. Missouri Botanical Garden, St Louis, USA. Grobler, Mrs A. South African National Biodiversity Institute, Pretoria, RSA. Hill, Dr T.R. University of KwaZulu-Natal, Pietermaritzburg, RSA. Hind, Dr D.J.N. Royal Botanic Gardens, Kew, England, UK. Klopper, Mrs R.R. South African National Biodiversity Institute, Pretoria, RSA. Lammers, Prof. T.G. University of Wisconsin, Oshkosh, USA. Lavranos, J. Apartado Postal 243, 8100 Loule, Portugal. Leistner, Dr O.A. 194 Griselda Rd, Murrayfield, 0184 Pretoria, RSA. Manning, Dr J.C. South African National Biodiversity Institute, Cape Town, RSA. Mfundisi, Dr K. Harry Oppenheimer Okavango Research Centre, Maun, Botswana. Retief, Dr E. South African National Biodiversity Institute, Pretoria, RSA. Roux, Dr J.P. South African National Biodiversity Institute, Cape Town, RSA. Schrire, Dr B. Royal Botanic Gardens, Kew, England, UK. Smithies, Mrs S.J. South African National Biodiversity Institute, Pretoria, RSA. Snijman, Dr D. South African National Biodiversity Institute, Cape Town, RSA. Steiner, Dr K.E. California Academy of Sciences, San Francisco, USA. Van Jaarsveld, E.J. South African National Biodiversity Institute, Cape Town, RSA. Van Wyk, Prof. B-E. University of Johannesburg, RSA. Vlok, J. Regalis Environmental Services, P.O. Box 1512, 6620 Oudtshoorn, RSA. Welman, Ms W.G. South African National Biodiversity Institute, Pretoria, RSA. Werger, Prof. M.J.A. Department Plant Ecology and Evolutionary Biology, Utrecht, The Netherlands. Yatskievych, Dr G. Missouri Botanical Garden, St Louis, USA. Yoshida, Dr N. Miyazaki University, Miyazaki, Japan. Date of publication of Bothalia 39,2: 6 October 2009. CONTENTS Bothalia 39,2 1. New species of Geissorhiza (Iridaceae: Crocoideae) from the southern African winter rainfall zone, range extensions, taxonomic changes, and notes on pollen morphology and floral ecology. P. GOLDBLATT, J.C. MANNING and I. NÄNNI ........................................ 123 2. Albuca crispa and A. grandis (Hyacinthaceae: Ornithogaloideae), two new species of subgenus Albuca, the rediscovery of A. albucoides (subgenus Osmyne), and the identity of A. reflexa. J.C. MANNING and P. GOLDBLATT. 153 3. Taxonomy of the genus Keetia (Rubiaceae–subfam. Ixoroideae–tribe Vanguerieae) in southern Africa, with notes on bacterial symbiosis as well as the structure of colleters and the ‘stylar head’ complex. P.M. TILNEY and A.E. VAN WYK ...................................................... 165 4. Allioni’s Aloe names (Asphodelaceae): nomenclature and typification. L. GUGLIELMONE, L. GALLO, M. MEREGALLI, G.F. SMITH and E. FIGUEIREDO ................................... 177 5. The botany of the Cunene–Zambezi Expedition with notes on Hugo Baum (1867–1950). E. FIGUEIREDO, M. SOARES, G. SEIBERT, G.F. SMITH and R.B. FADEN ............................... 185 6. New records of pteridophytes from Annobón Island. E. FIGUEIREDO, A. GASCOIGNE & J.P. ROUX .... 213 7. Notes on African plants: Euphorbiaceae. A new species of succulent Euphorbia from southern Angola. P.V. BRUYNS .... 219 Hyacinthaceae. Nomenclatural corrections in Ornithogaloideae. J.C. MANNING . 221 Hyacinthaceae. Drimia oliverorum (Urgineoideae), a new species from southern Namibia. J.C. MANNING and E.G.H. OLIVER ............................................... 225 Hyacinthaceae. Pseudogaltonia liliiflora (Ornithogaloideae), a new species from the Richtersveld, Northern Cape. J.C. MANNING, P. GOLDBLATT and G. WILLIAMSON .............. 229 Hyacinthaceae. A new subspecies of Pseudoprospero firmifolium (Hyacinthoideae) from KwaZulu- Natal. J.C. MANNING, D.G.A. STYLES and N.R. CROUCH . 231 Hyacinthaceae. A new species of Drimia (Urgineoideae) from the Knersvlakte, Western Cape, South Africa. D.A. SNIJMAN and A.D. HARROWER .............................. 233 Mimosaceae. A new species of Acacia from northern Mozambique. J.E. BURROWS and S.M. BURROWS . 222 Poaceae. A new subspecies of Trichoneura eleusinoides (Eragrostideae) and two new species of Panicum (Panicoideae–Paniceae) from South Africa. L. FISH ........................ 237 Pteridophyta. Otto Kuntze’s lycopod and fern collections from South Africa. J.P. ROUX . 227 Ranunculaceae.