INFORMATION TO USERS This reproduction was made from a copy of a document sent to us for microfilming. While the most advanced technology has been used to photograph and reproduce this document, the quality of the reproduction is heavily deper. .ru upon the quality of the material submitted. The following explanation of techniques is provided to help clarify markings or notations which may appear on this reproduction. 1.The sign or “target” for pages apparently lacking from the document photographed is “Missing Page(s)”. If it was possible to obtain the missing page(s) or section, they are spliced into the film along with adjacent pages. This may have necessitated cutting through an image and duplicating adjacent pages to assure complete continuity. 2. When an image on the film is obliterated with a round black mark, it is an indication of either blurred copy because of movement during exposure, duplicate copy, or copyrighted materials that should not have been filmed. For blurred pages, a good image of the page can be found in the adjacent frame. If copyrighted materials were deleted, a target note will appear listing the pages in the adjacent frame. 3. When a map, drawing or chart, etc., is part of the material being photographed, a definite method of “sectioning” the material has been followed. It is customary to begin filming at the upper left hand comer of a large sheet and to continue from left to right in equal sections with small overlaps. If necessary, sectioning is continued again—beginning below the first row and continuing on until complete. 4. For illustrations that cannot be satisfactorily reproduced by xerographic means, photographic prints can be purchased at additional cost and inserted into your xerographic copy. These prints are available upon request from the Dissertations Customer Services Department. 5. Some pages in any document may have indistinct print. In all cases the best available copy has been filmed. University M icrofilms International 300 N. Zeeb Road Ann Arbor, Ml 48106 8426371 Cichan, Michael Anthony VASCULAR CAMBIUM AND WOOD DEVELOPMENT IN SELECTED CARBONIFEROUS PLANTS The Ohio State University Ph.D. 1984 University Microfilms International300 N. Zeeb Road, Ann Arbor, Ml 48106 Copyright 1984 by Cichan, Michael Anthony All Rights Reserved PLEASE NOTE: In all cases this material has been filmed in the best possible way from the available copy. Problems encountered with this document have been identified here with a check mark V . 1. Glossy photographs or pages 2. Colored illustrations, paper or print ____ 3. Photographs with dark background _____ 4. Illustrations are poor copy _____ 5. Pages with black marks, not original copy _ 6. Print show s through as there is text on both sid es of p ag e _______ 7. Indistinct, broken or small print on several pages 8. Print exceeds margin requirements ______ 9. Tightly bound copy with print lost in spine ______ 10. Computer printout pages with indistinct print ______ 11. Page(s) ____________ lacking when material received, and not available from school or author. 12. P ag e(s)_____________ seem to be missing in numbering only as text follows. 13. Two pages numbered _____________ . Text follows. 14. Curling and wrinkled pages _______ 15. O ther _________________________________________ ________________________________ _ University Microfilms International VASCULAR C&flBJUB AID ROOD DEVELOPREHT I » SELECTED cmmSIFEEQUS FLAWS DISSERTATION Presented in Partial Fulfillment of the Requirements for the degree Doctor of Philosophy in the Graduate School of The Ohio State University By Hichael Anthony Cichan* B»A-* H-Sc- * * # # * The Chio State University 1984 Reading Committee; Approved By Thomas N„ Taylor Gary L- Floyd Daniel J. Crawford Ralph E- Boecner Thomas N Taylc Department of Botany C opyright by M ichael Anthony Cichan 1984 D U IC 1 U 0 1 ®enory of my mother# Cecelia mmoimimsims I ttould like to thank Dr s. David F® Brauer® Charles a. Good® G ilbert H® leisraan® and Benton H® Stidd for graceous- Ij alXo@iag ae to examine their fossil plant collections and providing some of the specimens used in this study® I bouM also like to thank the members of ay reading commit­ tee for helpful comments on this work. Financial assis­ tance fro® the Ohio State University Graduate School in the fore of a Graduate Student Alumni fiesearch heard and a Uni­ v e r s i t y Presidential Fellowship is also gratefully ack n o w l­ edged. This investigation was c a r r i e d out under the s u p e r­ v is io n o f Dr® Thomas N. Taylor® whose expertise and enthusiastic manner encouraged and stimulated my interest in the biology of fossil plants. A student c o u ld a sk no more than this of a teacher® and I thank him for his sup­ port® advice® and friendship® Finally® I would like to extend my deepest appreciation to my wife® Paula® and my children® David and Cecelia® for giving me the love and understanding that was so critical for the completion of this project® fWl O c to b er 2 8 4 , 195# ............ Born - Philadelphia* Pennsylvania 1978 Bo A. * LnS a H e C ollege* Philadelphia* Pennsylvania 1977 - 1979 ooooooooooooo^oo leaching assistant* Rutgers University* Newark* New Jersey 1978 - 1979 teaching Assistant* Butgers University* New Brunswick* Ken J e r s e y 1979 Ho Sc.* Butgers University* New Brunswick* New Jersey 1980 - 1981 0 ^ 0000.000000000 T each in g A s s is ta n t the Ohio State University* Colum bus* Ohio 1982 - 1983 oooooo<ooo<oooo0oo R ese a rc h A s s i s t a n t The Chi© State University* Columbus* Ohio 1983 - 198# ...o.o.oooooo.o. Presidential Fellow* The Ohio State University* Columbus* Ohio Professional Affiliations Botanical Society of America* Paleobotanical Section Botanical Society o f America* Structural/Developmental Section American Association for the Advancement of Science Palaeontological Society* London International Association of Hood Anatomists International Organization of Paleobotany The Society of the Sigma Si The Ohio Academy o f S c ie n c e The Torrey B o ta n ica l Club Research I n t e r e s t s Structure and evolution o f the vascular cambium Anatomy and morphology of Carboniferous plants Reproductive biology and evolution of angiosperms Electron microscopy of fossil pollen and spores A&rards and H onors 1983 The Ohio State University Presidential Fellowship 1983 The Ohio State University Graduate Student Alumni Research Association Award 1983 Isabel Co Cooksoa Award* Paleobotanical Section* Botanical Society -of America* best student paper presented at annual meeting Extramural Grants 1981 - 1982 Grant for student research* The Geological S o c ie ty of America; Project Title; Characterization of the megafossil flora of the Cranks Creek coal ball locality (Carboniferous) in eastern Kentucky. $600 1983 National Science Foundation Doctoral Dissertation Grant for Research in the F ie ld S c ie n c e s ; Project Title: The structure* organization* and activity of the vascular cambium in Pennsylvanian- age plants. $4060 Service to the Ohio State University 1980 Student representative on the Space and Facilities Committee (Department of Botany) 1981 Member of the Sailer lecture Committee (Department o f Botany) 1982 - 1983 Student representative on th e Colloquium Committee (Department of Botany) Publications 1979a Parthenocarpy in Cichoriua intvbus. Bot. Sac. Amer. Hisc. Ser. P u b l. 157s 7 . "(M. A. C ichan and B. F. P a ls e r ) . - v - 1979b Parthenocarpic achenes in Cichoriua iotyfcus L. B u ll, lea Jersey Acad. Sci. 38s 38. ( 8 . A. Cichan and B. P. Falser). 1980 Phylogenetic trees in plant systematics. O hio J. Sci. 80: 22. | f . Steussy, V. Funk* and M. A. Cichan). 1981a Oa th e s t r u c t u r e and morphology of the seed Pachvtesfca m uacii a . sp. (Bedullosales). fie?. Palaeobot. Palynol. 34s 359-367- (fl. A. Cichan and T. H. T a y lo r ). 1981b The application of scanning electron microscopy in the characterization of Carboniferous lycopod wood. Scanning Electron flicrosc. 1981. 3s 197-201. |KU 1. Cichan^ T= I . T a y lo r s and 2. L. Smoot) . 1981c In analysis of cambial development in Sphenophvllum. Chio J. Sci. 81s 29. 1981d Stauropteris in the Carboniferous of North America. B ot. Soc. Amer. fiisc. Ser. Publ. 160s 4 3. |M. A- Cichan and T. N. T a y lo r ) . 1981e The v a scu la r c a ib iu ® in Carboniferous plants: Sphenophylluta. Bot- Soc. Amer. ftisc. Ser. Pull. 160: 4 3 . 1982a Development of seedless and normal achenes in Cichcrium intybus (Compositae). Amer. J. Bot. 69- 885-895. p. A- Cichan and B. F. F a lse r) . 1982b Structurally preserved plants from southeastern K entucky: Stauaropfceris tiseriata sp- nov. Amer. J. Bot. 69: 1491-1496- (H. A- Cichan and T. N. Taylor). 1982c Plant-animal interactions in the Carboniferous: Wood borings in Premnorylon. Palaeogeography, Palaeo- climatology, Palaeoecology 39: 123-127. |M. A. Cichan and T. N. Taylor)- 1982d Vascular cambium development in SphenophyHum, a Carboniferous arthrophyte. IAWA Bull- 3: 155-160. (H. A- Cichan and I. N. Taylor). 1982e Book Beview: Paleobotany, Paleoecology, and Evolution. Volume 1- Karl J. Niklas (ed.). International Organization of Falaeolotany Newsletter #19. 1982f The e a r lie s t-k n o w n arthropod burrows in fossil wood Ohio J® S c i, 82s 17. fff® A® C ich an and T. B. T aylor) 1982g Sood b o rin g s i n Premaozylons Plant-animal interactions i n th e Carboniferous® Bot® Soc® Amer.