Åâðàçèàòñêèé ýíòîìîë. æóðíàë 4(1): 4351 © EUROASIAN ENTOMOLOGICAL JOURNAL, 2005 Ê ïîçíàíèþ ðîäà Brachyta Fairmaire, 1864 (Coleoptera, Cerambycidae) Ñèáèðè è Äàëüíåãî Âîñòîêà Contribution to our knowledge of the genus Brachyta Fairmaire, 1864 (Coleoptera, Cerambycidae) of Siberia and the Far East Ñ.Ý. ×åðíûø¸â, Â.Â. Äóáàòîëîâ S.E. Tshernyshev, V.V. Dubatolov Ñèáèðñêèé çîîëîãè÷åñêèé ìóçåé, Èíñòèòóò ñèñòåìàòèêè è ýêîëîãèè æèâîòíûõ ÑÎ ÐÀÍ, óë. Ôðóíçå 11, Íîâîñèáèðñê 630091 Ðîññèÿ. E-mail: [email protected]. Siberian Zoological Museum, Institute of Systematic and Ecology of Animals, Russian Academy of Sciences, Siberian Branch, Frunze str. 11, Novosibirsk 630091 Russia. Êëþ÷åâûå ñëîâà: æóêè-óñà÷è, Cerambycidae, Brachyta, ôàóíà Ñèáèðè è Äàëüíåãî Âîñòîêà, íîâûé âèä, íîâûé ïîäâèä. Key words: long-horn beetles, Cerambycidae, Brachyta, fauna of Siberia and the Far East, new species, new subspecies. Ðåçþìå. Ïðåäñòàâëåíû ðåçóëüòàòû èññëåäîâà- Reservation) are described. The new species is distin- íèÿ ñåðèéíîãî ìàòåðèàëà Brachyta èç Ñèáèðè è guished by the following characters: larger size, wide Äàëüíåãî Âîñòîêà Ðîññèè. Îáñóæäàåòñÿ íåîäíîðîä- legs, fine adpressed goldish pubescense, spotted pattern íîñòü òàêñîíîìè÷åñêîé ñòðóêòóðû âèäîâîé ãðóïïû on elytra, and endophallic structure of aedeagus pos- B. interrogationis. Îïèñàí íîâûé âèä B. danilevskyi sessing additional small dents near the tip. All speci- sp.n. ñ îñòðîâà Êóíàøèð, îòëè÷àþùèéñÿ áîëåå êðóï- mens which have been previously recorded for Japan íûìè ðàçìåðàìè, óòîëù¸ííûìè íîãàìè, òîíêèì as B. punctata (nec Fald., 1833) are considered to be êîðîòêèì ïðèæàòûì çîëîòèñòûì îïóøåíèåì, ïÿò- B. danilevskyi sp.n. B. interrogationis sochondensis íèñòûì ðèñóíêîì íàäêðûëèé è ñòðîåíèåì ýäåàãó- ssp.n. differs from the nominative form by the spotted ñà, èìåþùèì äîáàâî÷íûå ãðóïïû ìåëêèõ çóáöîâ ó pattern on its elytra, elongate pronotum with strong âåðøèíû. Ê íîâîìó âèäó îòíåñåíû âñå ýêçåìïëÿ- transverse impression on the distal part, and a narrow ðû, óêàçûâàåìûå äëÿ ßïîíèè êàê B. punctata (nec aedeagus. All eastern representatives of B. variabilis Fald., 1833). Èç þãî-çàïàäíîé ÷àñòè ×èòèíñêîé îá- with a distinct and even punctation on their elytra are ëàñòè (Ñîõîíäèíñêèé çàïîâåäíèê) îïèñàí íîâûé discussed as a good subspecies B. variabilis eurinensis ïîäâèä B. interrogationis sochondensis ssp.n., îòëè- (Tsherepanov, 1978) stat.n. A key to Brachyta of Siberia ÷àþùèéñÿ îò íîìèíàòèâíîé ôîðìû ïÿòíèñòûì ðè- and the Far East is provided, together with illustrations ñóíêîì íàäêðûëèé, âûòÿíóòîé êïåðåäè ïåðåäíåñ- of habits and male genitalia of the new taxa. Holotypes ïèíêîé ñ ðåçêèì äèñòàëüíûì âäàâëåíèåì è áîëåå of the taxa described are preserved in the collection of òîíêèì è óçêèì ýäåàãóñîì. Ïðåäëàãàåòñÿ ñ÷èòàòü the Siberian Zoological Museum of the Institute of Sys- âñåõ âîñòî÷íûõ ïðåäñòàâèòåëåé âèäà Brachyta vari- tematics and Ecology of Animals (Novosibirsk, Russia). abilis, îáëàäàþùèõ îòëè÷íîé îò íîìèíàòèâíîé ôîð- ìû ÷¸òêîé óïîðÿäî÷åííîé ïóíêòèðîâêîé íàäêðû- Ïðåäñòàâèòåëè ðîäà Brachyta øèðîêî ðàñïðîñò- ëèé, ïîâèäîì B. variabilis eurinensis (Tsherepanov, ðàíåíû â óìåðåííîé Ïàëåàðêòèêå, è îáû÷íû íà öâå- 1978) stat.n. Ïðèâåäåíà îïðåäåëèòåëüíàÿ òàáëèöà òóùåì ëåñíîì ðàçíîòðàâüå. Ýòî ñòðîéíûå æóêè ñ âèäîâ Brachyta Ñèáèðè è Äàëüíåãî Âîñòîêà, äàíû âûòÿíóòîé ãîëîâîé è ïåðåäíåñïèíêîé, èìåþùåé íà ðèñóíêè ãàáèòóñîâ è ãåíèòàëèé ñàìöîâ âíîâü îïè- áîêàõ çóáöåâèäíûå áóãîðêè, îáû÷íî ñ æ¸ëòûìè íàä- ñûâàåìûõ òàêñîíîâ. êðûëüÿìè ñ ðèñóíêîì èç ÷¸ðíûõ òî÷åê, ÷àñòî ñëè- âàþùèõñÿ â ïðîäîëüíûå ïîëîñû èëè ïîëíîñòüþ Abstract. Results of an investigation of Brachyta çàòåìíÿþùèõ íàäêðûëüÿ, ñ òîíêèìè óñèêàìè, äîñ- from Siberia and the Far East are presented. The com- òèãàþùèìè ïîëîâèíû òåëà è òîíêèìè íåäëèííûìè plexity of taxonomic structure of the B. interrogationis íîãàìè, çàäíèå ëàïêè êîòîðûõ êîðî÷å ãîëåíåé. species group is discussed. A new species, B. danilev- Äî ïîñëåäíåãî âðåìåíè ñ÷èòàëîñü, ÷òî â Ñèáè- skyi sp.n., from Kunashir Island, and a new subspecies ðè è íà Äàëüíåì Âîñòîêå Ðîññèè îáèòàþò îò òð¸õ B. interrogationis sochondensis ssp.n. from the south- [×åðåïàíîâ, 1996] äî ïÿòè [Danilevsky, 1997], à â west of Chitinskaya Oblast (Sohondinsky State ßïîíèè ÷åòûðå âèäà Brachyta [Kusama, Takakuwa, 44 Ñ.Ý. ×åðíûø¸â, Â.Â. Äóáàòîëîâ 1984], íî òàêñîíîìè÷åñêàÿ ñòðóêòóðà ðîäà äî ñèõ ïàðàòèïîâ âíîâü îïèñûâàåìûõ òàêñîíîâ áóäåò ïå- ïîð îñòà¸òñÿ íåâûÿñíåííîé è òðåáóåò ðåâèçèè [Äà- ðåäàíà â ñëåäóþùèå êîëëåêöèè: Çîîëîãè÷åñêèé èí- íèëåâñêèé, 1988].  íàñòîÿùåå âðåìÿ äëÿ Ñåâåðî- ñòèòóò ÐÀÍ, ã. Ñàíêò-Ïåòåðáóðã (ÇÈÍ); êîëëåêöèÿ Âîñòî÷íîé Àçèè èçâåñòíû ñëåäóþùèå âèäû è ïîä- Ì.Ë. Äàíèëåâñêîãî, ã. Ìîñêâà (êÌËÄ). âèäû ýòîãî ðîäà: Brachyta bifasciata bifasciata Ïðè öèòèðîâàíèè ìàòåðèàëà íå ïîêàçàí ïîäâèä Olivier, 1792, Brachyta bifasciata japonica Matsushita, Brachyta bifasciata japonica Matsushita, 1933, îïè- 1933, B. variabilis Gebler, 1817 (= B. eurinensis Tshe- ñàííûé èç ßïîíèè è íå âñòðå÷àþùèéñÿ â Ïðèìî- repanov, 1978), B. breiti Tippman, 1946, B. interroga- ðüå, åãî õàðàêòåðíûå ïðèçíàêè äàþòñÿ â îïðåäåëè- tionis Linnaeus, 1758 (= B. punctata Faldermann, 1833), òåëüíîé òàáëèöå. B. amurensis Kraatz, 1879, B. sachalinensis Matsumu- Èëëþñòðàöèè áûëè âûïîëíåíû ïî ýêçåìïëÿðàì ra, 1911. èç ñëåäóþùèõ ëîêàëèòåòîâ: Èçó÷åíèå áîëüøîãî ñåðèéíîãî ìàòåðèàëà âèäî- B. variabilis variabilis Êåìåðîâñêàÿ îáë., Êóçåäååâî; âîé ãðóïïû B. interrogationis ïîêàçàëî íåîäíîðîä- B. variabilis eurinensis stat.n. Õàáàðîâñêèé êðàé, Áóðåèí- íîñòü ýòîãî òàêñîíà â âîñòî÷íîé ÷àñòè ðåãèîíà. ñêèé çàïîâåäíèê (ðèñ. 2) è ãîëîòèï, ×èòèíñêàÿ îáëàñòü, Íåò ñîìíåíèÿ, ÷òî îïèñàííûé Ôàëüäåðìàííîì èç Øàõòîìà (ðèñ. 3); B. breiti Âîñòî÷íûé Ñàÿí, Ìîíäû; îêðåñòíîñòåé Èðêóòñêà B. punctata ÿâëÿåòñÿ ñèíî- B. interrogationis interrogationis Ôèíëÿíäèÿ, Ïåëêîñå- íèìîì B. interrogationis â ýòîé ÷àñòè àðåàëà ãðóï- íèåììè; ïû. Îäíàêî óæå â Âîñòî÷íîé Ñèáèðè òàêñîí «ðàñ- B. interrogationis sochondensis ssp.n. ãîëîòèï, ×èòèíñêàÿ ïàäàåòñÿ» íà ðÿä âèäîâ, õîðîøî îòëè÷àþùèõñÿ îò îáëàñòü, Ñîõîíäèíñêèé çàïîâåäíèê; B. amurensis Ïðèìîðüå, Êåäðîâàÿ Ïàäü; íîìèíàòèâíîé ôîðìû è ìåëêèìè ðàçìåðàìè (B. sa- B. sachalinensis Ïðèìîðüå, Êåäðîâàÿ Ïàäü; chalinensis, B. amurensis), è îêðàñêîé íîã è óñèêîâ B. danilevskyi sp.n. ãîëîòèï, îñòðîâ Êóíàøèð, Àë¸õèíî. (B. sachalinensis), è, êîíå÷íî æå, îïóøåíèåì òåëà è îêðàñêîé íàäêðûëèé, ïðåæäå âñåãî èõ ÷¸ðíûì Brachyta bifasciata bifasciata (Olivier, 1792) ðèñóíêîì. Ëþáîïûòíî, ÷òî â Ïðèìîðüå òèïè÷íûé Ðèñ. 17(1). B. interrogationis íå âñòðå÷àåòñÿ, è ñóùåñòâóåò ïðåä- ëîæåíèå âîîáùå èñêëþ÷èòü ýòîò âèä èç ôàóíû ßïî- Ìàòåðèàë. ×ÈÒÈÍÑÊÀß ÎÁËÀÑÒÜ: 2##, 4$$ ëåâûé áåðåã ðåêè Áóäþìêàí, 5 êì âûøå óñòüÿ, 1015 êì Þ íèè [Ëîáàíîâ, 2002: ñàéò ÇÈÍ]. Ìåæäó òåì, ÿïîíñ- ñ. Óðþïèíî, 7.7.2002, Ñ.Ý. ×åðíûø¸â, òàì æå, 23.6.2003, Â.Â. êèå ñïåöèàëèñòû ïðèâîäÿò â ñâîåé ëèòåðàòóðå Äóáàòîëîâ; ÏÐÈÌÎÐÜÅ: 3##, 6$$ îêð. ñ Êàìåíóøêà, [Kusama, Takakuwa, 1984] âèä B. punctata, êàê îáû÷- áåðåã ðåêè Ñóïóòèíêè, 28.6.1971, À.È. ×åðåïàíîâ. íûé äëÿ ßïîíèè è ïðèëåæàùèõ îñòðîâîâ. Îáëàäàÿ Ðàñïðîñòðàíåíèå. Âèä ðàñïðîñòðàí¸í îò Âîñòî÷íîé ñåðèéíûì ìàòåðèàëîì ñ îñòðîâà Êóíàøèð è îçíà- Ñèáèðè äî Êîðåè, Ñåâåðî-Âîñòî÷íîãî Êèòàÿ è ßïîíèè. êîìèâøèñü ñ îïèñàíèåì ÿïîíñêèõ æóêîâ, ìû óáå- äèëèñü, ÷òî ýòîò âèä íå òîæäåñòâåí B. interrogati- Brachyta variabilis variabilis (Gebler, 1817) onis, ñèíîíèìîì êîòîðîãî ÿâëÿåòñÿ B. punctata, à Ðèñ. 1, 17(2). çíà÷èò, òðåáóåò íîâîå íàçâàíèå è îïèñàíèå, êîòî- = Evodinus mannerheimii Motschulsky, 1860: 148. Tab. IX, ðîå ìû ïðèâîäèì íèæå êàê B. danilevskyi sp.n. fig. 26. Áîëüøîé ñåðèéíûé ìàòåðèàë B. interrogationis, Ìàòåðèàë. ÞÆÍÛÉ ÓÐÀË: 1$ çàïàäíûé áåð. îç. ñîáðàííûé â 1991 ãîäó â Ñîõîíäèíñêîì çàïîâåäíè- ×åáàðêóëü, 21.6.1994, À.Â. Áàðêàëîâ (ÑÇÌÍ). ÍÎÂÎÑÈÁÈÐÑ- êå íà þãî-çàïàäå ×èòèíñêîé îáëàñòè, ïðåäñòàâëåí ÊÀß ÎÁËÀÑÒÜ: 2##, 3$$ Ìàñëÿíèíñêèé ëåñõîç, ïèõòî- ñîâåðøåííî îòëè÷àþùåéñÿ ôîðìîé êàê ïî ÷¸ðíî- âûé ëåñ, íà öâåòàõ, 929.6.1956, Ã.Î. Êðèâîëóöêàÿ (ÑÇÌÍ); 1# Êàðàñóêñêèé ðàéîí, 1315 êì Ç Êàðàñóêà, ÞÇ îêð. îç. ìó ðèñóíêó íàäêðûëèé, òàê è ïî ñòðîåíèþ ïåðå- Êðîòîâàÿ Ëÿãà, 1.6.1998, Â.Â. Äóáàòîëîâ (ÑÇÌÍ). ÂÎÑÒÎ×- äíåñïèíêè è îïóøåíèþ íîã è íàäêðûëèé. Íåëüçÿ ÍÛÉ ÊÀÇÀÕÑÒÀÍ: 2##, 2$$ îêð. ï. Ãëóáîêîå, 4.5.1994, èñêëþ÷àòü, ÷òî ýòà ôîðìà ïðåäñòàâëÿåò ñîáîé ñà- Â.Ê. Çèí÷åíêî (ÑÇÌÍ). ÀËÒÀÉÑÊÈÉ ÊÐÀÉ: 16##, 20$$ ìîñòîÿòåëüíûé âèä, íî íå èçó÷èâ òèïû âîñòî÷íî- Êàòàíäû, 16.6.1920, Å. Ðîää (ÑÇÌÍ). ÃÎÐÍÛÉ ÀËÒÀÉ: 1#, 3$$ ð. Ïûæà, ï. Îáîãî, íà öâåòàõ îãîíüêîâ è ìàðüèíîãî ñèáèðñêèõ òàêñîíîâ ãðóïïû B. interrogationis, ìû êîðíÿ, 14.6.1969, À.Ç. Åðìîëåíêî (ÑÇÌÍ); 17##, 12$$ îç. ðåøèëè îïèñàòü å¸ êàê ïîäâèä B. interrogationis Òåëåöêîå, îêð. ï. Àðòûáàø, ñ öâåòîâ îãîíüêîâ è ñ òðàâû, sochondensis ssp.n. â íàäåæäå, ÷òî áóäóùàÿ ðåâèçèÿ 24.613.7.1969, À.Ç. Åðìîëåíêî (ÑÇÌÍ). ÊÅÌÅÐÎÂÑÊÀß ãðóïïû ïîçâîëèò óñòàíîâèòü ÷¸òêóþ òàêñîíîìè÷åñ- ÎÁËÀÑÒÜ: 4##, 11$$ îêð. ï. Êóçåäååâî, 28.5.1948, Äÿòëî- êóþ ñòðóêòóðó ðîäà. âà, 1421.6.1969, ñá.? (ÑÇÌÍ). ÕÀÊÀÑÈß: 3##, 2$$ îêð. ï. Áèðèê÷óëü, ñ äóäíèêà, àñòðû è òðàâû, 1.64.7.1969, Íå îáëàäàÿ èñ÷åðïûâàþùèì ìàòåðèàëîì, ìû íå Â.Á. Ïèâåíü (ÑÇÌÍ). ÇÀÏÀÄÍÛÉ ÑÀßÍ: 1#, 4$$ Êóðòó- ìîæåì ñäåëàòü âûâîä î ïðèñóòñòâèè íà Äàëüíåì øèáèíñêèé õð., 10 êì ÑÇ ï. Øèâèëèã, 11001300 ì., 6.7.1990, Âîñòîêå íîìèíàòèâíîãî B. interrogationis, íî ñ÷è- Ñ.Ý. ×åðíûø¸â, Í.À. Ãëàäêåâè÷ (ÑÇÌÍ). ÒÓÂÀ: 1$ Îâþðñ- òàåì âïîëíå âåðîÿòíûì, ÷òî äàííàÿ ôîðìà â ýòîì êèé ðàéîí, ð. Áàðëûê, 20002100 ì, Ä.Â. Ëîãóíîâ (ÑÇÌÍ); 3##, 6$$ Ñàÿíî-Òóâèíñêîå íàãîðüå, îêð. ï. Óø-Áåëäèð, ðåãèîíå íå âñòðå÷àåòñÿ âîîáùå. 3.7.1975, Ò.À. Äóïàë (ÑÇÌÍ). ßÊÓÒÈß: 2##, 3$$ 232 êì Ìàòåðèàëîì äëÿ ïîäãîòîâêè ðàáîòû ïîñëóæèë îò Õàíäûãè ïî òðàññå, 27.6.1985,