GENETICS | THE 2015 GSA HONORS AND AWARDS Previous Recipients of GSA Awards Thomas Hunt Morgan Medal 1981 Barbara McClintock and 1991 Armin Dale Kaiser 2003 David S. Hogness Marcus M. Rhoades 1992 Edward H. Coe, Jr. 2004 Bruce N. Ames 1982 Sewall Wright 1993 Ray D. Owen 2005 Robert L. Metzenberg 1983 Edward B. Lewis 1994 David D. Perkins 2006 Masatoshi Nei 1984 George W. Beadle and 1995 Matthew Meselson 2007 Oliver Smithies R. Alexander Brink 1996 Franklin W. Stahl 2008 Michael Ashburner 1985 Herschel L. Roman 1997 Oliver Evans Nelson, Jr. 2009 John Roth 1986 Seymour Benzer 1998 Norman H. Horowitz 2010 Alexander Tzagoloff 1987 James F. Crow 1999 Salome G. Waelsch 2011 James Haber 1988 Norman H. Giles 2000 Evelyn M. Witkin 2012 Kathryn V. Anderson 1989 Dan L. Lindsley 2001 Yasuji Oshima 2013 Thomas Douglas Petes 1990 Charles Yanofsky 2002 Ira Herskowitz 2014 Frederick M. Ausubel Genetics Society of America Medal 1981 Beatrice Mintz 1992 Maynard V. Olson 2004 Trudy F. C. Mackay 1982 Gerald R. Fink 1993 Jonathan R. Beckwith 2005 Steven J. Elledge 1983 Charles Yanofsky 1994 Leland H. Hartwell 2006 Victor Ambros 1984 David S. Hogness 1995 Eric Wieschaus 2007 Shirley Tilghman 1985 Philip Leder 1996 Elliot Meyerowitz 2008 Susan Lindquist 1986 Gerald M. Rubin 1997 Christine Guthrie 2009 Marian Carlson 1987 Sydney Brenner 1998 Ronald W. Davis 2010 Barbara Meyer 1988 David Botstein and 1999 Charles H. Langley 2011 John Carlson Ira Herskowitz 2000 Jack W. Szostak 2012 Joanne Chory 1989 Allan C. Spradling 2001 H. Robert Horvitz 2013 Elaine A. Ostrander 1990 Nancy Kleckner 2002 Andrew Z. Fire 2014 Angelika B. Amon 1991 Bruce S. Baker 2003 Jeffrey C. Hall George W. Beadle Award 1999 Michael Ashburner 2003 Gerald M. Rubin and 2009 Jay C. Dunlap 2000 John Sulston and Allan C. Spradling 2010 William M. Gelbart Robert Waterston 2004 Norbert Perrimon 2011 Joseph R. Ecker 2001 Gerald R. Fink 2005 Thomas C. Kaufman 2012 Therese Markow 2002 Robert Mortimer and 2006 Fred Sherman 2013 R. Scott Hawley André Goffeau 2007 Robert K. Herman 2014 Hugo J. Bellen 2008 Mark Johnston Genetics Society of America Elizabeth W. Jones Award for Excellence in Education 2007 Elizabeth W. Jones 2010 Utpal Banerjee 2013 A. Malcolm Campbell 2008 R. Scott Hawley 2011 Peter Bruns 2014 Robin Wright 2009 Sarah C. R. Elgin 2012 David A. Micklos Genetics, Vol. 200, 683–684 July 2015 683 Novitski Prize 2008 Thomas J. Silhavy 2011 Abby F. Dernburg 2014 Charles Boone 2009 Rodney Rothstein and Kent Golic 2012 Dana Carroll 2010 Thomas W. Cline 2013 Jonathan Pritchard 684 Honors and Awards.