W&M ScholarWorks Dissertations, Theses, and Masters Projects Theses, Dissertations, & Master Projects 1984 On the anatomy and phylogeny of the Zoarcidae (teleostei: perciformes) M. Eric. Anderson College of William and Mary - Virginia Institute of Marine Science Follow this and additional works at: https://scholarworks.wm.edu/etd Part of the Zoology Commons Recommended Citation Anderson, M. Eric., "On the anatomy and phylogeny of the Zoarcidae (teleostei: perciformes)" (1984). Dissertations, Theses, and Masters Projects. Paper 1539616550. https://dx.doi.org/doi:10.25773/v5-qgxc-9x74 This Dissertation is brought to you for free and open access by the Theses, Dissertations, & Master Projects at W&M ScholarWorks. It has been accepted for inclusion in Dissertations, Theses, and Masters Projects by an authorized administrator of W&M ScholarWorks. For more information, please contact [email protected]. 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University M'odrilms International 300 N. Zeeb Road Ann Arbor, Ml 48106 Reproduced with permission of the copyright owner. Further reproduction prohibited without permission. Reproduced with with permission permission of the of copyright the copyright owner. owner.Further reproductionFurther reproduction prohibited without prohibited permission. without permission. 8428464 Anderson, Michael Eric ON THE ANATOMY AND PHYLOGENY OF THE ZOARCIDAE (TELEOSTEI: PERCIFORMES) The College of William and Mary in Virginia Ph.D. 1984 University Microfilms International 300 N. Zeeb Road, Ann Arbor, Ml 48106 Reproduced with permission of the copyright owner. Further reproduction prohibited without permission. Reproduced with with permission permission of the of copyright the copyright owner. owner.Further reproductionFurther reproduction prohibited without prohibited permission. without permission. 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Other______________________________________________________ _ _ _ _ _ University Microfilms International Reproduced with permission of the copyright owner. Further reproduction prohibited without permission. Reproduced with with permission permission of the of copyright the copyright owner. owner.Further reproductionFurther reproduction prohibited without prohibited permission. without permission. ON THE ANATOMY AND PHYLOGENY OF THE ZOARCIDAE (TELEOSTEI: PERCIFORMES) A Dissertation Presented to The Faculty of the School of Marine Science The College of William and Mary in Virginia In Partial Fulfillment Of the Requirements for the Degree of Doctor of Philosophy by Michael Eric Anderson r 1984 Reproduced with permission of the copyright owner. Further reproduction prohibited without permission. APPROVAL SHEET This dissertation is submitted in partial fulfillment of the requirements for the degree of Doctor of Philosophy £/lCc Michael Eric Anderson Approved, August 1984 :k, Chairman G X . , J yAQmM Daniel M. Cohen, Natural History Museum of Los Angeles County Robert H. Gibbs, Jr^/, National Museum of Natural History John ierriner, Natioral Marine leries Service Evon P. Ruzecki Reproduced with permission of the copyright owner. Further reproduction prohibited without permission. TABLE OF CONTENTS Page ACKNOWLEDGMENTS................................................. vi LIST OF TABLES................................................. viii LIST OF FIGURES................................................. ix LIST OF PLATES.................................................. xvi ABSTRACT...................................................... xviii INTRODUCTION.................................................... 2 METHODS AND MATERIALS........................................... 5 RESULTS......................................................... 9 Relationships among Zoarceoidea............................. 9 Classification of Zoarcidae................................. 20 Character Matrix............................................ 23 Phylogeny of Zoarcidae...................................... 42 Zoarcidae (Taxonomy, Description)........................... 50 Artificial Key to Genera of Adult and Late Juvenile Zoarcidae.. 54 Generic Accounts............................................ 59 Subfamily Lycozoarcinae............ 59 Lycozoarces Popov, 1933......................... 59 Subfamily Zoarcinae..................................... 61 Tribe Zoarcini...................................... 62 Macrozoarces Gill, 1864.......................... 62 Zoarces Cuvier, 1829............................ 64 Tribe Gymnelini..................................... 67 iii Reproduced with permission of the copyright owner. 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Page Andriashevia Fedorov and Neyelov, 1978........... 67 Bllabria Shmidt, 1936........................... 69 Davidi.j ordania Popov, 1931....................... 71 Gymnelopsis Soldatov, 1922....................... 74 Gymnelus Reinhardt, 1834......................... 76 Hadropareia Shmidt, 1904........................ 80 Krusensterniella Shmidt, 1904.................... 82 Melanostigma Gunther, 1881....................... 84 Nalbantichthys Schultz, 1967..................... 88 Puzanovia Fedorov, 1975......................... 91 Tribe Lycodini...................................... 94 Aiakas Gosztonyi, 1977.......................... 94 Austrolycichthys Regan, 1913..................... 96 Austrolycus Regan, 1913......................... 98 Bothrocara Bean, 1891........................... 101 Bothrocarina Suvorov, 1935....................... 105 Crossostomus Lahille, 1908....................... 108 Dadyanos Whitley, 1951.......................... 110 Derepodichthys Gilbert, 1895..................... 113 Eucryphycus gen. nov............................ 116 Exechodontes DeWitt, 1977........................ 119 Hadropogonichthys Fedorov, 1982.................. 122 Iluocoetes Jenyns, 1842......................... 124 Lycenchelys Gill, 1884.......................... 127 Lycodapus Gilbert, 1891...... 132 iv Reproduced with permission of the copyright owner. Further reproduction prohibited without permission. Page Lycodes Reinhardt, 1831......................... 136 Lycodichthys Pappenheim, 1911.................... 140 Lycodonus Goode and Bean, 1883................... 143 Lycogrammoides Soldatov and Lindberg, 1929....... 145 Lyconema Gilbert, 1895.......................... 148 Maynea Cunningham, 1871......................... 150 Notolycodes Gosztonyi, 1977...................... 153 Oidiphorus McAllister and Rees, 1964............. 155 Ophthalmolycus Regan, 1913....................... 158 Pachycara Zugmayer, 1911........................ 160 Phucocoetes Jenyns, 1842........................ 163 Piedrabuenia Gosztonyi, 1977..................... 166 Plesienchelys gen. nov.........................