/IZ /avv^^-.-^ v.? FLORA MALESIANA LIBRARY JUL 20 1977 NEW YORK BOTANICAL GARDEN Presented by B. A. Krukoff to New York Botanical Garden July 1977 SERIES I - SpERMATOPHYTA Flowering Plants Vol. 7, part 4 Dedication - Revisions Addenda - Index INDEX TO REVISED FAMILIES Aceraceae 4: 3, 592 Dipsacaceae 4: 290 Passifloraceae 7: 405 . Pedaliaceae Actinidiaceae s. str. 4: 37 Droseraceae 4: 377; 5: 557 4: 216 Aizoaceae 4: 267 Elatinaceae 4: 203 Pentaphragmataceae . 4: 517 Alismataceae . 5: 317; 6: 915 Epacridaceae 6: 422 Pentaphylacaceae ... 5: 121 Amaranthaceae Ericaceae .... 6: 469, 943 Philydraceae 4: 5 4:69, 593; 6: 915 Erythroxylaceae . .5: 543 Phytolaccaceae .... 4: 229 Ancistrocladaceae ... 4: 8 Fagaceae 7: 265 Pittosporaceae 5: 345 Aponogetonaceae 4: 11; 7: 213 Ficoidaceae 4: 267 Plumbaginaceae . 4: 107 Balanophoraceae .... 7: 783 Flacourtiaceae Podostemaceae . 4: 65; 6: 963 Basellaceae 5: 300 5: 1, 565; 6: 943 Polemoniaceae 4: 195 Batidaceae 5: 414 Flagellariaceae 4: 245 Pontederiaceae 4: 255 Portulacaceae 7: 121 Betulaceae . 5: 207; 6: 917 Geraniaceae 6: 445 Bixaceae s. str 4: 239 Gnetaceae ... 4: 336; 6: 944 Primulaceae 6: 173 5: Burmanniaceae . 4: 13, 592 Gonystylaceae 4: 349 Proteaceae 147 Burseraceae Goodeniaceae Punicaceae 4: 226 5:209, 567; 6:917 5: 335, 567; 6: 949 Restionaceae 5: 416 Butomaceae 5: 118 Haemodoraceae . .5:111 Rhizophoraceae 5: 429; 6: 965 Byblidaceae 7: 135 Haloragaceae 7: 239 Salicaceae 5: 107 Callitrichaceae 4: 251 Hamamelidaceae .... 5: 363 Salvadoraceae 4: 225 Campanulaceae . 6: 107, 928 Hippocr^teaceae .... 6: 389 Sarcospermaceae .... 4: 32 Cannabinaceae .... 4: 223 Hydrocaryaceae .... 4: 43 Saururaceae 4: 47 Capparidaceae 6: 61 Hydrocharitaceae Scyphostegiaceae Caprifoliaceae 5: 381; 6: 952 5: 297; 6: 967 4: 175, 598; 6:928 Hydrophyllaceae .... 4: 207 Simaroubaceae . 6: 193, 968 Cardiopteridaceae . .7: 93 Hypericaceae 8: 1 Sonneratiaceae Celastraceae 6: 227, 389, 930 Icacinaceae 7: 1 4:280, 513; 6: 973 Centrolepidaceae .... 5: 421 Juglandaceae 6: 143 Sparganiaceae 4: 233 Ceratophyllaceae . .4: 41 Juncaceae 4: 210 Sphenocleaceae . .4: 27 Chenopodiaceae Juncaginaceae 4: 57 Stackhousiaceae .... 4: 35 4: 99, 594; 6: 932 Leeaceae 7: 755 Staphyleaceae 6: 49 Clethraceae 7: 139 Lemnaceae 7: 219 Stylidiaceae 4: 529 Cochlospermaceae . .4: 61 Loganiaceae ... 6: 293, 953 Styracaceae 4: 49 Combretaceae Lophopyxidaceae ... 7: 89 Taccaceae 7: 806 4: 533; 5: 564; 6: 932 Malpighiaceae 5: 125 Thymelaeaceae 6: 976 Connaraceae . 5: 495; 6: 933 Martyniaceae 4: 216 4: 349; 1, Convolvulaceae Molluginaceae 4: 267 Trapaceae 4: 43 4: 388, 599; 5: 558; 6: 936 Moringaceae 4: 45 Trigoniaceae 4: 59 Corynocarpaceae Myoporaceae 4: 265 Turneraceae 4: 235 4: 262; 5: 557 Myricaceae 4: 277 Typhaceae 4: 243 Crassulaceae 4: 197 Najadaceae 6: 157 Umbelliferae Cyperaceae 7: 435 Nyctaginaceae 6: 450 4: 113, 595; 5: 555; 6: 983 Datiscaceae 4: 382 Nyssaceae 4: 29 Valerianaceae 4: 253 Dichapetalaceae 5: 305; 6: 941 Ochnaceae 7: 97 Violaceae 7: 179 Dilleniaceae 4: 141 Oxalidaceae 7: 151 Xyridaceae ... 4: 366, 598 Dioscoreaceae 4: 293 Papaveraceae 5:114 Zygophyllaceae . .4: 64 TAXONOMICAL REVISIONS REPUBLIK INDONESIA REPUBLIC OF INDONESIA LEMBAGA ILMU PENGETAHUAN INDONESIA (L.I. P.I.) INDONESIAN INSTITUTE OF SCIENCES FLORA MALESIANA BEING AN ILLUSTRATED SYSTEMATICACCOUNTOFTHEMALESIAN FLORA / INCLUDING KEYS FOR DETERMINATION I DIAGNOSTIC DESCRIPTIONS I REFERENCES TO TH E LITE RATU RE / S YNO N YM Y I A N D DISTRIBUTION / AND NOTES ON THE ECOLOGY OF ITS WILD AND COMMONL Y CULTIVATED PLANTS PUBLISHED UNDER THE AUSPICES OF LEMBAGA BIOLOGI N ASION AL BOTANIC GARDENS OF INDONESIA / BOGOR /JAVA AND OF THE RIJKSHER BARIUM / LEYDEN /NETHERLANDS PREPARED ON AN INTERNATIONAL CO-OPERATIVE BASIS UNDER THE SUPERVISION OF SEVERAL DIRECTORS OF BOTANIC G A RDENS /KEEPERS OF HERBARIA AND VARIOUS PROMINENT BOTANISTS FOR THE PROMOTION OF BOTANICAL SCIENCE AND THE CULTURAL ADVANCEMENT OF THE PEOPLES OF SOUTH-EASTERN ASIA TO THE SOUTHWEST PACIFIC REGION SERIESI ^KCT^;;^^ VOLUME? SPERMATOPHYTA GENERAL EDITOR: Dr C. G. G. J. VAN STEENIS DIRECTOR OF THE FOUNDATION 'FLORA MALESIANA' PUBLISHED BY NOORDH OFF INTERNATIONAL PUBLISHING. LEYDEN THENETHERLANDS PRINTED IN THE NETHERLANDS 1972-1976 COPYRIGHT 1976 All rights reserved, including the right to reproduce this hook or parts thereof in any form ISBN 90 286 0615 7 PUBLICATION DATES Part 1 26 Jan. 1972 Part 2 12 Jan. 1973 Part 3 13 Sept. 1974 Part 4 24 June 197(X Printed in the Netherlands Flora Malcsiana Volume 7 Volume 7, preliminary matter, page (4), under publication dates, last line, change 1975 into 1976. CONTENTS Title-page <3) Contents <5) Dedication *7) Abbreviations and signs (I5j TAXONOMICAL REVISIONS in alphabetical sequence Aponogetonaceae by U.V^.'E.. van ^xuggQn 213 Balanophoraceae by B. Hansen 783 Byblidaceae by C.G.G.i. van SiQQms. 135 Cardiopteridaceae by H. Sleumer 93 Clethmceae by H. Sleumer 139 Cyperaceae (to be continued) by J. H. Kern 435 Fagaceae by E. Soepadmo (& C. G. G. J. van Steenis) 265 Haloragaceae by R. van der Meijden & N. Caspers 239 Icacinaceae by H. Sleumer 1 Leeaceae by C. E. Ridsdale ^55 Lemnaceae by F. van der Plas 219 Lophopyxidaceae by H. Sleumer 89 Ochnaceae by A. Kanis 97 Oxalidaceae by J. F. Veldkamp 151 Passifloraceae by W. J. J. O. de Wilde 405 Portulacaceae by R. Geesink 121 Taccaceae by E. Drenth 806 Violaceae by M. Jacobs & D. M. Moore 179 ADDENDA to volumes 4, 5, 6, and 7 Addenda, corrigenda et emendanda by C. G. G. J. van Steenis and collaborators 820 INDEX Index to scientific plant names by M. J. van Steenis-Kruseman 833 Dedicated to HERMAN JOHANNES LAM DEDICATION The completion of the seventh volume of this Flora gives me the occasion to dedicate this volume to Herman Johannes Lam, who from the beginning was intimately connected with the taxonomical study of the flora of the Malesian region, adopted the working team, provided for it a permanent niche in his institute, and finally played an important role when the perpetuating of its existence was threatened in 1958. Herman Lam was born in Veendam, January 3rd, 1892. His father was an organic chemist and taught chemistry at Veendam. There was a possibility that he would be attached to the Uni- versity at Groningen, but he accepted a new post in Rotterdam, in 1893, to set up the first municipal food-inspection department in Holland; this stood model for such inspections annex laboratories in other places. He also had a major share in the realisation of the Dutch 'Codex alimentarius'. Thus, it was at Rotterdam that Herman grew up and received a 'classical' education at the 'Gymnasium Erasmianum' (1904-1911). Though he did not have an enthusiastic teacher he decided to study natural history, at his father's suggestion at the University of Utrecht. This study lasted from 1911 1919. Just before the first World War (1914-1918) he managed to get his B.Sc, but during this war he was in military service part of each year. During the winter months he was allowed to pursue his academic studies, thanks to the support of his biology professors. It was quite an achievement to have this study crowned by a thick thesis (April 7th, 1919) within eight years under these difficult circumstances, as such would be the least required in peace-time. From this it can be deduced that Herman was an eager, devoted, and zealous personality, and during all his life he came up to this mark. In biology he felt not attracted or fit to devote his research to experimental work, and so his principal teacher became professor A. A. Pulle. As he had himself ideas to make a career in the tropics he got a training in acquiring form knowledge with plants of Surinam. Pulle, who tried to promote taxonomy of tropical plants at Utrecht, then the only centre in Holland where this was initiated, selected for him the family Verbenaceae of the Netherlands Indies, as Lam had in view to make a career there. The thesis was entitled: 'The Verbenaceae of the Malayan Archipelago, together with those of the Malayan Peninsula, the Philippines, the Bismarck Archipelago, and the Paiau, Marianne and Caroline Islands'. With these extensions beyond the boundaries of the former Netherlands Indies proper, he set a geographical standard delimitation which closely approaches the area covered by Flora Malesiana. Owing to the restricted wartime communications his voluminous thesis could only be based upon the materials in the herbaria of Utrecht, Leyden, and Berlin- Dahlem, without access to those at Kew and especially those at Bogor. In the brief chapter on plant geography he showed his interest in chorology and dispersal and the probable genesis of ranges. This was also reflected in the attached 'Stellingen' (theorems) on the Philippines as an area of junction of dispersal lines of Verbenaceae, origin and dispersal of Cocos and his defence of the then recently posed theory on continental drift by Wegener (1917), then a novel and rather wild idea, on which he had to give a colloquium for his professor Nierstrasz. This just fitted him, as Lam felt always attracted by new ideas and theories. In the brief chapter on taxon- omy he paid attention to affinities of genera, their place in the system, and derivation from others. This, again, is stressed in the 'Stellingen' where he posed an assumed polyphyletic derivation of Geunsia from Callicarpa\ and, furthermore, he made a plea for biosystematics and delimitation of taxa by means of incompatibility. In both fields he was obviously led to (1) This was premature jumping to conclusions. The disentangling of the Callicarpa-Geunsia complex is not yet solved and will require a great deal of detailed routine research, as I have recently pointed out. (7) Flora Malesiana deductions and theoretical reasoning. This characteristic facet of his scientific ambition, to synthesize main issues in addition to collecting factual material, was possibly to a certain degree due to his professor in zoology, H.