Additions to the Rapateaceae Bassett Maguire (*) Abstract duratis, marginibus prominenter searioso-mar­ ginatis, aurieulis ineonspieuis, seariosis, ea New taxa In the Rapateaceae are described herein: 3-4 mm longis; laminis 1.5-2.0 em latis, ad Spathantheae, Ceph-alostemoneae, Stegolepis piresii, Ku nhardtia radlata, Rapatea rugulosa, Rapated ulel var. 50 em longis, anguste aeuminatis, firme papy­ latif olia, and Rapatea ulei var. lati folia forma multlca­ raeeis, prominenter venatis; peduneulis axilla­ pltata. ribus, apparenter 5-8, ea 1. 5 mm diam, valde nervosis; floribus 1·2; spieulis ea 2 em longis, sepalis inelusis, braeteolis 16-18, gradatis. Preparatory to presentation of a monograph laneeolatis, ad 8 mm longis, subaeutis, of the essentially neotropieal monoeatyledo· enerviis, induratis; sepalis ea 18 mm longis, nous family Rapateaeeae, there are herein limbis anguste laneeolatis, induratis, enerviis: offered the validation of the tribes Spathan­ petalis apparenter latiobovatis . theae and Cephalostemoneae, whieh will be TYPE- Planta de 1 m, flor amarela, 1000 m elaborated upon in the monograph, and the alt, Serra Aracá, Territorio Amazonas, Brasil, prepublieation of additional speeies repre­ 2 Nov. 1975, João Murça Pires, 15.039 (60) senting the genera Stego/epis, Kunhardtia, and (holotype NY, isotype MG) . Rapatea, respeetively. Stego/epis piresii is assigned to the sub­ seetion Pauciflora of the seetion Pauciflora, Spathantheae Maguire, trlb. nov. Stegolepis, Rapateaeeae. lts closest relatives Braetea eephalaris solitaria, spathaeea, appear to be Stego/epis pu/chefia and S. mem­ elongata. spieis Jateraliter · adnatis; granis branacea of Amazonian Venezuela. pollinis elliptieis, bisuieatis; ovariis imperfeete The leaf sheaths of Stegolepis membrana­ 3-loeularibus; loeulo solitario maturo; semi­ cea are eonspieuously broad, to 5 em wlde; nibus oblongis, striatis, exappendieulatis. Spa­ those of S. pu/chefia are indurated and pro­ thanthus Desvaux. vided w i t h eonspieuous white - searious margins and prominent aurieles; those of S · Cephalostemoneae Maguire, trib. nov. membranacea are subindurated or submembra­ nous, and are provided with narrowly searious Braeteae eephalares duae; antheris ex­ margins and rounded, non-searious auricles. appendieulatis; granis pollinis zonisuleatis; The leaf blades of S. piresii are 1/ 2 to 2/3 the ovario 3-earpellato, quoque uniovulato; semi­ width of those of the two close rei atives. nibus appendieulatis vel exappendleulatis; follis Ali members of the genus oeeur over gramineis. Cephalostemon Robert Sehomburk, Roraima Formation sandstone or eonglomerate. Duckea Maguire. Kunhardtia radiata Maguire and Steyermark, Stegolepis piresii Maguire, sp. nov. sp. nov. Seetio Paucif/ora, subseetio Pauciflora Herbae perennes terrestres; foliis 7-14 dm referenda. Stego/epis pulchellae et S. membra­ longis; vaginis valde distiehis equitantibusque naceae affinis. in uno plano dispositis, valde eoriaeeis, eon­ Herpae perennes ad 60 em altae; vaginis duplicatis, eligulatis; laminis 5-12 dm longis, anguste laneeolatis, membranaeeis vel subin- 4·6 em latis; peduneulo axillari, solitario, apiee ( • ) - The New York Botanical Garden, USA. ACfA AMAZONIA 9(2) : 267-269. 1979 -267 valde dilatato et compressa; inflorescentia This is the second species of the rapatea­ globosa interdum aliquantum depresso-globo­ ceous genus Kunhardtia assigned to the sa, 7-9 em diam; bracteis subtendentibus late Schoenoeephalieae of the subfamily Saxo­ ovato-acuminatis, ca 3 em longis, 2.5 em latis,l fridericioideae. ad maturitatem reflexis; spiculis vulgo 60-75, sessilibus, radiati- porrectatis, 3. 0-4.5 em Rapatea rugulosa Maguire, sp. nov. longis; braeteolis 16-20, apieulatis, valde gradatis, imbrieatis, purpureo-rhodanthis; ba­ Seetio E/ongata assignata; Rapatea mem­ cteolis exterioribus ovato-faneeolatis, 5-6 mm branaceae affinis. longis, 3-nervatis ad apicem, bracteolis superis Herbae perennes ad 90 em altae; vag1n1s laneeolatis, ca 20 mm longis, ineonspieue anguste laneeolatis, submembranaeeis, margi­ 3-5-nervatis ad apicem; sepalis 3, liberis, nibus anguste seariosis, ea 12-14 em longis. anguste laneeolatis, enerviis, valde induratis; petiolis vulgo 8-10 em longis, 3-4 mm latis; petalis ad basim in tubo membranaeeo con­ laminis anguste laneeolatis, vulgo 3.5 em latis, natis, sursum liberis, segmentis anguste 60-80 em longis, longiacuminatis, conspicue lanceolatis, 3-4 mm latis, conspicue exsertis, transversa venatis et rugulosis; pedunculis porrectatis; antheris muriculatis, lineanbus, 15-30 em longis, ca 4 mm diam; capitulis admodum 15 mm longis; granis pollinis ca elongatis, 5-8 em longis, bracteis involueralibus 46 X 33 X 33 .um, monosulcatis, sporodermate laneeoiatis, ea 20 em longis, anguste a­ foveolato; avario triloculari, loculis pluri­ cuminatis, axibus capitulis adnatis; recepta­ ovulatis; seminibus subpyramidalibus, striatis, cuJo paleaceo, paleis scariosis, anguste lan­ ca 3 mm diam . ceolatis; pedicellis 1-2 mm longis, bracteolis 7-8, nongradatis, 12-15 mm longis, scarioso­ 'IYPE - Plant 1.5-2.0 m tall, leaves deep membranaceis, anguste lanceolatis. 7-nerva­ green above, silvery-white below, bracts tis, pungentiacuminatis; sepalis in tubo ca subtending head silvery green within, glaucous 10-12 mm longo connatis, laminis subinduratis, green without, sepals and corolla coral red, lanceolatis, ca 5-6 mm longis; petalis longi­ floral bracts buff, increasingly more broad in unguibus; antheris ca 5 mm longis, 4-locu­ age, anthers 2-celled, ovary pluriovulate; on laribus, seminibus non visis. large igneous outcrop bordering forest, 35 km S of Puerto Ayacucho at the "Tobogán ", 85 m TYPE - Corolla yellow, beside river, Iga­ alt, Territorio Amazonas, Venezuela, 4 May rapé Lages, Manaus-Caracarai Road, Km 130, 1977, J. A. Steyermark & Otto Huber 113851 Amazonas, Brazil, 9 May 1974, G. T. Prance, B. (holotype NV, isotype VEN) . W. Nelson, O. P. Monteiro, J. F. Lima 21055 (holotype, INPA; isotype, NV). DISTRIBUT.ION - Known only by two col­ There are two additional collections known lections from a limited locality in Orinocan to me, viz: Herb with 130 em long leaves, Amazonas, growing on thin, granite-derived perianth yellow, inflorescence and base of soils. VENEZUELA. Território Amazonas: herb plant with abundant clear gelatinous material; to 3 m high, flowers red, common on "bush area with sandstone rocks along river, scrub islands" and margins of I aja, Cano Fluta, 200 m to low forest, Km 130, Manaus-Caracarai Road, alt, 50 km SE of Puerto Ayacucho, 4 Mar 1951, Amazonas, Brazil 13 Nov. 1973, Berg et a/ Maguire, Cowan, Wurdack & Baumgartner P19538; (INPA, NV, U, US) Herb 1 m alt, 31834 (NV); Steyermark & Huber 113851, as flowers yellow, banks of Igarapé Lages, Km cited above. 130, Manaus-Caracarai Highway, Amazonas, Brazil, 13 Feb 1974, Steward et a/ P20264A. Kunhardtia radiata is closely similar to K. (INPA, NY) . rhodantha, b,tJing distinguished from the latter by the more nearly spherical, less depressed Rapatea rugulosa is similar to and closely heads, the much narrower petals, and alti­ related to R. membranacea of the Guyana Ro­ tudinal and ecologic restriction . raima sandstone regions of the Pakaraima 268- Mquire 'BtBLIO~:CAdo j I ~J p A Plateau, differing ehiefly in the shorter heads Ouestions immediate ,-a1 I Sé Wfi1c can be and narrower leaf blades whieh are borne on answered only by further field observations: distinct petioles, the petioles laeking in R. Does 216568 represent a normal subpopula­ membranacea. tion, or does it represent a teratoiogieal expression within a normal population ? lt is interesting that eollectors' notes indi­ cate the new species also grows on sandstone­ Obviously these questions cannot be derived soil. Prance (verbal eommunication) answered with the evidenee available. These states that this sandstone locality is the site extraordinary specimens must receive de­ of considerable local endemism. signation as fvrma multicapitata. Rapatea ulei Pilger, Notisbl. 6: 119. 1914. Rapatea ulei Pilger, var. lat ifolia Maguire, var. nov. Rapatea ulei was collected by UI e "N9 8822, in Sümpfen zwischen Flores und Tauru­ A var. ulei simili. sed laminis foliorum ea mã, Manáos". A second colleetion of the well 15 em latis. marked species also is known to me, this TYPE - Terrestrial herb, flowers yellow. obtained by Killip and Smith, "NQ 30183, dense in wet sandy soil by stream, Km 45, Manaus· forest. Manacs". The specimens of both Caracarai Road, Amazonas, Brazil, 5 Apr 1971, eollections are uniform and possibly represent G. T. Prance et ai. 11357 (holotype. INPA; a consistent, immediately regional form. isotype, NY) . Two further eol lections now at hand, made ADDITIONAL SPECIMENS - leaves to 80 em by Pranee and assoeiates, Nos. 3827 and tall, corolla yellow, flowers borne in mucilage; 11357, were obtained along the Manaus-Cara­ varzea forest, Igarapé Leão, road Manaus-Ca­ carai Road at kilometers 12 and 45, re­ racarai, Amazonas, Brazil, Km 12, 28 Dec. 1966, spectively. These represent again a uniform Prance, Pena & Ramos 3827 (INPA, NY); leaves facies, but are distinguished from the Ule and to 1.5 m long, corolla yellow, varzea forest, Killip and Smith specimens by their leaves 3 km west of Reserva Experimental Station which are twiee the width of the original, and, of INPA, Manaus-Caracarai Highway, Amazo­ thus, are conspieuously different in facies nas Brazil, 15 Aug 1974, Prance, Pennington, from it. These latter plants are designated as et ai 21656A (NY, US). Rapatea ulei var.