Arthropoda Selecta 23(2): 145188 © ARTHROPODA SELECTA, 2014 Heteropoda Latreille, 1804: new species, synonymies, transfers and records (Araneae: Sparassidae: Heteropodinae) Heteropoda Latreille, 1804: íîâûå âèäû, ñèíîíèìèÿ, êîìáèíàöèè è íàõîäêè (Araneae: Sparassidae: Heteropodinae) Peter Jäger Ï. ßãåð Arachnology, Senckenberg Research Institute, Senckenberganlage 25, 60325 Frankfurt am Main, Germany. E-mail: peter. [email protected] KEY WORDS: Huntsman spiders, Southeast Asia, systematics, revision. ÊËÞ×ÅÂÛÅ ÑËÎÂÀ: ïàóêè-ñïàðàññèäû, Þãî-Âîñòî÷íàÿ Àçèÿ, ñèñòåìàòèêà, ðåâèçèÿ. ABSTRACT. The genus Heteropoda Latreille, 1804 1988) comb.n., Pseudopoda straminiosa (Kundu, Bis- is revised by means of original descriptions and re- was et Raychaudhuri, 1999) comb.n., Gnathopalystes spective type material as well as of material recently denticulatus Saha et Raychaudhuri, 2007 comb.n., Olios received. Seven new species are described: H. pekkai bhattacharjeei Saha et Raychaudhuri, 2007 comb.n., sp.n. from Bhutan ($), H. opo sp.n. from caves in Olios debalae (Biswas et Roy, 2005) comb.n., Spario- Burma ($), H. cece sp.n. from a cave in Indonesia, lenus buxa (Saha, Biswas et Raychaudhuri, 1995) Kalimantan (#), H. duo sp.n. (#) from Indonesia, comb.n. (all ex Heteropoda). Heteropoda camelia Kalimantan, H. kusi sp.n. from a cave in Indonesia, Strand, 1914 belongs not to Heteropoda, but to an Kalimantan (#$), H. maukin sp.n. from a cave in unrecognisable genus of Sparassidae. Biswas & Ray- Indonesia, Kalimantan ($), H. obe sp.n. from a cave in chaudhuri [2005] illustrate not a H. venatoria, but an Indonesia, Sulawesi (#$). Lectotypes are designated unidentifiable species of Sparassidae. The following for: H. mediocris Simon, 1887 ($); H. kobroorica species described from juveniles are considered nomi- Strand, 1911 ($). Redescriptions are provided for: H. na dubia: H. annulipoda Strand, 1911 from Aru Is- listeri Pocock, 1900 from # and $ syntypes from lands, H. bimaculata Thorell, 1878 from Ambon, H. Christmas Island; H. nigriventer Pocock, 1897 from bivittata Thorell, 1887 from Sulawesi, H. megalopis recently collected material from Central Sulawesi; H. (Thorell, 1891) from Nicobar Islands, and H. panareti- mediocris Simon, 1887 from female lectotype from formis Strand, 1906 from Sumatra. Heteropoda zuviele New Guinea; H. languida Simon, 1887 from $ holo- Jäger, 2008 and H. simplex Jäger et Ono, 2000 are type from Burma; H. kabaenae Strand, 1911 from $ recorded the first time from Taiwan, H. boiei (Dole- holotype from Indonesia, Sulawesi; H. analis Thorell, schall, 1859) from Borneo, H. parva Jäger, 2000 from 1881 from $ holotype from Fly River, Papua New Kalimantan and Sarawak. Heteropoda beroni Jäger, Guinea. Following new synonymies are proposed: H. 2005 is recorded from the cave Gua Istani Taokala kobroorica Strand, 1911. syn.n. with H. analis Thorell (Sulawesi, Maros karst, Bantimurung). Heteropoda sp. 1881; H. regalis Roewer, 1938, syn.n. with H. medioc- cf. sumatrana Thorell, 1890 ($) is recorded from Java, ris; H. laurae Jäger, 2008 syn.n. with H. nigriventer Cibodas, described and illustrated. Pocock, 1900; Ocypete thoracica C.L. Koch, 1845, syn.n., Sarotes aulicus L. Koch, 1878, syn.n., Sarotes ÐÅÇÞÌÅ. Ðåâèçîâàí ðîä Heteropoda Latreille, truncus McCook, 1878, syn.n., H. venatoria japonica 1804: ïðèâåäåíû îðèãèíàëüíûå îïèñàíèÿ, èññëåäî- Strand, 1907, syn.n., H. venatoria chinesica Strand, âàíû òèïîâûå ýêçåìïëÿðû, èçó÷åíû íîâûå ìàòåðè- 1907, syn.n., H. venatoria maculipes Strand, 1907, àëû. Îïèñàíî 7 íîâûõ âèäîâ: H. pekkai sp.n. (Áóòàí, syn.n., H. nicki Strand, 1915, syn.n., H. nicki quala $), H. opo sp.n. (ïåùåðà â Áèðìå, $), H. cece sp.n. Strand, 1915, syn.n., H. tokarensis Yaginuma, 1961, (ïåùåðà â Èíäîíåçèè, Êàëèìàíòàí, #), H. duo sp.n. syn.n., H. andamanensis Tikader, 1977, syn.n., H. nico- (Èíäîíåçèÿ, Êàëèìàíòàí, #), H. kusi sp.n. (ïåùåðà barensis Tikader, 1977, syn.n. and H. hainanensis Li, â Èíäîíåçèè, Êàëèìàíòàí, #$), H. maukin sp.n. (ïå- 1991, syn.n. with Heteropoda venatoria (Linnaeus, ùåðà â Èíäîíåçèè, Êàëèìàíòàí, $), H. obe sp.n. 1767). The following new combinations are established: (ïåùåðà â Èíäîíåçèè, Ñóëàâåñè, #$). Âûäåëåíû Heteropoda megalopis (Thorell, 1891) comb.n. (ex ëåêòîòèïû äëÿ: H. mediocris Simon, 1887 ($); H. Spariolenus Simon, 1880); Barylestis manni (Strand, kobroorica Strand, 1911 ($). Ïåðåîïèñàíû: H. listeri 1906) comb.n., Pseudopoda aureola (He et Hu, 2000) Pocock, 1900 ïî ñèíòèïàì # è $ ñ î. Ðîæäåñòâà; H. comb.n., Pseudopoda shillongensis (Sethi et Tikader, nigriventer Pocock, 1897 íà îñíîâå íåäàâíî ñîáðàí- 146 P. Jäger íîãî ìàòåðèàëà èç öåíòðàëüíîãî Ñóëàâåñè; H. me- onyms of the pantropical H. venatoria (Linnaeus, 1767) diocris Simon, 1887 ïî ëåêòîòèïó $ èç Íîâîé Ãâè- or are to be transferred to other genera. Another spe- íåè; H. languida Simon 1887 ïî ãîëîòèïó $ èç Áèð- cies, H. pumilla Keyserling, 1880, was recently trans- ìû; H. kabaenae Strand, 1911 ïî ãîëîòèïó $ èç Èí- ferred to the genus Sparianthina Banks, 1929 [Jäger et äîíåçèè, Ñóëàâåñè; H. analis Thorell, 1881 ïî al., 2009]. Seven species are described by juveniles ãîëîòèïó ñ Fly River, Ïàïóà Íîâàÿ Ãâèíåÿ. Óñòà- only, thus have to be considered nomina dubia think- íîâëåíà íîâàÿ ñèíîíèìèÿ: H. kobroorica Strand, ing of the huge amount of mostly similar species in 1911, syn.n. = H. analis Thorell, 1881; H. regalis respect of somatic characters. In the present paper, 7 Roewer, 1938, syn.n. = H. mediocris; H. laurae Jäger, species are newly described, 15 are synonymised, 1 is 2008 syn.n. = H. nigriventer Pocock, 1900; Ocypete transferred to Heteropoda and 8 are transferred to oth- thoracica C.L. Koch, 1845, syn.n., Sarotes aulicus L er genera, resulting in a current total of 211 valid Koch, 1878, syn.n., Sarotes truncus McCook, 1878, Heteropoda species, including 15 nomina dubia. syn.n., H. venatoria japonica Strand, 1907, syn.n., H. Recently, Jäger [2008b] described 25 new Heteropo- venatoria chinesica Strand, 1907, syn.n., H. venatoria da species from Asia and Australia focussing on spe- maculipes Strand, 1907, syn.n., H. nicki Strand, 1915, cies with exceptional male palp conformations, e.g. of syn.n., H. nicki quala Strand, 1915, syn.n., H. tokarensis the Heteropoda javana-group. Eusemann & Jäger Yaginuma, 1961, syn.n., H. andamanensis Tikader, [2009] recorded a high variation in one widely distrib- 1977, syn.n., H. nicobarensis Tikader, 1977, syn.n. è uted species (H. tetrica). Jäger & Bayer [2009] de- H. hainanensis Li, 1991, syn.n. = Heteropoda venatoria scribed two new species from Laos, and Bayer & Jäger (Linnaeus, 1767). Óñòàíîâëåíû ñëåäóþùèå íîâûå [2009] recorded the male of H. ocyalina (Simon, 1887) êîìáèíàöèè: Heteropoda. megalopis (Thorell, 1891) for the first time. In the meantime new material was comb.n. (ex Spariolenus Simon, 1880); Barylestis trusted to the author and, additionally, type material of manni (Strand, 1906) comb.n., Pseudopoda aureola previously described species was either re-examined or (He et Hu, 2000) comb.n., Pseudopoda shillongensis examined for the first time resulting in new synonyms (Sethi et Tikader, 1988) comb.n., Pseudopoda stra- and redescriptions. miniosa (Kundu, Biswas et Raychaudhuri, 1999) comb.n., Gnathopalystes denticulatus Saha et Ray- Material and methods chaudhuri, 2007 comb.n., Olios bhattacharjeei Saha et Raychaudhuri, 2007 comb.n., Olios debalae (Biswas The examined spiders were preserved in 70% dena- et Roy, 2005) comb.n., Spariolenus buxa (Saha, Biswas tured ethanol. Examination and drawings were carried et Raychaudhuri, 1995) comb.n. (âñå ex Heteropoda). out with a Leica MZ 16 stereomicroscope with camera Heteropoda camelia Strand, 1914 ïðèíàäëåæèò íå ê lucida attachment. Female copulatory organs were dis- ðîäó Heteropoda, à ê ðîäó Sparassidae, êîòîðûé íå- sected and the sclerotised internal duct system was cleared in 96% DL-lactic acid (C H O ). All measure- âîçìîæíî èäåíòèôèöèðîâàòü. Biswas & Raychaud- 3 6 3 huri [2005] ïðîèëëþñòðèðîâàëè íå H. venatoria, à ments are in mm. Prosoma length/width means length/ íå îïðåäåëÿåìûé ïî ðèñóíêó âèä of Sparassidae. width of the dorsal shield of prosoma. Leg formula, leg Ðÿä âèäîâ, îïèñàííûõ ïî íåïîëîâîçðåëûì îñîáÿì spination pattern and size classes follow Jäger [2001]. ñ÷èòàþòñÿ nomina dubia: H. annulipoda Strand, 1911 Palp and leg lengths are listed as: total (femur, patella, (î-âà Àðó), H. bimaculata Thorell, 1878 (Àìáîí), H. tibia, metatarsus, tarsus). Arising points of tegular ap- bivittata Thorell, 1887 (Ñóëàâåñè), H. megalopis (Tho- pendages in males are described as clock-positions of rell, 1891) (Íèêîáàðñêèå î-âà) è H. panaretiformis the left pedipalp in ventral view. The part of the inter- Strand, 1906 (Ñóìàòðà). Heteropoda zuviele Jäger, nal duct system with glandular pores is called turning 2008 è H. simplex Jäger et Ono, 2000 âïåðâûå îòìå- point, as at this point the duct system changes its ÷åíû íà Òàéâàíå, H. boiei (Doleschall, 1859) íà direction. In schematic illustration of the courses it is Áîðíåî, H. parva Jäger, 2000 íà Êàëèìàíòàíå è â marked with T, the copulatory orifice with a circle, Ñàðàâàêå. Heteropoda beroni Jäger, 2005 îòìå÷åíà â and the end of the fertilisation duct in direction of the ïåùåðå Gua Istani Taokala (Ñóëàâåñè, êàðñò Maros, uterus externus with an arrow. As in Jäger [2005: 88], Bantimurung). Heteropoda sp. cf. sumatrana Thorell, slit sensilla close to the epigyne are illustrated as de- 1890 ($) îòìå÷åíà íà ßâå, Cibodas, îïèñàíà è ïðî- scriptive character. èëëþñòðèðîâàíà. Valid Heteropoda species are