Fossil Record 13 (1) 2010, 35–152 / DOI 10.1002/mmng.200900010 The ammonoids from the Argiles de Teguentour of Oued Temertasset (early Late Tournaisian; Mouydir, Algeria) Dieter Korn*,1, Jrgen Bockwinkel2 and Volker Ebbighausen3 1 Museum fr Naturkunde Berlin, Invalidenstraße 43, 10115 Berlin, Germany. E-mail: [email protected] 2 Dechant-Feinstraße 22, 51375 Leverkusen, Germany. E-mail: [email protected] 3 Engstenberger Hhe 12, 51519 Odenthal, Germany. E-mail: [email protected] Abstract Received 12 July 2009 The ammonoids from the Teguentour Shales (Tournaisian, Early Carboniferous) of Accepted 19 September 2009 Oued Temertasset (Mouydir, Algeria) are described monographically. The following Published 19 February 2010 new ammonoid taxa are introduced: Imitoceras dimidium n. sp., Imitoceras strictum n. sp., Triimitoceras tantulum n. sp., Acrocanites disparilis n. sp., Jdaidites cultellus n. sp., Pericyclus tortuosus n. sp., Pericyclus circulus n. sp., Pericyclus trochus n. sp., Pericyclus intercisus n. sp., Nodopericyclus n. gen., Nodopericyclus circumnodosus n. sp., Nodopericyclus deficerus n. sp., Ammonellipsites serus n. sp., Helicocyclus for- mosus n. sp., Helicocyclus inornatus n. sp., Helicocyclus laxaris n. sp., Ouaoufilalites creber n. sp., family Temertassetiidae n. fam., Temertassetia n. gen., Temertassetia temertassetensis n. sp., Temertassetia secunda n. sp., Temertassetia decorata n. sp., Temertassetia coarta n. sp., Jerania n. gen., Jerania jeranensis n. sp., Jerania sicilicula n. sp., Jerania pusillens n. sp., Jerania subvexa n. sp., Jerania persimilis n. sp., Kusinia n. gen., Kusinia falcifera n. sp., Bouhamedites insalahensis n. sp., Muensteroceras subparallelum n. sp., Muensteroceras multitudum n. sp., Follotites n. gen., Follotites folloti n. sp., Follotites stelus n. sp., Follotites flexus n. sp., family Rotopericyclidae n. fam., Eurites permutus n. sp., Eurites doliaris n. sp., Mouydiria n. gen., Mouydiria mouydirensis n. sp., Mouydiria scutula n. sp., Rotopericyclus kaufmanni n. sp., Roto- pericyclus rathi n. sp., Rotopericyclus wendti n. sp., Rotopericyclus lubesederi n. sp., subfamily Dzhaprakoceratinae n. subfam., Dzhaprakoceras punctum n. sp., Dzhapra- koceras amplum n. sp., Dzhaprakoceras vergum n. sp., Dzhaprakoceras biconvexum Key Words n. sp., Progoniatitinae n. subfam., Progoniatites uncus n. sp., Progoniatites pilus n. sp., Progoniatites paenacutus n. sp., Progoniatites globulus n. sp., Habadraitinae Ammonoidea n. subfam., Habadraites n. gen., Habadraites weyeri n. sp., Habadraites supralatus Mississippian n. sp., Primogoniatites n. gen., Primogoniatites fundator n. sp., Antegoniatitinae Carboniferous n. subfam., Antegoniatites n. gen., and Antegoniatites anticiparis n. sp.. The ammo- North Africa noids occur in three successive assemblages and are the richest ammonoid faunas of taxonomy the time interval worldwide. Introduction early Late Tournaisian, and most of the assemblages were rather poor in species numbers. The discovery of The Middle and early Late Tournaisian was regarded for an ammonoid fauna of this age with eleven new species a long time as an example of a period of low-diversity in the Anti-Atlas of Morocco, however, showed that in the evolutionary history of the Ammonoidea (Kono- there is a potential for a much more detailed overview valova 2005). At the beginning of the 21st century, for of the ammonoid evolution of that time interval (Korn instance, only about 25 species were known from the et al. 2003a). * Corresponding author # 2010 WILEY-VCH Verlag GmbH & Co. KGaA, Weinheim 36 Korn, D.: Ammonoids from the Argiles de Teguentour Middle and Late Tournaisian ammonoids are known scribed some individuals of Progoniatites, which prob- from many places in the world, but occur often in low- ably came from the same locality, under the name Go- diversity assemblages. An exception is the terminal niatites cf. crenistria. Later, Conrad (1984) presented a Tournaisian Erdbach Limestone of Germany (Holzapfel number of well-preserved specimens from the Oued Te- 1889; Schindewolf 1951), from which comparatively mertasset and other localities. These belong to various species-rich occurrences have been described. Time- genera such as Imitoceras, Rotopericyclus, Pericyclus, equivalents of this fauna are also very diverse, as seen Jerania, and Progoniatites. in the occurrences of Kyrgyzstan (Librovitch 1927; Po- Considerably rich early Late Tournaisian ammonoid pov 1965, 1968) and North Russia (Kusina 1980), for faunas were reported from the following geographic instance. areas: Early Late Tournaisian assemblages are much smaller, Indiana – The fauna of the Rockford Limestone (Hall and are limited in their geographic distribution. They are 1860; Miller 1891; Gutschick & Treckman 1957) known from the classic occurrences in the Calcaire de shows some resemblance with the newly described Calonne of Belgium (de Koninck 1844, 1880; Delpine assemblages. The Rockford fauna is composed of 1940), the ‘Carboniferous Limestone’ of Ireland (Crick the genera Imitoceras, Gattendorfia, Prodromites, 1899; Foord 1901, 1903), and the Rockford Limestone as Muensteroceras, and Protocanites. well as the Marshall Sandstone of the North American Kentucky – Of the three Osagean ammonoid assem- Midcontinent (Hall 1860; Miller & Collinson 1951; Mill- blages (Work & Mason 2003, 2004, 2005), it is par- er & Garner 1955). An outstanding site is the Borden ticularly the Nada Member of the Borden Formation Formation of Kentucky, which yielded well-preserved that is similar to the occurrences at Oued Temertas- ammonoids from three successive faunal complexes set. The fauna contains Polaricyclus and Winchello- (Work & Mason 2003, 2004, 2005). ceras (Work & Mason 2003). While Early Tournaisian (Ebbighausen et al. 2004; Ireland – Of the rich ammonoid fauna described by Bockwinkel & Ebbighausen 2006; Ebbighausen & Crick (1899) and Foord (1901, 1903), particularly Bockwinkel 2007) and Late Visan (Korn et al. 1999, the material from St. Doulagh’s (Co. Dublin) which 2007; Klug et al. 2006; Korn & Ebbighausen 2008) shows a close resemblance with a distinct horizon ammonoid Faunas from North Africa have been out- within the Argiles de Teguentour. The species lined in several monographs, the state of knowledge (names given by Foord and revised names) known of middle or early Late Tournaisian ammonoids from from this locality are the following: that region is still limited. A rich early Late Tournai- sian fauna was recently described by Korn et al. Pericyclus Foordi = Rotopericyclus foordi (2003a, 2005, 2007) from Bouhamed and Jdaid in the Pericyclus subplicatilis = Rotopericyclus subplicatilis Anti-Atlas of Morocco, being with twelve species the Pericyclus Bailyi = Rotopericyclus bailyi most diverse assemblage known so far from this time Pericyclus rotuliformis = Rotopericyclus rotuliformis unit with the genera Irinoceras, Triimitoceras, Pericy- Glyphioceras (Beyrichoceras) corpulentum clus, Orthocyclus, Helicocyclus, Jerania, Muenstero- = Eurites corpulentus ceras, Mouydiria, Progoniatites, Becanites, and Mero- Glyphioceras (Beyrichoceras) truncatum canites. In contrast to most of the other places, where = Dzhaprakoceras hibernicum the specimens occur in calcareous sedimentary rocks, the assemblage from Bouhamed is composed of Belgium – Two rock formations yielded ammonoids of shales, in which the specimens are preserved as limo- early Late Tournaisian age, the Calcaire de Vaulx nitic steinkerns. and the Calcaire de Calonne (de Koninck 1844, Ammonoids of early Late Tournaisian age have been 1880; Delpine 1940). known from the Mouydir in Algeria for decades. Ter- mier & Termier (1950) figured ‘Pericyclus rotuliformis’ The Calcaire de Vaulx yielded the following species (= Rotopericyclus kaufmanni n. sp.) and ‘Pericyclus di- (determinations by Delpine and revised names): latatus’(=Pericyclus trochus n. sp.), collected in the Acrocanites tornacensis ¼ Acrocanites tornacensis course of oil exploration near Hassi Habadra in the Muensteroceras rotella ¼ Xinjiangites rotella Mouydir. Follot (1951, 1952, 1953) published a fossil Muensteroceras perspectivum ¼ Xinjiangites perspectivus list of an occurrence at Oued Temertasset in the Mouy- “Goniatites”(Pericyclus ?) crenulatus ¼ indet. dir, in which he enumerated several goniatite taxa, Protocanites lyoni ¼ Becanites tornacensis among them the nomen nudum Pericycloides (which was renamed by Korn et al. 2003a into Progoniatites). The Calcaire de Calonne yielded the following species A description of Follot’s specimens, however, has unfor- (determinations by Delpine and revised names): tunately never been published, and Follot’s (1952, pl. 3) schematic drawings of some specimens are not clear Imitoceras rotatorium ¼ Imitoceras rotatorium enough to allow detailed examination. Short reports on Pericyclus princeps ¼ Pericyclus princeps the ammonoid faunas were later provided by Conrad Pericyclus ryckholti ¼ Bouhamedites ryckholti (1966, 1967). Pareyn & Conrad (1968) figured and de- Muensteroceras rotella ¼ Xinjiangites rotella museum-fossilrecord.wiley-vch.de # 2010 WILEY-VCH Verlag GmbH & Co. KGaA, Weinheim Fossil Record 13 (1) 2010, 35–152 37 Pericyclus princeps var. multiplicatus ¼ Temertassetia multiplicata Muensteroceras complanatum ¼ Jerania complanata Pericyclus divisus ¼ Progoniatites divisus The faunal lists show that the Calcaire de Vaulx has some resemblance with the lower assemblage described by Korn et al. (2010a) from the Grs de Kahla supr- ieur of Timimoun. The Calcaire