INSTITUT DES PARes NATIONAUX jlNSTITUUT DER NATIONALE PARKEN DU CONGO BELGE VAN BELGISCH CONGO Exploration du Pare National Albert MISSION G. F. DE WITTE (1933-1935) FASCICULE 92 Exploratie van het Nationaal Albert Park ZENDING G. F. DE WITTE (1933-1935) AFLEVERI NO 92 CALLIPHORIDAE (DIPTERA CYCLORRAPHA) Part II: RHINIINI BY FRITZ ZUMPT, Ph. D., F.R.E.S. (Johannesburg) BRUXELLES BRUSSEL 1958 1958 INSTITUT DES PARCS NATIONAUX j INSTlTUUT DER NATIONALE PARKEN DU CONGO BELGE VAN BELGISCHCONGO Exploration du Parc National Albert MISSION G. F. DE WITTE (1933-1935) FASCICU L.E 92 Exploratie van het Nationaal Albert Park ZENDING G. F. DE WITTE (1933-1935) AFL.EVERING 92 CALLIPHORIDAE (DIPTERA CYCLORRAPHA) Part Il: RH 1N Il N 1 BY FRITZ ZUMPT, Ph. D., F.R.E.S. (.Johannesburg) BRUXELLES BRUSSEL 1958 1958 1 M P R 1 MER 1 E H A Y E Z, s.p.r.1. 112. rue de Louvain, 112, Bruxelles 1 Gérant: M. Hayez, av. de l'Horizon, 39 Bruxelles 15 PARC NATIONAL ALBERT NATIONAAL ALBERT PARK J. MISSION G. F. DE WITTE 1933-1935 1. ZENDING G. F. DE WITTE 1933-1935 Fascicule 92 Aflevering 92 CALLIPHORIDAE (DIPTERA CYCLORRHAPHA) Part Il : RHINIINI EY fRITZ ZUMPT, Ph. D., f.R.E.S. (Johannesburg) PREFACE The first part of my revision of the Calliphoridae of the Ethiopian reglOn, published in 1956 as part 87 of the cc Exploration du Parc National Albert, Mission G. F. DE WITTE (1933-1935)", dealt with the Calliphorini and Chrysomyiini. 1 am now presenting the Rhiniini, the last tribe of the Calliphorinae as far as Africa South of the Sahara is concerned. The necessary remarks on the higher classification, on the morphological features of taxonomie importance (including the abbreviations), and a few on the variability have already been given in Part l, as well as my thanks ta all those institutions and private persons who lent me material or helped in sorne other respect. 4 PARC NATIONAL ALBERT Subfamily CALLIPHORINAE. RHINIINL PERIS (1952) has revised this tribe, which he regards as subfamily, on a world-wide basis. Unfortunately, ihis author has not taken into consideration the structure of the male terminalia as much as should be done in a modern revision of a fly-group. That this is necessary has become clear during the course of studies of the Palaerctic species (ZUMPT, 1956) and the Ethiopian ones. Several species have been detected which are only recognisable by the structure of this organ, whereas in others, it has been shown that the so-called outer features may vary to a high degree within the same species, that they may overlap within related forms, and that a clear recognition is sometimes only possible if the hypopygium is dissecled. The Rhiniini form a specialised group of the Calliphorinae, but they are nevertheless so closely related to them, that they should not be regarded as a higher unit than a tribe. There is a strong tendency in this group towards a reduction of the thoracic and abdominal chaetotaxy, and towards a stretching of the whole body, including the protruding of the epistome. The chaetotaxy of the head, however, is not decreased, but more or less increased, especially in the female sex. The male post-abdomen consists ilL the primitive Rkiniini of three distinct and separated (free) segments, but the anterior one is subject to a reduction, becoming more and more rudimentary, and can be totally wanting in sorne Rhiniini. There are two large genera which contain the most primitive species valued from a combination of a number of outer features and of the male genitalia, namely Isomyia and Rh'yncomya. Isomyia has the arista provided with long hairs on both sides. Assuming that the progressive reduction of the thoracic chaetotaxy is a sign of specialization, the evolution appears to have proceeded in two directions. The one leads to genera like Rhinia and Vanemdenia in which the upper aristal hairs are present, but the lower ones reduced (genera nos. 1-8); the other leads through Eurhyncomyia to Stegosoma which has the aristal hairs reduced on both sides (genera 9-16). Very little is known about the bionomics of the Rhiniini, but ail of them seem to be associated with developing stages of insects, especially termites, hymenoptera and orthoptera, on which the larvae feed as predators or parasites. The adults are commonly found on flowering plants. NATIONAAL ALBERT PARK 5 LIST OF VALID SPECIES OF RHINIINI KNOWN TO ME FROM THE ETHIOPIAN REGION. Recorded from Scientific name Belgian Congo 1 P.N.A. i. Isomyia pallens (CURRAN) •• ... ... ... ... ... + - 2. Isomyia jfavida (VILLENEUVE) .. ... ... ... ... + - .J.') Isomyia grossa (VILLENEUVE) ... ... ... ... ... + - 4. Isomyia oenlosa (VILLENEUVE) •• ... ... ... ... - - 5. Isomyia pubera (VILLENEUVE) .. ... ... ... ... + - 6. Isomyia jactatrix (VILLENEUVE) • ... ... ... .. + + 7. Isomyia calliphoroides (MALLOCH) ... ... ... ... + - 8. Isomyia tristis (BIGOT) ... ... ... ... .. ... + + 9. Isomyia connivens (VILLENEUVE) ... ... ... ... - - 10. Isomyia evanida (VILLENEUVE) . ... ... ... .. + - 11. Isomyia fasciculata (VILLENEUVE) .. , ... ... ... + - 12. l samyia nitida (CURRAN) ... ... ... ... ... .. + - 13. Isomyia cinerascens (VILLENEUVE) ... ... ... ... + - 14. Isomyia dubiosa (VILLENEUVE) ... ... ... ... + + 15. Isomyia pendula (MALLOCH) ... ... ... ... ... ? - 16. Isomyia deserti (KARSCH) ... ... ... ... ... ... + - 17. Isomyia eos n. sp. ... ... ... ... ... ... ... - - 111. Isomyia natalensis (VILLENEUVE) ... ... ... ... - - 19. Isomyia snyderi ZUMPT ... ... ... ... ... ... - - ;t0. II!/Ymyia nigripes (VILLENEUVE) • ... ... ... ... - - 21. Isamyia cuprapex (VILLENEUVE) ... ... ... ... + - ;t2. Isamyia terminata (WIEDEMANN) ... ... ... ... + - 23. Isomyia distinguenda (VILLINEUVE) •. ... ... ... + - 24. l samyia darwini (CURRAN) . ... ... ... ... ... - - 25. Isamyia cuthbertsoni (CURRAN) .. ... ... ... ... - - ;tG. Isomyia faini n. sp. ... ... ... ... ... ... ... - - 27. Isomyia longicauda (VILLENEUVE) ... ... ... ... - - 28. Isomyia angolensis (PERIS) • ... ... ... ... ... - - 29. Isomyia ellenbergi (SÉGUY) • ... ... ... ... ... - - 30. Isomyia occidental is (PERIS) ... ... ... ... ... - - 3i. Isomyia pharyge (SÉGUY) ... ... ... ... ... ... - - 32. Isomyia pluvialis (SÉGUY) •• ... ... ... ... ... - - :33. Isomyia solitaria (PERIS) .... ... ... ... ... ... + - :34. Thoracites cingulatus BEZZI ... ... ... ... ... - _. :35. Idiopsis aenea (FABRICIDS) • ... ... ... ... .. , + + 6 PARC NATIONAL ALBERT 1 Recorded from Scientific name 1 Belgian Congo P.N.A. 1 36. Idiopsis petiolata (MALLOCH) ... ... ... ... ... + - 37. Idiopsis viridis (TOWNSEND) ... ... ... ... ... - - 38. Idiopsis griseoviridis (MALLOCH) ... ... ... ... + - 39. Idiopsis prasina BRAUER and BERGENSTA~IJ\I ... - - 40. Cosmina punctUlata (WIEDEMANN) ... ... ... ... - - 41. Cosmina undulata MALLOCH ... ... ... ... ... + - 42. Cosmina rnargaritae PERIS •• ... ... ... ... ... + - 43. Cosmina gracilis CURRAN ... ... ... ... ... ... - - 44. Fainia albitarsis (MACQUART) ... ... ... ... ... - + 45. Fainia elongata (BEZZI) ... ... ... ... ... ... + + 46. Stomorhina apta CURRAN ... ... ... ... ... ... - - 47. Stomorhina arrnatipes (MALLOCH) ... ... ... ... - - 48. Stomorhina lunata (FABRICIUS) .. ... ... ... ... + + 49. Stomorhina atra (CURRAN) •• ... ... ... ... ... + - 50. Stomorhina chapini CURRAN ... ... ... ... ... + - 51. Stomorhina patrizii (PERIS) ... ... ... ... ... - - 52. Stomorhina guttata (VILLENEUVE) ... ... ... ... - - 53. Stornorhina rugosa (BIGOT) • ... ... ... ... ... + + 54. Stomorhina cribrata (BIGOT) ... ... ... ... ... + + - 55. Stomorhina tristriata (BECKER) .• ... ... '" ... - 56. Stomorhina celibe (PERIS) ... ... ... ... ... ... - - 57. Stornorhina deceptor (CURRAN) .. ... ... ... ... + - 58. Rhinia apicalis (WIEDEMANN) .. ... ... ... ... + - 59. Rhinia nigricornis (MACQUART) . ... ... ... ... + - 60. Rhinia coxendix VILLENEUVE ... ... ... ... ... + + 61. Vanemdenia africana PERIS ... ... ... ... ... - - 62. Eurhyncomyia diversicolor (BIGOT) ... ... ... ... - .. - 63. Pseudorhyncomyia braunsi (VILLENEUVE) ... ... - - 64. Rhyncomya dasyops BEZZI •• ... ... ... ... ... + - 65. Rhyncomya tetropsis (BIGOT) ... ... ... ... ... + - 66. Rhyncomya ituriensis n. sp. ... ... ... ... ... + - 67. Rhyncomya elegantula VILLENEUVE .. ... ... ... + - 68. Rhyncomya buccalis VILLENEUVE ... ... ... ... + - 69. Rhyncomya disclusa VILLENEUVE ... ... ... ... - - 70. Rhyncomya depressifrons VILLENEUVE ... ... ... - - 71. Rhyncomya currani n. n •... ... ... ... ... ... - - 72. Rhyncomya nigra PERIS ... ... ... ... ... ... - - 73. Rhyncomya messoria VILLENEUVE ... ... ... ... + - 1 NATIONAAL ALBERT PARK 7 1 1 Recorded from Scientific name 1 1 Belgia,n Congo 1 P.N.A. l , 74. Rhyncomya formosa PERIS .. ... ... ... ... .. - ­ 75. Rhyncomya hessei n. sp. ... ... ... ... ... .. - ­ 76. Rhyncomya minutalis (VILLENEUVE) ... ... ... - ­ 77. Rhyncomya maculata (MACQUART) ... ... ... .. - ­ 78. Rhyneomya interclusa VILLENEUVE .. ... ... .. - ­ 79. Rhyncomya discrepans VILLENEUVE • ... ... ... - ­ 80. Rhyneomya paradoxa n. sp. ... ... ... ... ... - ­ 81. Rhyneomya bieolor (MACQUART) • ... ... ... ... - ­ 82. Rhyneomya peraequa VILLENEUVE ... ... ... .. - ­ 83. Rhyneomya obtusa (BIGOT) • ... .. , ... ... ... + ­ 84. Rhyncomya soyauxi (KARSCH) ... ... ... ... ... + ­ 85. Rhyncomya stannocuprea SPEISER ... ... ... ... - ­ 86. Rhyncomya tristis SÉGUY ... ... ... ... ... ... ­ 87. Rhyncomya pruinosa VILLENEUVE ... .. , ... ... + ­ 1 88. Rhyncomya io PERRIS • ... ... ... ... ... .. - ­ - 1 ­ 89. Rhyncomya zumpti PERIS .. '" ... ... ... ... 1 90. Rhyneomya