Quick Guide for the Identification of Maryland Scarabaeoidea Mallory Hagadorn Dr. Dana L. Price Department of Biological Sciences Salisbury University This document is a pictorial reference of Maryland Scarabaeoidea genera (and sometimes species) that was created to expedite the identification of Maryland Scarabs. Our current understanding of Maryland Scarabs comes from “An Annotated Checklist of the Scarabaeoidea (Coleoptera) of Maryland” (Staines 1984). Staines reported 266 species and subspecies using literature and review of several Maryland Museums. Dr. Price and her research students are currently conducting a bioinventory of Maryland Scarabs that will be used to create a “Taxonomic Guide to the Scarabaeoidea of Maryland”. This will include dichotomous keys to family and species based on historical reports and collections from all 23 counties in Maryland. This document should be cited as: Hagadorn, M.A. and D.L. Price. 2012. Quick Guide for the Identification of Maryland Scarabaeoidea. Salisbury University. Pp. 54. Questions regarding this document should be sent to: Dr. Dana L. Price - [email protected] *All photographs are linked to their copyright holder. 1 Table of Contents Families of Scarabaeoidea of Maryland…………………………………….................... 6 Geotrupidae……………………………………………………………………………....... 7 Subfamily Bolboceratinae…………………………………………………………… 7 Genus Bolbocerosoma…………………………………………………....... 7 Genus Eucanthus……………………………………………………........... 7 Subfamily Geotrupinae……………………………………………………………… 8 Genus Geotrupes………………………………………………………........ 8 Genus Odonteus...………………………………………………………….. 9 Glaphyridae................................................................................................................ 9 Genus Lichnanthe.…………………………………………………………... 9 Hybosoridae………………………………………………………………………………… 10 Genus Germarostes…………………………………………………………. 10 Lucanidae…………………………………………………………………………………… 10 Subfamily Aesalinae……………………………………………………………........ 10 Genus Nicagus………………………………………………………………. 10 Subfamily Luncaninae………………………………………………………………. 10 Genus Dorcus…………………………………………………..................... 10 Genus Lucanus…………………………………………………………........ 11 Genus Platycerus……………………………………………………………. 12 Subfamily Syndesinae………………………………………………………………. 12 Genus Ceruchus…………………………………………………………….. 12 Scarabaeidae……………………………………………………………………………….. 13 Subfamily Aphodiinae……………………………………………………………….. 13 Tribe Aegialiini…………………………………………………………………… 13 Genus Aegialia……………………………………………………………….. 13 Genus Caelius………………………………………………........................ 13 *All photographs are linked to their copyright holder. 2 Tribe Aphodiini………………………………………………………………....... 14 Genus Acrossus………………………………………………..................... 14 Genus Agoliinus………………………………………………..................... 14 Genus Aphodius………………………………………………..................... 15 Genus Blackburneus…………………………………………..................... …… 15 Genus Blackburneus…………………………………………..................... 15 Genus Calamosternus………………………………………………………. 15 Genus Chilothorax…………………………………………………………… 16 Genus Colobopterus………………………………………………………… 16 Genus Dialytellus…………………………………………………………….. 16 Genus Dialytes……………………………………………………………….. 17 Genus Flaviellus……………………………………………………………... 17 Genus Irrasinus………………………………………………………………. 17 Genus Labarrus……………………………………………………………… 18 Genus Lechorodius………………………………………………………….. 18 Genus Melinopterus…………………………………………………………. 18 Genus Oscarinus……………………………………………....................... 19 Genus Otophorus……………………………………………………………. 19 Genus Oxyomus……………………………………………………………... 19 Genus Pardalosus…………………………………………………………… 20 Genus Planolinellus…………………………………………………………. 20 Genus Planolinus……………………………………………………………. 20 Genus Pseudagolius………………………………………......................... 21 Genus Stenotothorax………………………………………........................ 21 Genus Strigodius…………………………………………………………….. 21 Genus Teuchestes…………………………………………......................... 22 Tribe Didactyliini…………………………………………………………………. 22 Genus Aidophus……………………………………………………………... 22 Tribe Eupariini……………………………………………………………………. 22 Genus Aphotaenius………………………………………………………….. 22 Genus Ataenius……………………………………………………………… 23 Genus Pseudataenius………………………………………………………. 23 Genus Saprosites……………………………………………………………. 24 Tribe Psammodiini……………………………………………………………….. 24 Genus Pammodius…………………………………………........................ 24 Genus Rhyssemus…………………………………………........................ 25 Genus Pleurophorus………………………………………………………… 25 Subfamily Cetoniinae……………………………………………………………….. 26 Tribe Cetoniini…………………………………………………………………… 26 Genus Euphoria……………………………………………………………… 26 *All photographs are linked to their copyright holder. 3 Tribe Cremastocheilini…………………………………………………………... 28 Genus Cremastocheilus…………………………………………………….. 28 Genus Psilocnemis………………………………………………………….. 28 Tribe Gymnetini………………………………………………………………….. 29 Genus Cotinis………………………………………………......................... 29 Tribe Trichiini……………………………………………………………………... 29 Genus Gnorimella……………………………………………………………. 29 Genus Osmoderma………………………………………………………….. 29 Genus Trichiotinus…………………………………………………………… 30 Genus Trigonopeltastes…………………………………………………….. 31 Tribe Valgini………………………………………………………………………. 31 Genus Valgus………………………………………………......................... 31 Subfamily Dynastinae………………………………………………........................ 31 Tribe Cyclocephalini……………………………………………........................ 31 Genus Cyclocephala………………………………………………………… 31 Genus Dyscinetus…………………………………………………………… 32 Tribe Dynastini…………………………………………………………………… 32 Genus Dynastes……………………………………………………………... 32 Tribe Oryctini……………………………………………………………………... 32 Genus Strategus…………………………………………………………….. 32 Genus Xyloryctes……………………………………………………………. 33 Tribe Pentodontini………………………………………………........................ 33 Genus Aphonus……………………………………………………………… 33 Genus Euetheola…………………………………………………………….. 34 Genus Tomarus……………………………………………………………… 34 Tribe Phileurini…………………………………………………………………… 35 Genus Phileurus……………………………………………………………... 35 Subfamily Melolonthinae……………………………………………………………. 35 Tribe Diplotaxini…………………………………………………....................... 35 Genus Diplotaxis……………………………………………........................ 35 Tribe Hopliini……………………………………………………………………… 36 Genus Hoplia…………………………………………………………………. 36 Tribe Macrodactylini……………………………………………………………... 37 Genus Dichelonyx…………………………………………………………… 37 Genus Macrodactylus………………………………………....................... 38 Tribe Melolonthini………………………………………………………………... 38 Genus Phyllophaga…………………………………………....................... 38 Genus Polyphylla…………………………………………………………….. …………….. 40 Tribe Sericini……………………………………………………………………… 41 Genus Maladera……………………………………………………………... 41 *All photographs are linked to their copyright holder. 4 Genus Nipponoserica……………………………………………………….. 41 Genus Serica…………………………………………………………………. 41 Subfamily Rutelinae…………………………………………………....................... 42 Tribe Anomalini............................................................................................. 42 Genus Anomala……………………………………………………………… 42 Genus Popillia………………………………………………………………... 44 Genus Strigoderma…………………………………………........................ 45 Tribe Rutelini……………………………………………………………………... 45 Genus Cotalpa……………………………………………………………….. 45 Genus Parastasia……………………………………………………………. 45 Genus Pelidnota……………………………………………………………... 46 Subfamily Scarabaeinae……………………………………………....................... 46 Tribe Ateuchini…………………………………………………………………… 46 Genus Ateuchus……………………………………………......................... 46 Tribe Canthonini…………………………………………………………………. 46 Genus Canthon………………………………………………...................... 46 Genus Melanocanthon………………………………………………………. 47 Tribe Coprini……………………………………………………………………… 47 Genus Copris………………………………………………………………… 47 Genus Dichotomius…………………………………………....................... 48 Tribe Onthophagini………………………………………………………………. 48 Genus Onthophagus………………………………………………………… 48 Tribe Phanaeini…………………………………………………………………... 49 Genus Phanaeus……………………………………………....................... 49 Trogidae……………………………………………………………………………………... 50 Subfamily Troginae………………………………………………………………….. 50 Genus Omorgus……………………………………………………………… 50 Genus Trox…………………………………………………………………… 51 References………………………………………………………………………………….. 53 *All photographs are linked to their copyright holder. 5 Table of Contents Families of the Scarabaeoidea of Maryland Geotrupidae Glaphyridae Hybosoridae Geotrupes hornii Lichnanthe vulpina Germarostes globosus Lucanidae Passalidae Lucanus capreolus Odontotaenius disjunctus Scarabaeidae Trogidae Onthophagus taurus Trox spinulosus *All photographs are linked to their copyright holder. 6 Table of Contents GEOTRUPIDAE Subfamily Bolboceratinae Genus: Bolbocerosoma Species: Bolbocerosoma farctum Dichotomous Key: Scarab Beetles of South Carolina Key to Tribe Bolboceratini Page 51 http://bugguide.net/node/view/255129/bgimage Genus: Bolbocerosoma Species: Bolbocerosoma tumefactum Dichotomous Key: Scarab Beetles of South Carolina Key to Tribe Bolboceratini Page 51 http://bugguide.net/node/view/429013/bgimage Genus: Eucanthus Species: Eucanthus lazarus Similar to: E. impressus Dichotomous Key: Scarab Beetles of South Carolina Key to Tribe Bolboceratini Page 51; Skip to 9(5) http://bugguide.net/node/view/345282/bgimage *All photographs are linked to their copyright holder. 7 Table of Contents Genus: Eucanthus Species: Eucanthus subtropicus Dichotomous Key: Scarab Beetles of South Carolina Key to Tribe Bolboceratini Page 51; Skip to 9(5) http://bugguide.net/node/view/209128/bgimage Subfamily Geotrupinae Genus: Geotrupes Species: Geotrupes splendidus Similar to: G. balyi, G. semiopacus Dichotomous Key: Scarab Beetles of South Carolina Key to Tribe Geotrupini Page