1 . !" #$% "&' ()( " * * + ,- &. " /&" & !" 0 1 2&# # # 311# # 4#5 &+6 1 & 3 8 2* & & *+ 22 " 1 ##'1 & *+) 2 2 %''&$* 8 25 & *+ " / " " 2 % &+ 22 1 & !2" &+ 1 & 1 #'1 25 & *+ 22" / # 2 1 , 1 ##'1 */ 8 25 & * + 22" / , 1 #* % % %''& $*25 %''&$*" / 21 # 8 ;<= ;2 *1- '/ >5* + 1(!# 1 "? */ 25' '. " )(%-% *@ 2 8 "" ;3 % # "? , 8@& !, &% ,, %-% 6841 2 A' , 0 & . ' ( 2 + , Kingdom Animalia, Phylum Arthropoda, Class Malacostraca, Order Decapoda, Family Hymenoceridae, Genus Hymenocera '/ /&6 " Hymenocera picta Dana, 1852 /* Hymenocera elegans Heller, 1861 / " Harlequin shrimp, Painted shrimp, Clown shrimp, Dancing shrimp, Harley Shrimp 2 0 * &" , &"5 Hymenocera 1 26 Hymenoceridae 1 . [] : http://th.wikipedia.org/wiki ,-. /- 15 . 2555 2 0 * + ,- %.&*#5 &2 5 '& 2 '&2);d 2#" 0 * + 2 3 !2" picta *#$%2$% 2);d& 1(*" '/ # #, 2 elegans *#'& 22);d )( 0 1 ,- (, 2 #;e. - 1-2 , '/ 2-5 ) *#& #%( 3-60 ; (1-18 )4 k 2-5 l5 - , 2 .2/ !!"" .& - @ 5 ) . 1%#&? 18-26 / %# 1 !* + 2 ! " 1! "!&' &! "1! "& " ,- ! "% , 100-5,000 ; (, " #2 2"* + ! " * 0 " 2"1&5 17-24 ") , (zoea) ,- 2##2*% %/"! 2 ,- 8@'' 2 6 %#'% , " 5 28 - 56 */ 2"1 ) , (zoea) 0 * (postlarva) '/ */, (%-) 5 ,1 !"&%2 .5 " !% (Echinoderms) !2" " & (sea urchin) (brittle sea stars) 2& (sea star) 0 ' 1 22& */ /, 5 8@ " 1 # - ,' (, 21*#&%' #) 0 r &12 1(! # " s#)2'"&& s '/ t%" 2'" s #" 0 %"? 1 , /2 . 2 1;e ,- / %" !& '/ 2 "!"1 ,- " 11 ,- 1 &2 " 5 2 "u 6 - '& Indo-west Pacific 12 & ;e'/ ,- # @ # - Central-Eastern Pacific k*& y12 & "'/" 2 & ,2'" '/ '/'/ ,- "$5 12 ,- '/ ,- &2# @" ' ($*& 1) 2 Sammy De Grave (2010). "Hymenocera Latreille, 1819" . World Register of Marine Species . http://www.marinespecies.org/aphia.php?p=taxdetails&id=204637 . Retrieved August 7, 2011. 3 Sammy De Grave, N. Dean Pentcheff, Shane T. Ahyong et al. (2009). "A classification of living and fossil genera of decapod crustaceans". Raffles Bulletin of Zoology Suppl. 21: 1V109. http://rmbr.nus.edu.sg/rbz/biblio/s21/s21rbz1-109.pdf 4 . FlorentZs guide to the tropical reefs:Harlequin shrimp. [Online] Available http://reefguide.org/harlequinshrimp.html. ,-. /- 22 . 2555 5 . Blue Harlequin Shrimp ( Hymenoceraelegans ). [Online] Available http://www.theaquariumwiki.com/Hymenocera_elegans ,-. /- 22 . 2555. 3 !& 6 ) ;dd y (;e) ( ,- 2) (;e" ( ,- ") (") %* + !&) $*& 1 (Hymenocerapicta Dana, 1852) 2*"1*#& Indo-Pacific, East Africa, Red Sea, Great Barrier Reef, Australia, Philippines, Indonesia, Tuamotus Islands, Hawaiian Islands, Eastern Pacific, Panama, Red Sea 2 Polynesia 7 ($*& 2) $*& 2 2*"1 ( )8 8@ &#, " "? &2 " # / ? /1 * */5'u'#)" !"$ # @" ' - , 2 .2/ !!"" 0 2## compound eyes ($*& 3) # " (ommatidium) 1- 2 " #$*! $*& '. 1 0 2##$* '$* (mosaic image) $*!" 1 2 "u1 #% %/ !' $*! &-5'u" '/ '#'6 ! (rostrum)" - 2#" 0 2 " %/ cephalothorax (head # thorax) 2 & (abdomen)8 6 . ,`ab : ,`ac d /`e`. [] : http://www.zoobras.com. ,-. /- 15 . 2555. 7 . Harlequin Shrimp Hymenocerapicta - Dana, 1852 [Online] Available http://en.microcosmaquariumexplorer.com/wiki/Harlequin_Shrimp. ,-. /- 22 . 2555. 8 `f g f`h , . Phylum Arthopoda. [Online] Available.http://zoo.sci.ku.ac.th/html_T ,-. /- 19 . 2555 4 $*& 3 2 8@& ! 2 9 ' %"2' %/ antennule %"&"u ! %/ antenna 10 & ,' 5 %" (pereiopod) 2 %"2 2!&-' & 0 (claws) -' #%'/ "' 2 0 +e 2 1- 3 %" 0 2" # (pleopod) "" ,- ;3! "22 222* ' (uropod) 8@ &2 .2 uropod 1 2" '/ " "'2 " '. 8@2 "'" *62 *6 !" 1 / . *6 1 5'"" *6 # @5 &*#" *6 1;e& '. ! $5 &2! " &% " *6 8@5 '/ '/2!"1;e 15 ($*& 4) *6 *6 5'"" *6 *6 $*& 4 %2 "'" *62 *6 15 6 1 2* r3 !6(8 *2 # */ 5' 0 ,- 5'" # & ,5 2 " &6&2& 1 #1 + 2&6 ! & ,*"2"* + 5 1 12x24x15 , ,- 30 5'& 0 ' 1 /,1 /2"@ 5 / ! " ## @ " ,-! "5 * 15-20 0 ,- 2 2;3 1! " 0 ) , (Zoea) 1 , #1 *"2"! #5 #") #1 1.5 #6 5" ,- 1,000 " 1;3 0 @ 900-2,000 / 9 Raabe, C. and L. Raabe, .The care and breeding of the Harlequin Shrimp. [Online] Available. http: //www.chucksaddiction.com/harlequinshrimp.html ,-. /- 22 . 2555. 10 j cakg`c f`lmcnop. /f/qc ] http://www.fisheries.go.th/cf-rayong ,-. /- 19 . 2555 5 % , * " * & ,' 13 %/ " 12 (zoea I - zoea XII) 2%- (postlarva) 5 5 * 1) ,u(* 41 ,-5 #% %. 30-33 ppt @'$ ,- 30 oC 2 # 3-20 ; 2 & " 1 , 5'' k* & 2;3 1 & " / 25' & 0 ' ' 0 , .? @ 3 ) 5'' 1 /, u" ,-2 & u" ,- 50% 2 u" ,- 100% & 1 % , */ 5'%#1&-5' 1 # . (, 6 u *2 # 1 #%- .1 / / *61 2553 @ ,"'"6(8&+ #5' & * (, u(6(8 #'- .1&15&2& & 11 Reef Culture Technology &y !&-6(8 , %#& ' *6 5'! "@ 100 u( 3,000 ;' 1%# ! "&! #21 * &% *6 0 20 ;3! " 1-2 " 1u & & *;3& & 1 , 5 30 u( 50 & ,' 11 5 %#1(1 *" ? 12 !!" ## * , &!6(8" ' , *#" !"#@ 2 &! !" " / "%/ & # 0 u(%- (40-60 ) ' 3 2## %/ ; " # & 2' ., ; " # & 2 & " #' .*#" k 19.0, 4.7 2 13.7 ). 13 1 ! ' 14 &- 6(8 # *#"5 ' / 2 ' 1;3 &% %.& 24-34 psu (practical salinity units = ppt) 15 2@'$ ,-1 30 oC *#" (!"# ) 2 1 # 11 j cakg`c f`lmcnopc. (.) /f/qc . [] http://www.fisheries.go.th ,-. /- 15 . 2555. 12 Reef Culture Technology. Ornamental Shrimps. [Online] Available.http://www.rcthawaii.com/research/8.htm. ,-. /- 15 . 2555. 13 ka / r, kss , hk, ,fkj c /t akm u`,r. `a (Hymenocerapicta Dana, 1852). [] :http://www.fisheries.go.th/cf-rayong/images/downloads/shrimp.pdf ,-. /- 27 . 2555. 14 Cheng C., Yen M. and Chen Y. 2010. Effects of temperature, salinity and feeding on nursing of harlequin shrimp (Hymenocera picta ) larvae. Journal of Taiwan Fisheries Research. 18(2): 57-64. [Online] Available : http://www.tfrin.gov.tw/ct.asp?xItem=251715&ctNode=835&mp=3. ,-. /- 27 . 2555. 15 James, T. .Water Salinity Testing. [Online] Available http://www.ehow.com/about_6626245_water-salinity-testing.html. ,-. /- 29 . 2555. 6 * &% 5 6(8% , , & # & -2!"2 " 5' & * (, & ,5 & 0 ) ,2%- (postlarvae) */&-5' & * , & &!u !" *@ ! , 5 %1 , 1!* &% * , 5 ##d */&+$* 5'u -!" 2##&-; *@ #$ % &'() +1%& " 80 l 5 6 22*"1! '" y 2 ;dd u( '&2);d '& 2'& 2 2 311# &6&"& !2" ) 2;dd 1 , !2" #), ; , , 6 2y -' #' 4 0 )/,5'" !2" '$*2> & 16 +1 ,%")/, & u( 200-330 '' 4 "l& & , & & ,, 0.5 u( 2 * + & & , 1,471 140 -' # !" ' 500 #& 15% & " 6 $*5 * */ 5 *@ 8@-% '%/ % 5 2% "' "@% 5 ' 4 24.98 '/%" 48 '' 4 17 * + !&)/,- & 1 1 )(1 #1+ %" 800-1,000 #& 1 , % 1,950 #&, 1.5-2 , % 2,750 #& 2 *"2"* + % 3,830 #&-' # & - 1y ' 4 1 , % 15,950 #&, 1.5-2 , % 19,800 #& 2 *"2"* + % 27,880 #& 18 * ! + *# 2%% > *.6. 2535 '/6&*@ / "& & ! @1 *.6. 2549 -' 5'& & 0 %& ' / #"5"' ( "! , 1 & 8 2'@ */2#*+6 ' / #" 0 & &!1 16 Wabnitz, C., Taylor, M., Green, E., Razak, T. 2003. From Ocean to Aquarium.UNEP-WCMC, Cambridge, UK. [Online] Available http://www.unep-wcmc.org/resources/publications/UNEP_WCMC_bio_series/17.htm. ,-. /- 20 . 2555. 17 . Harlequin Shrimp Hymenocerapicta [Online] Available :http://www.freshmarine.com/harlequin-shrimp.html. ,-. /- 22 . 2555. 18 . [] : http://www.thaionlinemarket.com. ,-. /- 22 . 2555. 7 ** + '/1 - '/&" */6(8 19 )(311# & " 5'" !" u *2 * + ! , */e 1 #1+ 2 8 & 1( 5'" &u * * + !2 u( )(!"! 0 &%%"$5 / ! " 1 1#@A''" &6!2" "%'" &6)( >2*/>&5* + (The Convention on International Trade in Endangered Species of wild Fauna and Flora) '/ !) (CITES) 20 2 '$* */ 8 + 2& *+ (International Union of conservation or Nature and Natural Resources '/ world Conservative Union, IUCN) )( 0 % '" &6&1 , (, e' */ 8 & *+ 21 !"*#"5 ,-5 " crustacean (, & # 2 "2 &1u - 5' 0 %% " 8 22 / .? , 1 u# &6 && *#" Great Barrier Reef & &6 ! #% '!"%( # ,2 "l 1985 *#"2 1 28 u( 13.8 5 "&" 27 l 5 & (, ,2 "l 1998 ,! #&+*1$ + !2" * y % $ 2 * & k* A' (Acanthaster planci ) 23 ($*& 5) 23' 5 + , 0 " ' (& "1 -55 %#% 24 ! /1& . 0 ' 19 ,aag. ,`aq|. 2553. jafrkmc /- ,/,acmakmrg` f.j. 2549 200 . 20 . The CITES species. [Online] Available : http://www.cites.org/eng/disc/species.php. ,-. /- 15 . 2555. 21 IUCN. 2012. IUCN Red List of Threatened Species. Version 2012.2. [Online] Available: www.iucnredlist.org. Search from 22 October 2012. 22 /c. 2543. cm-,`a//~,.a|/,/f- `e`d e `. .c `a (Harlequin Shrimp) [] http://past.talaythai.com/issue/save/save03.html. ,-. /- 29 . 2555 23 . 2555. Researchers report decline of coral covering Great Barrier Reef. [Online] Available http://azstarnet.com. ,-. /- 15 . 2555. 24 Pratchett, M.S. 2005. Dynamics of an outbreak population of Acanthaster planci at Lizard Island, Northern Great Barrier Reef (1995-1999). Coral Reefs, 24: 453-462. 8 $*& 5 A' ()) 2 Great Barrier Reef &u&-A' ( )25 ) u $* # 1*#2 0 1u&- 2 2% &#- ,-!21 # (, */ ,5 & uu * &% 5 5' 0 1 5 *@ 16(8 &% &5 &6!& 2 " &6 *& ,6(8 */'2 &5 * + / ? 5 *@ 0 d " 2 -!5' # &6& ' ( 1u @ " &-5'%@ !"5 % &" 8 %1! -' -' #e " %"& 2e -' '% " 1(%- 1 21&+# */* ## * , */ 0 2##5 *@ "! 25 France-Presse. 2012. Australia admits neglect of Great Barrier Reef. Cosmos. [Online] Available :www.cosmosmagazine.com. ,-. /- 15 . 2555. .