PUBLICACIONES DEL INSTITUTO GEOLÓGICO Y MINERO DE ESPAÑA Serie: CUADERNOS DEL MUSEO GEOMINERO. Nº 9 ADVANCES IN TRILOBITE RESEARCH ADVANCES IN TRILOBITE RESEARCH IN ADVANCES ADVANCES IN TRILOBITE RESEARCH IN ADVANCES planeta tierra Editors: I. Rábano, R. Gozalo and Ciencias de la Tierra para la Sociedad D. García-Bellido 9 788478 407590 MINISTERIO MINISTERIO DE CIENCIA DE CIENCIA E INNOVACIÓN E INNOVACIÓN ADVANCES IN TRILOBITE RESEARCH Editors: I. Rábano, R. Gozalo and D. García-Bellido Instituto Geológico y Minero de España Madrid, 2008 Serie: CUADERNOS DEL MUSEO GEOMINERO, Nº 9 INTERNATIONAL TRILOBITE CONFERENCE (4. 2008. Toledo) Advances in trilobite research: Fourth International Trilobite Conference, Toledo, June,16-24, 2008 / I. Rábano, R. Gozalo and D. García-Bellido, eds.- Madrid: Instituto Geológico y Minero de España, 2008. 448 pgs; ils; 24 cm .- (Cuadernos del Museo Geominero; 9) ISBN 978-84-7840-759-0 1. Fauna trilobites. 2. Congreso. I. Instituto Geológico y Minero de España, ed. II. Rábano,I., ed. III Gozalo, R., ed. IV. García-Bellido, D., ed. 562 All rights reserved. No part of this publication may be reproduced or transmitted in any form or by any means, electronic or mechanical, including photocopy, recording, or any information storage and retrieval system now known or to be invented, without permission in writing from the publisher. References to this volume: It is suggested that either of the following alternatives should be used for future bibliographic references to the whole or part of this volume: Rábano, I., Gozalo, R. and García-Bellido, D. (eds.) 2008. Advances in trilobite research. Cuadernos del Museo Geominero, 9. Instituto Geológico y Minero de España, Madrid, 448 pp. Adrain, J.M. 2008. A global species database of Trilobita: progress, results, and revision of the Treatise. In: Rábano, I., Gozalo, R. and García-Bellido, D. (eds.), Advances in trilobite research. Cuadernos del Museo Geominero, 9. Instituto Geológico y Minero de España, Madrid, 27-28. Cover: Monotaxic cluster of complete specimens of Asaphellus toledanus, a species named after the Toledo province, Spain. Lower Oretanian (mid Darriwilian) slates of the Valongo Formation, Canelas quarry, Arouca, Portugal. © INSTITUTO GEOLÓGICO Y MINERO DE ESPAÑA C/ Ríos Rosas, 23. 28003 Madrid Tel.: +34 91 349 5700, Fax: +34 91 442 6216 www.igme.es NIPO 657-08-018-1 ISBN 978-84-7840-759-0 Depósito Legal: 25280-2008 Fotocomposición: Inforama, S.A. Príncipe de Vergara, 210. 28002 MADRID Imprime: Ibergraphi 2002, S.L.L. Mar Tirreno, 7 bis. 28830 SAN FERNANDO DE HENARES (Madrid) CONTENTS Preface ....................................................................................................................................................... 9 Acknowledgements .................................................................................................................................. 11 THE STATUS OF THE CRUZIANA (TRILOBITE TRACE FOSSIL) STRATIGRAPHY IN WESTERN GONDWANA: THE MIXING OF LOWER AND UPPER ORDOVICIAN ELEMENTS IN THE CENTRAL ANDEAN BASIN OF SOUTH AMERICA..................................................................................................... 13 G.F.Aceñolaza and S. Heredia CRUZIANA YINI YANG, A PERI-GONDWANAN TRILOBITE TRACE WITH NEW RECORDS IN THE ORDOVICIAN OF SOUTH AMERICA AND IBERIA.................................................................................... 19 G.F.Aceñolaza, A.A. Sá and J.C. Gutiérrez-Marco A GLOBAL SPECIES DATABASE OF TRILOBITA: PROGRESS, RESULTS, AND REVISION OF THE TREATISE ............................................................................................................................................ 27 J.M. Adrain SEGMENT VARIATION IN A CAMBRIAN TRILOBITE IN THE CONTEXT OF THE BURGESS SHALE FAUNA ............................................................................................................................................ 29 M. Balint and N.C. Hughes TREMADOCIAN TRILOBITE ASSEMBLAGES FROM THE ARGENTINE CORDILLERA ORIENTAL. A PRELIMINARY ANALYSIS...................................................................................................................... 33 D. Balseiro and L. Marengo ASPECTS OF SOME LATE MIDDLE CAMBRIAN TASMANIAN FAUNAL ASSEMBLAGES ...................... 41 C.J. Bentley and J.B. Jago FLUCTUATIONS IN TRILOBITE PALAEOBIODIVERSITY DURING THE DEVONIAN IN THE ARDENNES MASSIF ...................................................................................................................... 49 A. Bignon, C. Crônier and A. van Viersen MIDDLE CAMBRIAN TRILOBITES OF LA LAJA FORMATION, WESTERN ARGENTINA.......................... 55 O.L. Bordonaro and B.R. Pratt NEW POST-CAMBRIAN TRILOBITE ASSOCIATIONS FROM MONGOLIA................................................ 57 P. Budil, O. Fatka, R. Feist, M. Mergl and C. Minjin DALMANITOID AND ACASTOID TRILOBITES AND THEIR EVOLUTION IN THE PRAGUE BASIN (CZECH REPUBLIC)......................................................................................................................... 67 P. Budil, F. Hörbinger, O. Fatka and M. Mergl LINEAR CLUSTERS OF ARTICULATED TRILOBITES FROM LOWER ORDOVICIAN (ARENIG) STRATA AT BINI TINZOULIN, NORTH OF ZAGORA, SOUTHERN MOROCCO....................................................... 73 B.D.E. Chatterton and R.A. Fortey ORDOVICIAN TRILOBITES FROM VALONGO ANTICLINE: AN HISTORICAL AND SCIENTIFIC HERITAGE ................................................................................................................................................... 79 H. Couto and A. Lourenço 3 A QUANTITATIVE COMPARISON OF THE ONTOGENY OF TWO CLOSELY RELATED TRINUCLEIDAE TRILOBITES FROM UPPER ORDOVICIAN................................................................................................. 85 C. Crônier, A. Delabroye and L. Miclot TRILOBITE BIOSTRATIGRAPHY AND BIODIVERSITY PATTERNS THROUGH THE MIDDLE-UPPER CAMBRIAN TRANSITION IN THE KYRSHABAKTY SECTION, MALYI KARATAU, SOUTHERN KAZAKHSTAN............................................................................................................................................. 91 G.Kh. Ergaliev, V.A. Zhemchuzhnikov, F.G. Ergaliev, L.E. Popov, M. Ghobadi Pour and M.G. Bassett PARDAILHANIA THORAL, 1947 (SOLENOPLEUROPSINAE, TRILOBITA) FROM CAESARAUGUSTAN (MIDDLE CAMBRIAN) OF CADENAS IBÉRICAS (NE SPAIN)................................................................... 99 J. Esteve, R. Gozalo, E. Liñán and J.B. Chirivella Martorell ENROLMENT IN MIDDLE CAMBRIAN SOLENOPLEUROPSINAE TRILOBITES........................................ 105 J. Esteve, S. Zamora, R. Gozalo and E. Liñán PYRITIZED TRILOBITE FAUNAS FROM THE ORDOVICIAN OF NEW YORK STATE: BEECHER’S TRILOBITE BED AND THE WHETSTONE GULF FORMATION NEAR LOWVILLE...................................... 109 Ú.C. Farrell and D.E.G. Briggs EXOSKELETAL CONFIGURATIONS OF CAMBRIAN AND ORDOVICIAN AGNOSTID TRILOBITES: EXAMPLES FROM THE BARRANDIAN AREA OF CZECH REPUBLIC....................................................... 113 O. Fatka, P. Budil, M. Szabad and J. Bruthansová POSITION OF TRILOBITES IN CAMBRIAN ECOSYSTEM: PRELIMINARY REMARKS FROM THE BARRANDIAN REGION (CZECHIA) ........................................................................................................... 117 O. Fatka, M. Szabad, P. Budil and V. Micka ORDOVICIAN TRILOBITE GENUS HUNGIOIDES KOBAYASHI, 1936....................................................... 123 O. Fatka, P. Budil, I. Rábano, A.A. Sá, P. Kraft, U. Linnemann, J. Marek and K. Drost ASSESSING THE HYPOTHESIS OF A THIRD TAGMA IN SCUTELLUID TRILOBITES: ARGUMENTS FROM ONTOGENETIC, FUNCTIONAL, AND EVOLUTIONARY PERSPECTIVES ....................................... 127 R. Feist and R. Lerosey-Aubril DIGGING, SCRATCHING, AND WALKING IN THE STAIRWAY SANDSTONE OF CENTRAL AUSTRALIA: ‘ARTHROPOD’ ICHNOFOSSILS FROM THE ORDOVICIAN ....................................................................... 135 S. Gibb THE CAMBRIAN OF THE CADENAS IBÉRICAS (NE SPAIN) AND ITS TRILOBITES................................. 137 R. Gozalo, E. Liñán, J.A. Gámez Vintaned, M.E. Dies Álvarez, J.B. Chirivella Martorell, S. Zamora, J. Esteve and E. Mayoral EARLY ORDOVICIAN TRILOBITES FROM THE IBERIAN CHAINS, NE SPAIN: A TRIBUTE TO WOLFGANG HAMMANN .................................................................................................. 153 W. Hammann, J.J. Álvaro and D. Vizcaïno MID TO LATE ORDOVICIAN TRILOBITE PALAEOECOLOGY IN A MUD DOMINATED EPICONTINENTAL SEA, SOUTHERN NORWAY......................................................................................... 157 T. Hansen 4 CLADISTIC ANALYSIS OF PTYCHASPIDID TRILOBITES (UPPER CAMBRIAN), AND A NEW HYPOTHESIS OF INGROUP RELATIONSHIPS ........................................................................................... 167 T. Hegna THE FIRST TRILOBITES IN LAURENTIA AND ELSEWHERE...................................................................... 171 J.S. Hollingsworth FENESTRASPIS, AN UNUSUAL DALMANITID TRILOBITE FROM THE LOWER DEVONIAN OF BOLIVIA ................................................................................................................................................ 177 D.J. Holloway and M. da G.P. de Carvalho TRUNK SEGMENTATION OF THE EARLY CAMBRIAN EODISCID TRILOBITE NEOCOBBOLDIA CHINLINICA................................................................................................................................................ 181 P.S. Hong, N.C. Hughes, G. Fusco and X.-G. Zhang MORPHOLOGICAL