Taxonomy and Biogeography of Late Cretaceous Gastropoda Dissertation Zur Erlangung des Doktorgrades der Naturwissenschaften im Fachbereich Geowissenschaften der Universität Hamburg vorgelegt von Steffen Kiel aus Köln Hamburg 2001 Als Dissertation angenommen vom Fachbereich Geowissenschaften der Universität Hamburg auf Grund der Gutachten von Prof. Dr. Klaus Bandel und Prof. Dr. Christian Spaeth Hamburg, den 2. November 2001 Prof. Dr. U. Bismayer Dekan des Fachbereichs Geowissenschaften CONTENTS Contents ....................................................................................................................1 Introduction...............................................................................................................4 Material.....................................................................................................................5 Torallola ................................................................................................................5 Temalac .................................................................................................................6 Additional material................................................................................................8 Methods ....................................................................................................................9 Taxonomy ...............................................................................................................10 Subclass Archaeogastropoda THIELE, 1925 ........................................................10 Slit-bearing (selenimorph) groups................................................................10 Family Pleurotomariidae SWAINSON, 1840..............................................11 Family Temnotropidae COX, 1960...........................................................13 Family Scissurellidae GRAY, 1847...........................................................13 Family Fissurellidae FLEMMING, 1822.....................................................15 Superfamily Trochoidea RAFINESQUE, 1815...............................................15 Family Turbinidae RAFINESQUE, 1815 ....................................................16 Subfamily Liotiinae ADAMS & ADAMS, 1854 .........................................16 Subfamily Colloniinae COSSMANN, 1916 ................................................18 Subfamily Turbininae RAFINESQUE, 1815 ...............................................19 Family Trochidae RAFINESQUE, 1815......................................................22 Subfamily Tegulinae KURODA, HABE & OYAMA, 1971..........................22 Subfamily Eucyclinae KOKEN, 1897........................................................23 Subfamily Margaritinae STOLICZKA, 1868 ..............................................30 Subfamily Trochinae RAFINESQUE, 1815 ................................................32 Subfamily Solariellinae POWELL, 1951....................................................37 Subfamily Umboniinae ADAMS & ADAMS, 1854 ....................................40 Family Skeneidae CLARK, 1851...............................................................41 Family uncertain.......................................................................................43 Superfamily Cirroidea COSSMANN, 1916 ....................................................44 Family Cirridae COSSMANN, 1916...........................................................44 Subfamily Hesperocirrinae HAAS, 1953...................................................44 Subclass Neritimorpha GOLIKOV & STAROBOGATOV, 1975 ..............................45 Superfamily Neritopsoidea RAFINESQUE, 1815...........................................45 Family Neritopsidae RAFINESQUE, 1815..................................................46 Superfamily Neritoidea RAFINESQUE, 1815 ................................................47 Family Neritidae RAFINESQUE, 1815 .......................................................47 Family Pileolidae, n. fam..........................................................................51 Subclass Caenogastropoda COX, 1960................................................................53 Order Cerithiimorpha GOLIKOV & STAROBOGATOV, 1975 ............................53 Superfamily Campaniloidea DOUVILLÉ, 1904.............................................53 Family Campanilidae DOUVILLÉ, 1904....................................................54 Superfamily Cerithioidea FÉRRUSAC, 1819.................................................58 Family Vermetidae RAFINESQUE, 1815 ...................................................58 Family Litiopidae FISCHER, 1885.............................................................60 Family Provannidae WARÉN & PONDER, 1991........................................61 Order Ptenoglossa GRAY, 1853 .......................................................................64 1 Superfamily Cerithopsoidea GOLIKOV & STAROBOGATOV, 1975 ..............64 Family Cerithiopsidae ADAMS & ADAMS, 1854......................................65 Family Eumetulidae GOLIKOV & STAROBOGATOV, 1975 .......................65 Superfamily Triphoroidea GRAY 1847 ........................................................70 Family Triphoridae GRAY 1847 ...............................................................70 Superfamily Janthinoidea LAMARCK, 1810 .................................................71 Family Epitoniidae BERRY, 1810.............................................................71 Family Nystiellidae CLENCH & TURNER 1952 ........................................72 Superfamily Zygopleuroidea WENZ, 1939...................................................74 Family Protorculidae BANDEL 1991.........................................................74 Order Strombimorpha BANDEL, 1991 .............................................................74 Superfamily Stromboidea RAFINESQUE, 1815.............................................74 Family Aporrhaidae MÖRCH, 1852 ..........................................................74 Family Strombidae RAFINESQUE, 1815....................................................78 Family Pugnellidae KIEL & BANDEL, 1999 .............................................80 Family Xenophoridae TROSCHEL, 1852...................................................80 Order Neomesogastropoda BANDEL, 1991......................................................82 Family Naticidae FORBES, 1838...............................................................82 Family Cypraeidae RAFINESQUE, 1815....................................................83 Family Trichotropidae GRAY, 1850 .........................................................84 Superfamily Cassoidea LATREILLE, 1825....................................................84 Family Ranellidae GRAY, 1854................................................................85 Order, superfamily and family uncertain.........................................................88 Order Neogastropoda THIELE, 1928................................................................90 Family Cancellariidae GRAY, 1853..........................................................90 Family Turridae SWAINSON, 1840 ...........................................................92 Family Volutodermidae PILSBRY & OLLSON, 1954.................................96 Family Athletidae PILSBRY & OLLSON, 1954..........................................98 Family Volutidae RAFINESQUE, 1815 ......................................................99 Family Pseudolividae FISCHER, 1884.....................................................101 Family Pyrifusidae BANDEL & STINNESBECK, 2000 .............................101 Subclass Heterostropha FISCHER, 1885.............................................................111 Order Allogastropoda HASZPRUNAR, 1985 ...................................................111 Family Mathildidae DALL, 1889 ............................................................111 Family Architectonicidae GRAY, 1840...................................................116 Family Amphitomariidae BANDEL, 1996...............................................118 Family Ebalidae WARÉN, 1994 ..............................................................120 Family Pyramidellidae GRAY, 1840.......................................................121 Order uncertain..............................................................................................122 Family Misurinellidae BANDEL, 1994....................................................122 Order Opisthobranchia MILNE-EDWARDS, 1848...........................................124 Family Actaeonellidae ZILCH, 1959.......................................................124 Family Acteonidae D’ORBIGNY, 1842 ...................................................125 Family Ringiculidae FISCHER 1883........................................................127 Informal group Bullomorpha..................................................................129 Order Pulmonata CUVIER, 1817 ....................................................................132 Suborder Archaeopulmonata MORTON, 1955............................................132 Family Ellobiidae ADAMS & ADAMS in PFEIFFER, 1854.......................132 Family Siphonariidae GRAY, 1840.........................................................133 Suborder Stylommatophora SCHMITT, 1855..............................................133 Biogeography........................................................................................................135 Distribution-patterns