Tanglegram of proctophyllodid feather mites (Proctophyllodidae) and their avian hosts ●● Sphyrapicus varius AD1156 Analloptes sp n ex Xiphorhynchus flavigaster eburneirostris ●● ● AD704 Vingrassia velata ex Anas crecca ●● 276 ● Picoides pubescens 340 277 AD1006 Ingrassia philomachi ex Calidris pugnax ●● 345 275 ●● Dendrocopos major AD600 Xolalges sp n ex Coccyzus americanus ●● 341 ●● Colaptes auratus 274 AD1204 Xolalgoides sp n ex Vireo hypochryseus ●● 342 ●● ●● 344 273 Ramphastos sulfuratus AD1199 Xolalgoides sp n ex Cyanocorax sanblasianus 343 ●● Pandion haliaetus AD765 Ingrassiella ex Catharus fuscater ●● AD1172 Rhamphocaulus sinuatus ex Campylopterus excellens ●● ●● Steatornis caripensis 267 AD1158 Rhamphocaulus sp n aff sinuatus ex Anthracothorax prevostii ●● 338 ●● Campylopterus excellens AD1150 Rhamphocaulus vachoni ex Phaethornis superciliosus ●● 339 337 Ramphocaulini 271 ●● ● 268 Amazilia rutila AD1145 Toxerodectes sp n ex Amazilia candida ● 335 272 ● 270 ●● Amazilia candida AD1236 Trochilodectes bicuspidus ex Phaethornis superciliosus ● 10 Trochilidae AD1138 Trochilodectes buconvexus ex Amazilia rutila ●● 336 ●● Anthracothorax prevostii 331 269 AD1170 Allodectes paulicaulus ex Campylopterus excellens ●● ●● Phaethornis superciliosus 334 10 AD1149 Allodectes wetmori ex Phaethornis superciliosus ●● ●● Prunella atrogularis AD1146 Xynonodectes sp n ex Amazilia candida ●● 332 266 ● 333 ●● Prunella modularis AD1213 Xynonodectes sp n ex Amazilia rutila ● AD648 Montesauria sp n macronous group ex Illadopsis fulvescens ●● split Apodidae/Trochilidae ●● Passer hispaniolensis AD1195 Tyrannidectes berlai ex Myiarchus nuttingi ●● (Apodidae lack proctophyllodids 265 ● Pterodectinae ● Passer domesticus AD1181 Amerodectes sp n aff atyeoi ex Pitangus sulphuratus ●● 329 ● ● 330 316 and not included here) ● Tiaris olivaceus AD1211 Amerodectes sp n ex Myiozetetes similis ● 327 263 ●● Thraupis episcopus AD556 Metapterodectes toxostomae ex Toxostoma rufum ●● 264 AD1040 Amerodectes sp n aff caribaeus ex Empidonax minimus ●● 328 326 262 ●● Thraupis abbas AD1203 Amerodectes sp n ex Vireo hypochryseus ●● 318 ● ● Sporophila torqueola ●● AD1049 Amerodectes sp n aff plumbeus ex Hylocichla mustelina 321 NW ●● ●● 325 258 Cardinalis cardinalis AD1032 Amerodectes sp n aff turdinus and plumbeus ex Catharus fuscescens 324 Passeroidea 261 ●● 222 emberzoid ●● Habia rubica AD680 Amerodectes dumetellae ex Dumetella carolinensis 323 260 AD1174 Amerodectes gracilis ex Psarocolius montezuma ●● 320 317 Passerida ●● Cyanocompsa parellina 322 259 AD1046 Amerodectes sp n aff geothlypis ex Setophaga pensylvanica ●● ●● 319 Pterodectini 4,7 Pheucticus ludovicianus AD760 Pterodectes rutilus ex Hirundo rustica ●● Proctophyllodidae ●● Zonotrichia albicollis AD755 Alaudicola bureschi ex Lullula arborea ●● 208 ●● AD617 Montesauria leioplax ex Cossypha natalensis ●● 242 Emberiza schoeniclus AD1155 Platyacarus sp n aff sittasomi ex Xiphorhynchus flavigaster eburneirostris ●● 253 256 ●● Emberiza spodocephala AD1238 Platyacarus dontocoronius probably ex Dendrocolaptes certhia ●● 315 1 257 ●● 254 Emberiza aureola AD1176 Nycteridocaulus sp n ex Platyrinchus cancrominus ●● ●● Emberiza leucocephalos AD1243 Nycteridocaulus sp n ex Empidonax sp2 flaviventris probably ●● 314 255 312 6 ●● Emberiza citrinella AD1212 Nycteridocaulus sp n aff lamellus ex Myiozetetes similis ●● ●● 313 223 ●● Vermivora peregrina AD1194 Nycteridocaulus lamellus ex Myiarchus nuttingi 308 243 Emberiza AD1139 Anisophyllodes pipromorphae ex Mionectes oleagineus ●● ●● 249 Wilsonia pusilla AD1233 Anisophyllodes sp n aff elaeniae ex Empidonax sp1 ●● 309 250 ●● Dendroica caerulescens AD1179 Anisophyllodes sp n ex Pitangus sulphuratus ●● 310 252 ● ●● 311 248 ● Dendroica pensylvanica AD1237 Anisophyllodes sp n ex Myiarchus tuberculifer 209 251 ●● ●● Dendroica magnolia AD1460 Proctophyllodes detruncatus ex Corvus macrorhynchos detruncatus group 228 AD715 Proctophyllodes detruncatus ex Corvus monedula detruncatus group ●● 306 ●● Seiurus aurocapilla 307 244 AD712 Proctophyllodes detruncatus ex Corvus corone detruncatus group ●● 305 Proctophyllodinae ●● Sturnella magna AD1638 Proctophyllodes vitzthumi ex Sitta europaea detruncatus group ●● ● ●● 245 ● Icterus pustulatus AD1470 Proctophyllodes weigoldi ex Turdus merula weigoldi group 247 304●● 295 Euphonia ●● Agelaius phoeniceus AD1459 Proctophyllodes weigoldi ex Turdus merula weigoldi group 246 AD751 Proctophyllodes scolopacinus ex Scolopax rusticola weigoldi group ●● 302 ●● 8 Psarocolius montezuma AD963 Proctophyllodes aff scolopacinus ex Scolopax minor weigoldi group ●● 303 ● 296 ● Fringilla montifringilla AD638 Proctophyllodes sp ex Sheppardia bocagei caulifer group ●● 210 241 ●● AD743 Proctophyllodes doleophyes ex Phylloscopus trochilus caulifer group ●● Fringilla coelebs 297 ●● 301 ●● Euphonia hirundinacea AD875 Proctophyllodes aff cotyledon ex Phoenicurus erythronotus caulifer group 2 195 AD1469 Proctophyllodes sp n ex Cyanoptila cyanomelana caulifer group ●● 298 ●● Pyrrhula pyrrhula 229 AD997 Proctophyllodes rubeculinus ex Erithacus rubecula caulifer group ●● 299 239 ●● Uragus sibiricus AD812 Proctophyllodes caulifer ex Luscinia svecica caulifer group ●● 300 3 240 224 5 ●● Carpodacus mexicanus AD1472 Proctophyllodes vassilevi ex Acrocephalus scirpaceus caulifer group ●● AD1658 Proctophyllodes sp n ex Anthus campestris caulifer group ●● 230 ●● Carduelis spinus Muscicapoidea+Passeroidea 238 AD1175 Proctophyllodes attenuatus ex Psarocolius montezuma weigoldi group ●● ●● Carduelis pinus Fringillidae 232 237 AD1144 Proctophyllodes habiae ex Habia rubica weigoldi group ●● 294 236 ●● Carduelis tristis AD1249 Proctophyllodes icteri ex Icterus pustulatus weigoldi group ●● ● 174 235 ●● Carduelis cannabina AD1242 Proctophyllodes sp n aff gularis ex Tiaris olivaceus musicus group ● 284 AD912 Proctophyllodes longiphyllus ex Cardinalis cardinalis caulifer group ●● 231 ●● Loxia curvirostra 233 AD1198 Proctophyllodes sp n ex Cyanocorax sanblasianus weigoldi group ●● 292290 ●● 285 211 Carduelis chloris ●● 293 234 AD1251 Proctophyllodes stoddardi ex Vireo flavoviridis caulifer group 291 289 Proctophyllodes ●● Carduelis sinica AD1633 Proctophyllodes longiphyllus ex Cyanocompsa parellina caulifer group ●● ●● Coccothraustes coccothraustes AD1185 Proctophyllodes sporophilae ex Sporophila torqueola anthi group ●● 286 280 ● ●● Motacilla alba AD1239 Proctophyllodes sp n ex Euphonia hirundinacea thraupis group ● AD1178 Proctophyllodes thraupis ex Thraupis episcopus cana thraupis group ●● 287 153 ●● Anthus spinoletta 4 225 AD1154 Proctophyllodes thraupis ex Thraupis abbas thraupis group 288●● 8 227 ●● 226 Anthus trivialis AD1147 Proctophyllodes aff dendroicae ex Cardellina pusilla quadratus group ●● ●● Anthus campestris AD1140 Proctophyllodes dendroicae ex Setophaga magnolia quadratus group 283●● 281● ●● Regulus satrapa AD1148 Proctophyllodes quadratus ex Oreothlypis peregrina quadratus group ● 221 AD1047 Proctophyllodes sp n aff breviquadratus ex Setophaga caerulescens quadratus group 282●● ●● Regulus regulus AD1130 Proctophyllodes sp n ex Sinosuthora webbiana pinnatus group ●● ●● Bombycilla cedrorum AD1002 Proctophyllodes schoenicli ex Emberiza schoeniclus pinnatus group ●● ●● Luscinia svecica AD995 Proctophyllodes sylviae ex Sylvia atricapilla pinnatus group ●● 238 AD1128 Proctophyllodes pinnatus ex Chloris sinica pinnatus group ●● 217 ●● Erithacus rubecula AD873 Proctophyllodes pinnatus ex Carduelis cannabina pinnatus group ●● ●● 275 Phoenicurus phoenicurus ● 270 218 220 AD866 Proctophyllodes pinnatus ex Chloris chloris pinnatus group ●276 212 ●● Phoenicurus erythronotus ● 262 196 215 219 AD998 Proctophyllodes neopinnatus ex Loxia curvirostra pinnatus group ●277 ●● Saxicola rubetra AD970 Proctophyllodes troncatus ex Passer domesticus pinnatus group ●278● 257 251 3 279● 271 ●● Cossypha natalensis AD859 Proctophyllodes aff troncatus ex Passer hispaniolensis pinnatus group ● 213 216 AD734 Proctophyllodes clavatus ex Sylvia borin pinnatus group ●● ●● Sheppardia bocagei AD847 Proctophyllodes clavatus ex Sylvia curruca pinnatus group 274●● 263 ●● Muscicapa striata AD1007 Proctophyllodes cetti ex Cettia cetti pinnatus group ●● 272 214 207 ●● Cyanoptila cyanomelana AD846 Proctophyllodes clavatus ex Acrocephalus schoenobaenus pinnatus group ●●273 197 ● Passerida ●● Catharus fuscater AD1261 Proctophyllodes ludovicianus ex Lanius excubitor pinnatus group ● 205 AD738 Proctophyllodes spini ex Spinus spinus pinnatus group ●● 258 175 211 ●● Catharus ustulatus AD625 Proctophyllodes canadensis ex Sitta canadensis pinnatus group ●● 266 264 212 ●● 210 Catharus fuscescens AD772 Proctophyllodes spini ex Spinus tristis pinnatus group ●●267 253 ●● Hylocichla mustelina AD1054 Proctophyllodes spini ex Spinus pinus pinnatus group ●●268 265 269●● 206 ●● Turdus migratorius AD924 Proctophyllodes sp n vegetans sensu AB66 ex Haemorhous mexicanus pinnatus group Muscicapoidea 209 AD923 Proctophyllodes pheuctici ex Pheucticus ludovicianus pinnatus group ●● 202 ●● Turdus merula 208 AD996 Proctophyllodes ciae ex Emberiza citrinella pinnatus group ●● ●● Turdus philomelos 261 207 AD815 Proctophyllodes ciae ex Emberiza leucocephalos pinnatus group ●● 239 ●● Turdus viscivorus AD1636 Proctophyllodes sp n ex Uragus sibiricus pinnatus group ●● 259 252 260 ●● Dumetella carolinensis AD1027 Proctophyllodes megaphyllus ex Prunella atrogularis pinnatus group ●● 204 AD1132 Joubertophyllodes ampullaceus ex Emberiza aureola Joubertophyllodes group ●● 5 198 ●● Toxostoma rufum 203 AD854 Joubertophyllodes proximus ex Emberiza schoeniclus