FERNANDO ANGELINI WORLD CATALOGUE OF THE TRIBE AGATHIDIINI (COLEOPTERA, LEIODIDAE, LEIODINAE) ESTRATTO dagli ANNALI del MUSEO CIVICO di STORIA NATURALE “G. DORIA” Vol. 102 - 22 NOVEMBRE 2010 GENOVA 2010 1 FERNANDO ANGELINI * WORLD CATALOGUE OF THE TRIBE AGATHIDIINI (COLEOPTERA, LEIODIDAE, LEIODINAE) INTRODUCTION During the last 30 years, the tribe Agathidiini Westwood, 1838 has been the subject of many taxonomic studies and catalogues (by F. Angelini, J. Cooter, L. De Marzo, H. Hoshina, K. B. Miller, A. F. Newton, S. B. Peck, E. E. Perkovsky, M. Perreau, Z. Švec and Q. D. Wheeler) resulting in a moltiplication of the species number and in a better knowledge of the status and distribution of many taxa. New species have been mostly described from Indo-Malayan and Northern Neotropical Regions, less from Nearctics, Palearctics (especially Eastern Palearctics) and Afrotropics. Meanwhile, about 220 papers were published, of which over 110 including the description of more or less 760 new species and 7 new genera/subgenera, and 6 devoted to systematic revisions and geogra- phical catalogues (WHEELER 1979, ANGELINI 1995c and 2004, PER- REAU 2004, WHEELER & MILLER 2005, MILLER & WHEELER 2005b). Nowadays, in my opinion the knowledge of world Agathidiini is quite satisfactory. Furthermore, I presume that in future new spe- cies will be described mainly from Northern Neotropics, from some area of Indo-Malayan Region and from Easterner Palearctics, with a likely 10-15% increase to the present total number. Currently, I think that more studies are still necessary only for a better interpre- tation of Indo-Malayan and Eastern Palearctic species ascribed to the genus Anisotoma and Indo-Malayan species of the genera Cyr- toplastus and Stetholiodes. * Via De Reggio, 142 - 72021 Francavilla Fontana (Brindisi), Italy. E-mail: [email protected] 2 F. ANGELINI Taking in account what has been published during the last 30 years, I regard as necessary (and I am supported in my opinion by epistolary contacts with the above named colleagues) to publish a world catalogue. NOMENCLATORIAL NOTES AND STRUCTURE OF THE CATALOGUE Until January 1st, 2010 the total number of Agathidiini stood at 945 species/subspecies (Table 1). Taxa are listed in the catalogue in alphabetical order; as litera- ture and distributional data have already been published for most species, only essential data are listed here, viz.: author, year and page of description, type locality, type depository (the first abbrevia- tion/coden refers to the depository where the holotype -and often paratypes as well- is housed; further abbreviations, if present, para- types’ depository) and distribution. Terms and boundaries of the geographical regions are those defined by ANGELINI (2004: 137) and LÖBL & SMETANA (2004), par- ticularly for Europe (including European Russia), Caucasus, Siberia and China. In the References 512 titles are quoted; the catalogue lists 1.130 names: 13 valid genera (1 in “incertae tribus”), 6 generic names which are synonyms or preoccupied, 8 valid subgenera, 2 subge- nera names considered synonyms, 945 valid species and subspecies (1 species in “incertae sedis” and 1 species in “incertae tribus”), 156 specific names which are synonyms, homonyms, nomina nuda or referred to renamed taxa. Type depository is given for 933 valid and for 106 synonymized species. On the contrary, it remained unknown for the following 12 species: Agathidium angusticolle Reitter, 1898, A. bescidicum Reitter, 1885, A. insulare Portevin, 1928, A. mexicanum Hendrichs, 1979, A. microps Portevin, 1907, A. oniscoides Palisot de Beauvois, 1817, A. similare Reitter, 1898, Amphicyllis globus Fabricius, 1792, Anisotoma confusa Horn, 1880, A. nigripennis Reitter, 1884, Cyrtoplastus irre- gularis Reitter, 1898 and C. successor Reitter, 1898. The species Anisotoma nigripennis Reitter, 1884 and the genus Cainosternum Notman, 1921 are placed respectively in “incertae sedis” and “incertae tribus”. WORLD CATALOGUE OF THE TRIBE AGATHIDIINI 3 Tab. 1 - Distribution of the species and subspecies of the Tribe Agathidiini. Pale- arctic and Indo-Malayan species of the genus Agathidium are grouped by subgenera. The specialists of Nearctic and northern Neotropical fauna have not adopted this treatment. The totals in the various columns do not correspond with the aggregate for each genus/subgenus because some species are present in two regions. Region northern nearctic palaearctic oriental afrotropical northeastern Total Taxa neotropical australian Afroagathidium 3 3 6 Agathidium 41 59 274 439 1 800 subg. Agathidium 137 240 370 subg. Chaetoceble 1 1 subg. Cyphoceble 20 5 25 subg. Euryceble 6 1 7 subg. Macroceble 14 47 61 subg. Microceble 5 117 121 subg. Neoceble 98 24 115 Agathidium north. 41 41 Neotropical reg. Agathidium 59 59 Neartic region Amphicyllis 2 2 Anisotoma 15 15 22 8 55 Besuchetionella 3 17 20 Cyrtoplastus 11 5 16 Decuria 1 1 1 2 Gelae 5 3 8 Liodopria 2 8 10 Pseudoagathidium 1 9 10 Sphaeroliodes 1 1 2 Stetholiodes 1 5 8 14 Total 62 78 322 488 12 1 945 Moreover, the following new synonyms are established: Agathidium festivum Betta, 1847 = n. syn. of Agathidium nigri- penne (Fabricius, 1792); Anisotoma graminis (Gistel, 1857) = n. syn. of Anisotoma humera- lis (Fabricius, 1792); Anisotoma lucifuga (Gistel, 1857) = n. syn. of Anisotoma orbicu- laris (Herbst, 1791). 4 F. ANGELINI The type of Agathidium pallidum Say, 1824 is lost or destroyed (ORD 1859, MAwdSLEY 1993); then the name was doubtfully associa- ted with Hydnobius matthewsii Crotch, 1874 by HORN (1880: 281) and by HATCH (1929b: 11). Nevertheless, in my opinion the description given by SAY (1824: 91) is sufficiently clear to assume that the type was not referable to Hydnobius matthewsii (now Macrohydnobius), a species belonging to the tribe Sogdini. Then, I think that Say’s species really belongs to Agathidiini. Its type locality was reported as “Engineer Cantonment”, which was the winter encampment of the 1819-20 Expedition, led by Major Stephen H. Long, to explore the southern Rocky Mountains (BEIDLEMAN 1986). Archaeological investigations, carried out by the Nebraska State Historical Society, established the site of the encampment on the bank of the Mis- souri River, a few miles north of present-day Omaha (CARLSON et al. 2004, PECK & COOK 2009). ANGELINI & PECK (2000) presented a phylogenetic study of generic relationships, which was criticized, but without any impro- vements, by MILLER & WHEELER (2005b). Abbreviations follow ARNETT & SAMUELSON (1986) and EVEN- HUIS (2009). For type depository non reported by them, new abbre- viations are given. ADC-MSNG A. Dodero Collection in Museo Civico di Storia Naturale “G. Doria”, Genoa, Italy AMNH American Museum of Natural History, New York, New York, USA APC A. Pütz Collection, Eisenhüttenstadt, Germany ASC A. Smetana Collection, Ottawa, Canada BMNH The Natural History Museum, London, Great Britain CAS California Academy of Sciences, Department of Entomology, San Francisco, California, USA CCC C. Carlton Collection, Fayetteville, USA CMNC Canadian Museum of Nature, Ottawa, Canada CMNH Carnegie Museum of Natural History, Pittsburgh, Pennsylvania, USA CNCI Canadian National Collection of Insects, Ottawa, Canada CNUIC Chungnam National University, Insect Collection, Daejeon, Korea CUIC Cornell University Insect Collection, Ithaca, New York, USA DBC D. Boukal Collection, Ceske Budejovice, Czech Republic DENH University of New Hampshire, Entomological Museum, Durham, New Hampshire, USA DIE Deutsches Entomologisches Institut im ZALF, Müncheberg, Germany WORLD CATALOGUE OF THE TRIBE AGATHIDIINI 5 EEPC E. E. Perkovsky Collection, Kiev, Ukraine EMEC University of California, Essig Museum of Entomology, Berkeley, California, USA EUMJ Ehime University, College of Agriculture, Matsuyama, Japan FAC-MSNG F. Angelini Collection in Museo Civico di Storia Naturale “G. Doria”, Genoa, Italy FGAC F. G. Andrews Collection, CDFA Plant Pest Diagnostics Center, Sacramento, California, USA FMC F. Montemurro Collection, Taranto, Italy FMNH Field Museum of Natural History, Chicago, Illinois, USA FSCA Florida State Collection of Arthropods, Gainesville, Florida, USA GIC G. Israelson Collection, Lund, Sweden HDC H. Daffner Collection, Eching, Germany HFC H. Franz Collection, now in Naturhistorisches Museum, Vienna, Austria HNHM Hungarian Natural History Museum, Budapest, Hungary IBV Institut of Biology, Vladivostok, Russia IRSNB Institut Royal des Sciences Naturelles, Bruxelles, Belgium IZCAS Chinese Academy of Sciences, Institute of Zoology, Beijing, China JCC J. Cooter Collection, Hereford, Great Britain JKC J. Klapperich Collection, Bonn, Germany JLCC J. L. Carr Collection, Calgary, Canada JRAC J. R. A. Baranowski Collection, University of Lund, Lund, Sweden. JRC J. Ru˚žicˇka Collection, Prague, Czech Republic JSC J. Schneider Collection, Prague, Czech Republic JSCC J. Schuh Collection, Klamath Falls, Oregon, USA KSIC K. Stephan Collection, Red Oak, Oklahoma, USA KUEC Kyushu University, Faculty of Agriculture, Fukuoka, Japan MCZ Museum of Comparative Zoology, Harvard University, Cambridge, Massachusetts, USA MGHNL Musee Guymet d’Histoire Naturelle, Lyon, France MHNG Muséum d’Histoire Naturelle, Geneva, Switzerland MNHAH Museum of Nature and Human Activities, Hyôgo, Japan MNHN Muséum National d’Histoire Naturelle, Paris, France MRAC Musée Royal de l’Afrique centrale, Tervuren, Belgium MRSNT Museo Regionale di Scienze Naturali, Turin, Italy MSC M. Schülke Collection, Berlin, Germany MSNG Museo Civico di Storia Naturale