United States National Museum Bulletin 185 Checklist of the Coleopterous Insects of Mexico, Central America, the West Indies, and South America By RICHARD E. BLACKWELDER SMITHSONIAN INSTITUTION • WASHINGTON, D. C. • 1957 Contents Page (Part 1: xii + pp. 1-188) Preface, - v Introduction 1 Systematic list 11 (Part 2: iii+pp. 189-341) (Part 3: iv+pp. 343-550) (Part 4: iii+pp. 551-763) (Part 5: iv+pp. 765-925) (Part 6: vi+pp. 927-1492) Bibliography 927 Journals and abbreviations 1 345 Corrigenda 1 389 New names proposed 1 447 Index to genera and higher categories 1451 PUBLICATION DATES OF PARTS Part 1—March 7, 1944 Part 2—June 30, 1944 Part 3—May 21, 1945 Part 4—May 13, 1946 Part 5—October 31, 1947 Part 6—May 15, 1957 SMITHSONIAN INSTITUTION UNITED STATES NATIONAL MUSEUM Bulletin 185 CHECKLIST OF THE COLEOPTEROUS INSECTS OF MEXICO, CENTRAL AMERICA THE WEST INDIES, AND SOUTH AMERICA Part 1 COMPILED BY RICHARD E. BLACKWELDER UNITED STATES GOVERNMENT PRINTING OFFICE WASHINGTON : 1944 For sale by the Superintendent of Documents, U. S. Government Printing Office Washington, D. C. - Price 35 cents ADVERTISEMENT The scientific publications of the National Museum include two series, knowTi, respectively, as Proceedings and Bulletin. The Proceedings series, begun in 1878, is intended primarily as a medium for the publication of original papers, based on the collections of the National Museum, that set forth newly acquired facts in biology, anthro- pology, and geolog3^, with descriptions of new forms and revisions of limited groups. Copies of each paper, in pamphlet form, are distributed as pub- lished to libraries and scientific organizations and to specialists and others interested in the different subjects. The dates at which these separate papers are published are recorded in the table of contents of each of the volumes. The series of BuUeiins, the first of v/hich was issued in 1875, contains separate publications comprising monographs of large zoological groups and other general systematic treatises (occasionally in several volumes), fauna! works, reports of expeditions, catalogs of type specimens, special collections, and other material of similar nature. The majority of the volumes are octavo in size, but a quarto size has been adopted in a few instances in which large plates v/ere regarded as indispensable. In the Bulletin series appear volumes under the heading Contributions from the United States National Herbarium, in octavo form, published by the National Museum since 1902, which contain papers relating to the botanical collec- tions of the Museum. The present work forms No. 185 of the Bulletin series. Alexander Wetmore, Assistant Secretary, Smithsonian Institution. a CONTENTS Preface: Tasre In English v In Spanish vii In Portuguese ix In French xi Introduction 1 List of families and higher categories 2 List of localities 5 Acknowledgments 8 Systematic list 11 Order Coleoptera: Suborder Archostemata 11 Family Cupesidae 11 Suborder Adephaga 11 Family Rhysodidae 11 Family Cicindelidae . 11 Family Carabidae 20 Family Paussidae 72 Family Haliplidae 72 Family Dy tiscidae 72 Family Gyrinidae 80 Suborder Polyphaga 83 Family Catopidae 83 Family Colonidae 84 Family Clambidae 84 Family Leiodidae 84 Family Limnebiidae 85 Family Sphaeriidac 85 Family Ptiliidae 85 Family Scydmaenidae 87 Family Pselaphida e 88 Family Micropeplidae 97 Family Scaphidiidae 98 Family Silphidae 99 Family Staphylinidae 100 Family Hydroiihilidae 168 Family Synteliidae 175 Family Monoedidae 175 Family Histeridae 175 [Suborder POLYPHAGA to l)e continued in Part 2] PREFACE The large number of interesting and important species of insects to be found in Latin America, and the number of entomologists who have studied them, make it a matter of surprise that almost no general compilations have been produced that might aid the local students in making identifications of their specimens or in planning campaigns for the control of pests. Be- lieving that a list of the known species is a basic rec[uirement for the iden- tification of the insects and therefore for all branches of entomology, the Smithsonian Institution and the United States National Museum have undertaken the publication of this checklist for the purposes of aiding the entomologists in this region to identify local insects and of encouraging the publication of new data concerning these insects. The number of species of beetles recorded from America south of the United States is estimated to be near 50,000. In order to keep a list of this large number within reasonable size, it has been necessary to make considerable compromises with what might be considered to be an ideal checklist. With the arrangement selected, the following data are made available: A systematic arrangement of families and genera; an alpha- betical list of the species in each genus, with synonyms and varieties of each; a citation to the original publication of each name; and an indica- tion of the known geographical distribution by countries. In addition to these a preliminary bibliography of p11 papers on the taxonomy of the beetles of Latin America will be included in the final section of this work. In most details this checklist conforms with similar lists in other parts of the world. Family names end in -idae, subfamily names in -inae, tribes in -ini, and subtribes in various ways. All synonyms are in italics, and subgenera are treated in the same manner as synonyms. All names for categories lower than species (such as subspecies, variety, form, phase, or aberration) are listed under the specific name but preceded by a letter to indicate the rank assigned to them (s, v, f, p, or a, respectively). Cita- tions to original publications follow the abbreviated author's name, thus: Cupes latrcillei Sol. 49-4G0. The 49 indicates the year of publication (1849), and the 466 indicates the page. By referring to the bibliography one will find under Sober one or more papers published in 1849. The page will generally indicate which is the one concerned. Where the author's name and the reference are enclosed in parentheses this signifies that the origijial author and reference are unknown to the compiler, the name being referred to bv the author cited. VI PREFACE Geographic distribution is indicated by listing the names of all coun- tries from which the species has been recorded. A question mark before a country name indicates that the record is probably erroneous. Two country names both preceded by question marks indicate doubt as to which is the proper locality. A question mark alone indicates that no locality record has been found by the compiler but that the species probably is from our region. If a general term such as South America is given, no more specific locality was found or else the locality could not be identified on available maps. Distribution outside of Latin America is indicated by one or two additional localities, as U. S. A. or Old World (or Hawaii, Australia, or Africa). Island records are listed separately, often with a footnote reference to theu- location (geographically, not politically). All these locality records are listed in a fixed order starting in the north and proceeding southward and eastward. First Baja California and the rest of Mexico (with neighboring islands), then thi'ough Central America and South America lo Chile and neighboring islands, and finally the West Indies from the Bahamas and Cuba to Grenada. If there are only West Indian and South American records, the West Indian ones come first and aU follow from north to south as before. PROLOGO A pesar del gran numero de importantes e interesantes especies de in- sectos quo se encuentran en los paises latinoamericanos, y del numero de entomologos que los han estudiado, es sorprendente el que no se encuentre casi ninguna recopilacion general que pueda ayudar a los estudiantes en sus identificaciones de ejemplares o en el trazado de campanas para la extir- pacion de las plagas. En la creencia de que un registro de los ejemplares conocidos es im requisito fundamental para identincar los insectos y poi consiguiente para las distintas ramas de entoraologia, la Institucion Smith- soniana ha acometido la publicacion de este catalogo con el doble proposito de asistir a los entomologos de la region en sus identificaciones y de fomen- tar las publicaciones de nuevos datos concernientes a estos insectos. Se calcula que el numero anotado de los ejemplares de coleopteros en Latinoamerica llega a casi cincuenta mil. Para poder compilar una lista justipreciada de un numero tan grande ha sido necesario ilevar a cabo considerables arreglos con lo que pudieramos llamar un catalogo ideal. A continuacion damos, con la distribucion seleccionada, los datos asequibles: Un arreglo sistematico de familias y generos; una lista aifabetica de las especies de cada genero con los sinonimos y las distintas variedades de cada una; cita de la primera, publicacion del nombre de cada una, y una identi- ficacion, por paises, de la distribucion geograt'ica que se conoce. Ademas de esto se incluye, en la seccion final de este trabajo, una bibliografia prehminar de los aportes en la taxonomia de los coleopteros de Latino- america. Este catdlogo concuerda, en la mayoria de los detalles, con otros similares de distintas partes del mundo. Los nombres de la familia terminan en -idae, los de la subfamilia en -inae, los de los grupos en -ini, y los de los sub- grupos de varias maneras distintas. Todos los sinonimos estan en italica y los subgeneros se tratan de igual modo, Todos los nombres de las cate- gorias menos importantes que la especies, (tales como la subespecies, variedad, forma, fase, o aberracion) se encuentran a continuacion de la especie a que especificamente pertenecen, mas estan precedidos por una letra, (ya bien sea csta s, v,j, p, o a, respectivamcnte) la c^ue indica el rango f[ue se le ha consignado.