Cenozpic Pectinids of Alaska, Iceland, and Other Northern Regions GEOLOGICAL SURVEY PROFESSIONAL PAPER 553 Cenozoic Pectinids of Alaska, Iceland, and Other Northern Regions By F. STEARNS MAcNEIL GEOLOGICAL SURVEY PROFESSIONAL PAPER 553 Evolution of indigenous Alaskan pectinid stocks^ of migrant stocks from other areas in the Pacific region^ and of boreal Pacific species that subsequently migrated to the Arctic and northern Atlantic regions UNITED STATES GOVERNMENT PRINTING OFFICE, WASHINGTON : 1967 UNITED STATES DEPARTMENT OF THE INTERIOR STEW ART L. UDALL, Secretary GEOLOGICAL SURVEY William T. Pecora, Director Library of Congress catalog-card No. GS 66-321 For sale by the Superintendent of Documents, U.S. Government Printing Office Washington, D.C. 20402 - Price $1.00 (paper cover) CONTENTS Page Systematic paleontology Continued Abstract_ ________________________________________ 1 Genus CMamys Roeding Continued Introduction. ______________________________________ 1 Subgenus CMamys Roeding Continued Page Pectinidae as guide fossils__________________________ 1 Group of C. (C.) rubida (Hinds)..________ 21 Pectinidae of Alaska._______________________________ 2 Chlamys (Chlamys) rubida (Hinds)____ 21 Migration routes__________________________________ 3 Chlamys (Chlamys) rubida jordani (Ar­ Systematic concept.________________________________ 4 nold) 22 Systematic paleontology.____________________________ 5 Chlamys (Chlamys) rubida hindsii (Car­ Genus Delectopecten Stewart______________________ 5 penter) __________________________ 23 Delectopecten maddreni MacNeil, n. sp_________ 5 Chlamys (Chlamys) cf. C. (C.) rubida Genus Polynemamussium Habe___________________ 6 hindsii (Carpenter)_______________ 23 Polynemamussium alaskense (Dall)____________ 6 Chlamys (Chlamys) rubida prerubida Polynemamussium davidsoni (Dall)____________ 7 MacNeil, n. subsp________________ 23 Genus Arctinula Thiele__________________________ 7 Group of C. (C.) beringiana (Middendorft) _ _ 24 Arctinula groenlandica (Sowerby)_____________ 8 Chlamys (Chlamys) beringiana (Mid- Genus Chlamys Roeding-________________________ 8 dendorff) _________________ 24 Subgenus Leochlamys MacNeil, n. subgen______ 9 Chlamys (Chlamys) beringiana colvillen- Chlamys (Leochlamys) tugidakensis MacNeil sis MacNeil, n. subsp____________ 26 n. sp________________________________ 10 Chlamys (Chlamys) beringiana graui Subgenus Swiftopecten Hertlein_______________ 11 MacNeil, n. subsp_________-______ 26 Chlamys (Swiftopecten) donmilleri MacNeil, Chalmys (Chlamys) beringiana strate- n. sp________________________________ 12 gus (Dall). ------------------- 26 Chlamys (Swiftopecten) cf. C. (S.) donmilleri Chlamys (Chlamys) beringiana unalas- MacNeil___________________________ 13 kae MacNeil, n. subsp__ ___________ 27 Chlamys (Swiftopecten) sp _______________ 13 Chlamys (Chlamys) wainwrightensis Chlamys (Swiftopecten) swifti kindlei (Dall) _ 13 MacNeil, n. sp___________________ 27 Subgenus Chlamys Roeding-_________________ 13 Group of C. (C.) picoensis (Waterfall) _ _ _ _ _ 28 Group of C. ("C.") hastata (Sowerby) _____ 14 Chlamys (Chlamys) picoensis (Water­ Chlamys ("Chlamys") hastata (Sow- fall) -.------------------------ 28 by) _____________________________ 14 Chlamys (Chlamys) cf. C. (C.) picoensis Chlamys ("Chlamys") hastata hericius chinkopensis Masuda and Sawada___ 28 (Gould)__.______________________ 14 Chlamys (Chlamys) picoensis kino- Chlamys ("Chlamys") pugetensis (Old- shitai Kubota_--_-------_-------- 29 royd) ------_________________ 15 Group of C. (C.) hanaishiensis Masuda____ 29 Group of C. ("C.") tauroperstriata Sacco____ 15 Chlamys (Chlamys) cf. C. (C.) hanai­ Chlamys ("Chlamys") harmeri Altena__ 15 shiensis Masuda__---_-_---------_ 29 Chlamys ("Chlamys") breidavikensis Chlamys (Chlamys) hanaishiensis am- MacNeil, n. sp___________________ 16 chitkana MacNeil, n. subsp________ 30 Group of C. ("C.") gloriamaris (Dubois)___ 16 Chlamys (Chlamys) pseudislandica Mac­ Chlamys ("Chlamys") tjornesensis Mac­ Neil, n. sp_--------------_------- 31 Neil, n.sp_______________________ 16 Chlamys (Chlamys) pseudislandica plaf- Group of C. ("C") cosibensis (Yokoyama). 17 keri MacNeil, n. subsp____________ 32 Chlamys ("Chlamys") cf. C. ("C.") Chlamys (Chlamys) pseudislandica ar- cosibensis (Yokoyama) __ _ _________ 17 conis MacNeil, n. subsp___________ 33 Chlamys ("Chlamys") coatsi MacNeil, Group of C. (C.) islandica (Muller)________ 33 n. sp____________________________ 17 Chlamys (Chlamys) islandica islandica Chlamys ("Chlamys") coatsi middle- (Muller) 33 tonensis MacNeil, n. subsp.________ 18 Chlamys (Chlamys) islandica thulensis Chlamys ("Chlamys") cf. C. ("C.") MacNeil, n. subsp__________-----_ 34 coatsi middletonensis MacNeil_______ 18 Chlamys (Chlamys) islandica kanagae Chlamys ("Chlamys") chaixensis Mac­ MacNeil, n. subsp_______________ 35 Neil, n.sp_______________________ 19 Chlamys (Chlamys) islandica powersi Chlamys ("Chlamys") lioica (Dall)____ 19 MacNeil, n. subsp_______-_------- 35 Chlamys ("Chlamys") trinitiensis Mac­ Chlamys (Chlamys) islandica erythro- Neil, n. sp_______________________ 20 comata (Dall)_______----__----_-- 36 Chlamys ("Chlamys") aff. C. ("C.") Chlamys (Chlamys) islandica albida trinitiensis MacNeil.______________ 21 (Dall) 36 in IV CONTENTS Systematic paleontology Continued Systematic paleontology Continued Genus CMamys Roeding Continued page Genus Patinopecten Dall Continued page Subgenus CMamys Roeding Continued Subgenus Lituyapecten MacNeil-_____________ 41 Other species assigned to Chlamys. ________ 36 Patinopecten (Lituyapecten) n. sp________ 41 Chlamys sp___-__---______-------__ 36 Subgenus Mizuhopecten Masuda._____________ 41 Chlamys (?) washburnei (Arnold) _____ 36 Patinopecten (Mizuhopecten) skonunensis ? Chlamys sp______________________ 37 MacNeil, n. sp_______________________ 42 Chlamys"! nuwokensis MacNeiL_______ 37 Genus Fortipecten Yabe and Hatai._______________ 42 Genus Vertipecten Grant and Gale________________ 38 Fortipecten mollerensis MacNeil, n. sp_ ________ 43 Vertipecten lachenbruchi MacNeil, n. sp________ 39 Fortipecten hallae (Ball)_____________________ 44 Vertipecten popofensis MacNeil, n. sp __________ 39 Genus Miyagipecten Masuda.____________________ 44 Vertipecten n. sp.7-------------------------- 40 Miyagipecten alaskensis MacNeil, n. sp_ _______ 45 Genus Patinopecten Ball._________-________----__ 40 Subgenus Patinopecten Ball._________________ 41 Localities. _________________________________________ 45 Patinopecten (Patinopecten) cf. P. (P.) References.________________________________________ 49 caurinus (Gould)_____________________ 41 Index _____________________________________________ 53 ILLUSTRATIONS [Plates follow index] PLATES 1-25. Pectinids from 1. Aleutian Islands, Alaska Peninsula, Katalla district* (Alaska). 2. Alaska Peninsula, Shumagin Islands (Alaska). 3. Alaska Peninsula, Seward Peninsula, Yakataga district (Alaska). 4. Alaska Peninsula, Yakataga district, Malaspina district, Arctic coast (Alaska), Queen Charlotte Islands (British Columbia). 5. Alaska Peninsula, southeastern Alaska, Katalla district (Alaska). 6. Aleutian Islands, Malaspina district, Lituya district, Arctic coast (Alaska), Queen Charlotte Islands (British Columbia). 7. Trinity Islands, Middleton Island, Lituya district (Alaska). 8. Trinity Islands, Middleton Island (Alaska). 9. Trinity Islands, Yakataga district (Alaska). 10. Yakataga district, Lituya district (Alaska). 11. Trinity Islands, Middleton Island, Lituya district (Alaska). 12. Middleton Island (Alaska). 13. Middleton Island (Alaska). 14. Aleutian Islands, Middleton Island (Alaska). 15. Aleutian Islands, Arctic coast (Alaska). 16. Aleutian Islands, Trinity Islands, southeastern Alaska (Alaska). 17. Aleutian Islands (Alaska). 18. Aleutian Islands, southeastern Alaska, Seward Peninsula, Arctic coast (Alaska), Greenland, Iceland. 19. Southeastern Alaska, St. Lawrence Island, Arctic coast (Alaska), Iceland. 20. Aleutian Islands, Alaska Peninsula, Arctic coast, southeastern Alaska (Alaska). 21. Puget Sound (Washington), southeastern Alaska, Pribiloff Islands (Alaska), Okhotsk Sea. 22. Aleutian Islands, Sitkalidak Island off Kodiak, southeastern Alaska (Alaska), California. 23. Arctic coast (Alaska). 24. St. Lawrence Island, southeastern Alaska (Alaska), ?Bering Sea, California, Kurile Islands, Iceland, Netherlands. 25. Iceland. *Katalla district, Yakataga district, Malaspina district, and Lituya district are west to east in the outcrop of Tertiary rocks bordering the northern Gulf of Alaska. CENOZOIC PECTINIDS OF ALASKA, ICELAND AND OTHER NORTHERN REGIONS By F. STEARNS MAC^EIL ABSTRACT paper contains the first published illustration of the Oligocene to Recent pectinids from Alaska contained in the holotype of Pecten (Ghlamys) erythrovomatus Dall collections of the U.S. Geological Survey are herein described from the Okhotsk Sea and the first published photo­ and figured. New species from Iceland and the Queen Charlotte graph of the lectotype, herein designated, of Pecten is- Islands, British Columbia, and a new subspecies from Greenland landwus var. behringiana Middendorff. An attempt is are included. This paper also contains the first published figure made to rectify the nomenclature of northern Pacific of the holotype of Pecten (Chlamys) erythrocomatus Dall and the first published photograph of the lectotype, herein designated, pectinids in view of recent Japanese contributions, but of Pecten islandicus var. behringiana Middendorff. admittedly a much better knowledge of the Eussian An attempt