NOMENCLATURAL CHANGES IN THE STAPHYLINIDAE (INSECTA: COLEOPTERA) LEE H. HERMAN Curator, Division of Invertebrate Zoology American Museum of Natural History BULLETIN OF THE AMERICAN MUSEUM OF NATURAL HISTORY Number 264, 83 pp. Issued June 21, 2001 Price: $7.60 a copy Copyright q American Museum of Natural History 2001 ISSN 0003-0090 CONTENTS Abstract ....................................................................... 3 Introduction .................................................................... 4 Type Species ................................................................. 5 Leptotyphlinae ............................................................... 5 Megalopsidiinae .............................................................. 6 Omaliinae ................................................................... 6 Staphylininae ................................................................. 8 Steninae .................................................................... 11 Tachyporinae ................................................................ 11 Spelling Changes .............................................................. 11 Osoriinae ................................................................... 11 Oxytelinae .................................................................. 12 Piestinae .................................................................... 12 Staphylininae ................................................................ 12 Umlauts .................................................................... 12 New Combinations ............................................................. 13 Micropeplinae ............................................................... 13 Omaliinae .................................................................. 13 Osoriinae ................................................................... 13 Oxyporinea ................................................................. 13 Oxytelinae .................................................................. 14 Staphylininae ................................................................ 14 Tachyporinae ................................................................ 15 Synonyms .................................................................... 19 Euaesthetinae ............................................................... 19 Micropeplinae ............................................................... 19 Omaliinae .................................................................. 19 Osoriinae ................................................................... 22 Oxytelinae .................................................................. 22 Proteininae .................................................................. 24 Staphylininae ................................................................ 24 Steninae .................................................................... 32 Tachyporinae ................................................................ 34 Homonyms ................................................................... 36 Euaesthetinae ............................................................... 36 Leptotyphlinae .............................................................. 36 Micropeplinae ............................................................... 36 Omaliinae .................................................................. 36 Osoriinae ................................................................... 39 Oxytelinae .................................................................. 41 Staphylininae ................................................................ 42 Steninae .................................................................... 54 Tachyporinae ................................................................ 54 Special Problems .............................................................. 57 Omaliinae .................................................................. 57 Staphylininae ................................................................ 59 Acknowledgments ............................................................. 60 References .................................................................... 61 Addendum .................................................................... 83 2 2001 HERMAN: NOMENCLATURAL CHANGES, STAPHYLINIDAE 3 ABSTRACT More than 775 nomenclatural changes are proposed and nomenclatural problems are dis- cussed for the Staphylinidae. Two generic names are proposed as replacement names: Torobus for species formerly in Trigonopselaphus, and Ryvkinius for the preoccupied Mesoporus Ryvkin. Type species are ®xed or problems with the type species are discussed for: Entomoculia, Eumegalopsidia (an unavailable name), Anthobium, Lesteva, Phyllodrepoidea, Aleioglyphes- thus, Bolitogyrus, Cephalonthus, Diatrechus, Euremus, Indoquedius, Philothalpus, Philon- thopsis, Pseudoremus, Leptophius, Stenus (Nestus), Coproporus, and Paratachinus. The type species for Entomoculia, Anthobium, Phyllodrepoidea, Leptophius, and Stenus (Nestus) are ®xed under provisions of article 70.3 of the Code. Emendations are cited or one of multiple original spellings adopted in Bledius, Bryoporus, Carpelimus, Edaphus, Eleusis, Gauropterus, Indosorius, Lispinus, Lithocharodes, Medon, Osorius, Philonthus, Piestus, Pinophilinus, Platydracus, Priochirus, Prognathoides, Quedius, Stenus, and Trigonurus. New combinations are proposed in Arrhenopeplus (3), Dialycera (2), Eusphalerum (14), Nacaeus (19), Pseudoxyporus (4), Homalotrichus (1), Thinodromus (2), Gabrius (1), Platyd- racus (57), Torobus (9), Bryophacis (7), Ischnosoma (16), Lordithon (66), Ryvkinius (1), and Sepedophilus (244). Fifty-nine new synonyms are listed for Micropeplus (1), Anthophagus (2), Eusphalerum (1), Phloeonomus (1), Eleusis (1), Leptochirus (1), Neolosus (1), Osorius (5), Priochirus (1), An- otylus (1), Bledius (1), Carpelimus (1), Ochthephilus (1), Apoquedius (1), Diatrechus (1), Erichsonius (2), Gabrius (2), Gastrisus (1), Hesperus (2), Heterothops (2), Leptacinus (1), Notolinus (1), Phallolinus (1), Philonthus (6), Quedius (5), Staphylinus (2), Tasgius (1) Xan- tholinus (4), Stenus (7), Sepedophilus (1), and Tachinus (1). Polyphematiana E. Strand is a junior synonym of Trigonopselaphus Gemminger and Harold. Eleven names are resurrected from synonymy, one each in Phyllodrepoidea, Anotylus, Ox- ytelus, Loncovilius, Philonthus, Platydracus, Lordithon, Mycetoporus, and Tachinus and two in Stenus. Under provisions of article 23.9.1 of the Code, 28 junior synonyms are protected in Am- phichroum (1), Carpelimus (1), Deleaster (1), Proteinus (1), Bisnius (1), Leptacinus (1), Me- galinus (1), Neobisnius (1), Ocypus (1), Philonthus (3), Quedius (4), Tasgius (2), Stenus (5), Bolitobius (1), Lordithon (1), Mycetoporus (1), and Tachyporus (2). Provisions of the same article protect 5 junior homonyms in Eusphalerum (1), Xylodromus (1), Quedius (1), Tachinus (1), and Tachyporus (1). Under provisions of article 23.9.3 of the Code, 15 junior synonyms will be referred to the Commission for rulings under the plenary power. Pending the outcome of these cases, use of the junior name is maintained. The affected species are in Eusphalerum (1), Lesteva (1), Omalium (1), Phloeostiba (1), Xylodromus (1), Anotylus (1), Bledius (1), Carpelimus (1), Ocypus (1), Philonthus (1), Quedius (4), and Lordithon (1). Under provisions of article 23.9.5 of the Code, 64 junior primary homonyms will be referred to the Commission for rulings under the plenary power. Pending the outcome of these delib- erations, use of the junior name is maintained. The species are in Eusphalerum (1), Manner- heimia (1), Omaliomimus (1), Omaliopsis (1), Omalium (4), Phyllodrepa (1), Pycnoglypta (1), Xylodromus (1), Anotylus (1), Bledius (1), Carpelimus (1), Oxyporus (1), Belonuchus (2), Bisnius (1), Ca®us (2), Cheilocolpus (1), Diatrechus (1), Endeius (1), Gabrius (5), Hesperus (2), Leptacinus (1), Nordus (1), Paederomimus (1), Philonthus (10), Platydracus (3), Quedius (2), Staphylinus (6), Xantholinus (2), Xanthopygus (1), Xenopygus (2), Carphacis (1), Copro- porus (1), Tachinomorphus (1), and Tachinus (3). Replacement names are proposed for 90 preoccupied names in 37 genera: Euaesthetus (1), Leptotyphlus (1), Micropeplus (1), Eusphalerum (2), Lesteva (1), Olophrum (1), Omalium (2), Clavilispinus (1), Eleusis (3), Holotrochus (3), Leptochirus (1), Lispinus (4), Osorius (3), Priochirus (2), Thoracochirus (1), Anotylus (2), Carpelimus (2), Platystethus (1), Atanygnathus (1), Bisnius (1), Diochus (1), Gabrius (3), Hesperus (2), Heterothops (1), Leptacinus (3), Philonthus (19), Quedius (4), Staphylinus (3), Tasgius (1), Dianous (1), Bolitobius (1), Car- phacis (1), Coproporus (3), Lordithon (3), Sepedophilus (5), Tachinus (4), Tachyporus (1). 4 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY NO. 264 Eight names are resurrected to replace preoccupied names in Anotylus (1), Philonthus (1), Quedius (1), Stenus (2), Lordithon (1), Tachinomorphus (1), and Tachinus (1). Twenty-six senior homonyms, in Hapalaraea (1), Omalium (2), Phyllodrepa (1), Bledius (1), Oxytelus (1), Bisnius (1), Quedius (1), Staphylinus (15), Carphacis (1), Sepedophilus (1), and Tachinus (1), are regarded to be ``nomina dubia'' and the junior names conserved by ®at, without sanction by the Code. Psephidonus is the older name, but application to the Commission will be made to reject that name in favor of the younger Geodromicus. Tachyporiniformes is an unavailable name. INTRODUCTION Discussed in the