AllM SCANNER TEST CHART#2 Spectra 4 Pi ABCDEFGHIJKLM~~OPORSTUWXYZ~~~~~~~~~~~~~~OP~~~~~~Y~". /?SO123456768 Times Roman 4 PT ABCDEFOHIIKLUNOPQRSTLVWXYZ~~~~~~~~~~~~~~~~~P~P~~~~WX~Y/1601234567%9 6 PT ABCDEFGH1JKLMN0PQRSTUVWXYZabcdefgh1jklmnopqstuvwxyz", /1$0123456789 8 PT ABCDEFGHIJKLMNOPQRSTUVWXYZabcdefgh1jklmnopqrstuvwxyz;:",./?$Ol23456789 10 PT ABCDEFGHIJKLMNOPQRSTUVWXYZabcdefghijklmnopqrstuvwxyz;:",./?$Ol23456789 / Century Schoolbook Bold 4 FT ABCDEFCHIJKLINOPQRSTUVWXYZ~~~~~~~~~~II~~~~~::',.'?M~~~S~~~~~~ 6 PT ABCDEFGHIJKLMNOPQRSTUVWXYZabcdefahiiklmno~arstuvwxvz::'~../?$Ol23456l89 Bodoni Italic (H(I,PfLIII/kI &!>OIPX5?L i UXl/.td,fghc,rhuUn nqyr~ii,t lii /ablZlii(lP ABCDEFGHIJKLMNOPQRSTUVIYXYZ(I~~~~~~~~~~I~~~~~~~~IL~~,, /'SO123456789 A BCDEFGHIJKLMNOPQRSTUVWX YZabcdefghijklmnopyrstuuxyz;:",./?$0123456789 ABCDEFGHIJKLMNOPQRSTUVWXYZabcdefgh~klmnopqrstueu;xyz;:';./?SO Greek and Math Symbols AB~IEI~HIK~MNO~~~PITY~~XVLLP)ISS~B~~~A~UO~~~PPPPX~~~-,5*=+='><><i'E =#"> <kQ)<G White Black Isolated Characters 65432 A4 Page 6543210 MESH HALFTONE WEDGES A4 Page 6543210 665432 CONTBIBUTIONS FEOM THE MUSEUi\d OF PALEONTOLOGY (C&& of ConkibvtiOne from the Yweum of Gedogg) UNIVERSITY OF MICHIGAN THE COLOR PATTERNS OF FOSSIL CEPHALO- PODS AND BRACHIOPODS, WI!I'H NOTES ON GASTEROPODS AND PELECYPODS BY AUG. F. FOERSTE UNIVERSITY OF MICHIGAN ANN ARBOR ROCHESTER INSTITUTE OF TECHNOLOGY, ONE LOME CT W s E38L SEE 9 ~~~~ 2358 zgsp EH2 t 3ms 8 2 3 & sE2Z 53EL B83L BE3 9 2::: 2::: 285 9 gg,Bab EE 2 t s3zr BBE & :/; E 3 5 Z 32EL d SB50 COhTRIBUTIONS FROM THE MUSEUM OF PALEONTOLOGY (Continuationof Contributions from the Museum of Geology) UNIVERSITY OF MICHIGAN Editm: EUGENES. MC~AR~EY The series of contributions from the Museum of Paleontology was inaugurated to provide a medium for the publication of papers based entirely or principally upon the collections in the Museum. When the qhmber of ages issued is sdcient to make a volume, a title-page and a tabf e of contents will be sent to libraries on the mailing list, and also to individuals upon request. Communica; tions with reference to exchange or purchase of copies should be directed to the Librarian, General Library, University of Michi- gan. The Stratigraphy and Fauna of the Hackberry Stage of the U per Devonian, by Carroll Lane Fenton and Mildred ~gamsFenton. Pages xi + 260, 45 plates, 9 text figures and 1 map. Cloth. $2.75 net. VOLUMEI1 Consisting of 14 papers. With 41 plates, 39 text figures and 1map. Pp. ix + 240. Cloth $3.00. Postage extra. Parts sold separately in paper covers 1. A Possible E lanation of Fenestration in the Primitive Reptilian Sku"E , with Notes on the Temporal Region of the Genus Dimetrodon, by E. C. Case. Pages 1-12, with 5 illustrations. Price $.3O. 2. Occurrence of the Collingwood Formation in Michigan, by R. Ruedemann and G. M. Ehlers. Pages 13-18. Price, $.15. 3. Silurian Ce halopods of Northern Michigan, by Aug. F. Foerste. %'ages 19-104, with 17 plates and 2 text figures. Price, $I.(?. 4. A Specimen of Stylemys nebrascensis hidy, with the Skull Preserved, by E. C. Case. Pages 87-91, with 7 text figures. Price, $20. 5. Note on a New Species of the Eocene Crocodilian Allognatho- suchus, A. wartheni, by E. C. Case. Pages 93-97, with 1 plate and 1 text figure. Price, $20. 6. Two New Crinoids from the Devonian of Michigan, by G. M. Ehlers. Pages 99-104, with 1 plate. Price, $.20. 7. New Brachiopods from the Warsaw Formation of Wayne County, Kentucky, by G. M. Ehlers and M. S. Chang. Pages 105-111, with 1 plate. Price, $20. (Continued on inside of back cover) VOL. 111, NO. 6, pp. 109-150 (5 pls.) FEB~UAEY20,1930 THE COLOR PATTERNS OF FOSSIL CEPHALO- PODS AND BRACHIOPODS, WITH NOTES ON GASTEROPODS AND PELECYPODS By AUG. F. FOERSTE CONTENTS Acknowledgments I. Introduction 11. Brachiopods List of species 111. Pelecypods IV. Gasteropods List of Paleozoic species V. Cephalopods List of American species 1. Breviconic Ordovician, Silurian and Devonian cephalopods List of Bohemian Silurian cyrtoceroids 2. Phragmoceroids 3. Ordovician and Silurian orthoceroid cephalopods 4. Devonian and Carboniferous orthoceroid cephalopods 5. Carboniferous cyrtoceroid cephalopods 6. Silurian trochoceroid cephalopod 7. Coiled cephalopods from Carboniferous, Liassic and Tertiary strata VI. Crinoid and trilobite VII. Earliest appearance of color markings in different groups of animals VIII. Color markings as an index of relatively shallow seas Bibliography . ACKNOWLEDGMENTS THE incentive toward the studies recorded in this paper was furnished by the loan from Professor E. C. Case of the Uni- versity of Michigan of several specimens showing color patterns. Two of these were specimens of Orthoceras showing transverse zigzag color lines and one was a fragment of limestone retaining parts of two valves of a terebratuloid brachiopod, both of which showed concentric color lines. These were supplemented by a loan from Professor Carl Dunbar of Yale University of 109 110 Aug. F. Foerste several specimens of Orthoceras, belonging to two species, both of which showed color markings, one of these having zigzag lines, and the other, relatively straight transverse ones. Finally, Dr. H. Hedstrom of the Geological Survey of Sweden loaned me two specimens of a small erect cyrtoceroid in which the steeply in- clined color lines formed a zigzag pattern only on the ventral side of the conch. I had previously been permitted by Dr. Jan Koliha to study the unrivaled collection of cephalopods described by Barrande, a considerable number of which show color markings; similar markings occur also on specimens formerly borrowed from Dr. Rudolf Ruedemann and Professor T. E. Savage. To all of these I here express my appreciation of the favors so generously granted. The following studies are concerned chiefly with color markings in cephalopods and brachiopods, but a few notes on gasteropods and pelecypods are added, enough to give some notion of the rela- tive number of color-marked species known at present and their stratigraphic distribution. The bibliography at the end includes all the more important papers bearing on this subject, especially those listing species from the widely scattered literature. Among these the papers by Deecke, Leidhold, Newton, Oppenheim and Richter will be found very useful. Fossils retaining evidences of former coloration are always of interest, especially when they belong to the remote past. In fossils of more recent age the original colors are frequently pre- served, but in earlier fossils, especially those of Paleozoic age, the former presence of colors is usually indicated only by varying shades of light and dark. Sometimes these traces of color are in the form of dots or blotches, irregularly scattered, or arranged according to some definite pattern. At other times they may take the form of radiating or concentric lines or bands, or of combinations of the two. Occasionally the lines are of a zigzag form, or the patterns may be even more intricate. In the more recent fossils all the colors at present known from animals of a similar type are to be expected, but in fossils from Cephalopods and Brachiopods Paleozoic strata the colors usually found are codned to various shades of brown or are nearly black. Occasionally the browns verge on red. In some cases there is evidence that the color shown by a particular fossil is determined in some manner by the mineral matter present in the inclosing matrix. Occasionally this may be indicated by a single individual. For instance, specimens have been found which possessed brownish color markings where preserved in whitish limestone, but which had almost black mark- ings where extending into a blackish matrix. All traces of color tend to disappear with the increasing age of a deposit of fossils, especially during the recrystallization of the calcareous components of the shell. Hence color patterns are retained more frequently by the more recent species. This is shown especially by the gasteropods. A total of 293 species of gasteropods has been recorded as retaining color markings. Of these, 224 species occur in the Tertiary, 25 in the Mesozoic, and 44 in the Paleozoic. Even among Paleozoic strata great differences are shown, the Carboniferous retaining 33 species of gasteropods with color markings; the Devonian, 5 species; the Silurian, 3 species; and the Ordovician, also 3 species. In cephalopods, on the contrary, almost all color-marked species belong to the Paleozoic. This is readily understood when it is remembered that color markings are extremely rare among all the strongly coiled cephalopods, both nautiloid and ammonoid. They are not numerous among straight conchs, and they appear to occur most frequently among the breviconic cyrtoceroids, which are known only from Paleozoic strata. In a similar manner, only two brachiopods preserving color patterns have been recorded from the Tertiary. But this type of life was much more abundant in Paleozoic and Mesozoic times, from which a larger number of species with color patterns is reported. The general vertical distribution of the different types of shelI- bearing animals is indicated on the following page. The very imperfect character of the data secured so far is shown by the fact that of the 33 color-marked species of gastero- 112 Aug. F. Foerste pods recorded at present from the Carboniferous of the entire world, 15 were discovered by J. Brookes Knight only recently, and at a single horizon, in the Pennsylvanian division of the Carboniferous, near St. Louis, Missouri. In a similar manner, of the 25 color-marked species of cephalopods known so far from the Silurian of the entire world, 22 were found in a relatively small area north of Prague, formerly in Bohemia, now a part of Czecho- slovakia. Similar rich but isolated localities probably await dis- covery elsewhere. Tertiary Mesozoic Paleozoic Total Brachiopods . 2 10 11 23 Pelecypods . 6 12 7 25 Gasteropods . 224 25 44 293 C2phalopods.. 1 1 39 41 Total. 233 48 101 382 The imperfection of our knowledge regarding color markings in fossil shells is shown also by the irregular vertical distribution of the gasteropods in the Tertiary and Mesozoic.